EXHIBIT A
Joint Filing Statement
We, the undersigned, hereby express our agreement that the attached Schedule 13D/A is filed on behalf of each of us.
Dated as September 22, 2016 | ||||||||
THE COLUMN GROUP, LP | THE COLUMN GROUP GP, LP | |||||||
By: | The Column Group GP, LP | |||||||
By: | /s/ Peter Svennilson | By: | /s/ Peter Svennilson | |||||
Name: | Peter Svennilson | Name: | Peter Svennilson | |||||
Title: | Managing Partner | Title: | Managing Partner | |||||
THE COLUMN GROUP II, LP | THE COLUMN GROUP II GP, LP | |||||||
By: | The Column Group II GP, LP | |||||||
By: | /s/ Peter Svennilson | By: | /s/ Peter Svennilson | |||||
Name: | Peter Svennilson | Name: | Peter Svennilson | |||||
Title: | Managing Partner | Title: | Managing Partner | |||||
By: | /s/ Peter Svennilson | |||||||
Peter Svennilson | ||||||||
By: | /s/ David V. Goeddel | |||||||
David V. Goeddel |