Portolan Capital Management 13F-HR Quarterly holdings report by institutional manager

The Securities and Exchange Commission has not necessarily reviewed the information in this filing and has not determined if it is accurate and complete.
The reader should not assume that the information is accurate and complete.

UNITED STATES SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549
FORM 13F

FORM 13F INFORMATION TABLE

OMB APPROVAL

OMB Number:		3235-0006
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COLUMN 1	COLUMN 2	COLUMN 3	COLUMN 4	COLUMN 5			COLUMN 6	COLUMN 7	COLUMN 8
			VALUE	SHRS OR	SH/	PUT/	INVESTMENT	OTHER	VOTING AUTHORITY
NAME OF ISSUER	TITLE OF CLASS	CUSIP	(x$1000)	PRN AMT	PRN	CALL	DISCRETION	MANAGER	SOLE	SHARED	NONE
AARON'S, INC.	COM	002535300	3,257	142,960	SH		SOLE		142,960	0	0
ABIOMED, INC.	COM	003654100	1,764	12,155	SH		SOLE		12,155	0	0
ACTIVISION BLIZZARD, INC.	COM	00507V109	7,219	121,374	SH		SOLE		121,374	0	0
AKAMAI TECHNOLOGIES, INC.	COM	00971T101	19,733	215,681	SH		SOLE		215,681	0	0
ALARM.COM HOLDINGS, INC.	COM	011642105	3,524	90,580	SH		SOLE		90,580	0	0
ALEXANDRIA REAL ESTATE EQUITIES, INC.	COM	015271109	10,170	74,199	SH		SOLE		74,199	0	0
ALPHABET, INC.	COM	02079K107	15,219	13,088	SH		SOLE		13,088	0	0
ALTA EQUIPMENT GROUP, INC.	COM	02128L106	1,336	312,777	SH		SOLE		312,777	0	0
AMAZON.COM, INC.	COM	023135106	15,643	8,023	SH		SOLE		8,023	0	0
AMERICAN PUBLIC EDUCATION, INC.	COM	02913V103	4,646	194,135	SH		SOLE		194,135	0	0
AMERICAS CAR MART, INC.	COM	03062T105	6,323	112,214	SH		SOLE		112,214	0	0
ANGI HOMESERVICES, INC.	COM	00183L102	2,347	447,003	SH		SOLE		447,003	0	0
ANIKA THERAPEUTICS, INC.	COM	035255108	319	11,032	SH		SOLE		11,032	0	0
APPLIED GENETIC TECHNOLOGIES CORPORATION	COM	03820J100	1,561	475,796	SH		SOLE		475,796	0	0
AXOGEN, INC.	COM	05463X106	1,706	164,073	SH		SOLE		164,073	0	0
AXT, INC.	COM	00246W103	2,240	697,879	SH		SOLE		697,879	0	0
BANDWIDTH, INC.	COM	05988J103	23,830	354,136	SH		SOLE		354,136	0	0
BIO-RAD LABORATORIES, INC.	COM	090572207	23,016	65,656	SH		SOLE		65,656	0	0
BLACKLINE, INC.	COM	09239B109	1,606	30,534	SH		SOLE		30,534	0	0
BOINGO WIRELESS, INC.	COM	09739C102	2,523	237,749	SH		SOLE		237,749	0	0
BRIGHAM MINERALS, INC.	COM	10918L103	592	71,539	SH		SOLE		71,539	0	0
CACI INTERNATIONAL, INC.	COM	127190304	8,310	39,356	SH		SOLE		39,356	0	0
CAL-MAINE FOODS, INC.	COM	128030202	3,249	73,870	SH		SOLE		73,870	0	0
CASELLA WASTE SYSTEMS, INC.	COM	147448104	5,520	141,316	SH		SOLE		141,316	0	0
CASTLE BIOSCIENCES, INC.	COM	14843C105	4,044	135,654	SH		SOLE		135,654	0	0
CHEGG, INC.	COM	163092109	8,901	248,772	SH		SOLE		248,772	0	0
CHINA ONLINE EDUCATION GROUP	COM	16954L105	1,044	35,997	SH		SOLE		35,997	0	0
CISCO SYSTEMS, INC.	COM	17275R102	1,964	49,971	SH		SOLE		49,971	0	0
CLOUDERA, INC.	COM	18914U100	5,769	733,089	SH		SOLE		733,089	0	0
CROWDSTRIKE HOLDINGS, INC.	COM	22788C105	1,986	35,668	SH		SOLE		35,668	0	0
CYRUSONE, INC.	COM	23283R100	4,020	65,096	SH		SOLE		65,096	0	0
DOLLAR GENERAL CORPORATION	COM	256677105	4,956	32,819	SH		SOLE		32,819	0	0
DOLLAR TREE STORES, INC.	COM	256746108	7,396	100,662	SH		SOLE		100,662	0	0
DOMO, INC.	COM	257554105	590	59,372	SH		SOLE		59,372	0	0
DOUGLAS EMMETT, INC.	COM	25960P109	2,227	73,008	SH		SOLE		73,008	0	0
E.L.F. BEAUTY, INC.	COM	26856L103	6,799	690,921	SH		SOLE		690,921	0	0
EBIX, INC.	COM	278715206	2,028	133,582	SH		SOLE		133,582	0	0
EHEALTH, INC.	COM	28238P109	12,769	90,673	SH		SOLE		90,673	0	0
ELECTRONIC ARTS	COM	285512109	11,344	113,243	SH		SOLE		113,243	0	0
EMERGENT BIOSOLUTIONS, INC.	COM	29089Q105	7,618	131,659	SH		SOLE		131,659	0	0
ENCORE CAPITAL GROUP, INC.	COM	292554102	2,708	115,839	SH		SOLE		115,839	0	0
EVERBRIDGE, INC.	COM	29978A104	16,435	154,525	SH		SOLE		154,525	0	0
EXELIXIS, INC.	COM	30161Q104	4,346	252,359	SH		SOLE		252,359	0	0
FACEBOOK, INC.	COM	30303M102	8,132	48,755	SH		SOLE		48,755	0	0
GENMARK DIAGNOSTICS, INC.	COM	372309104	3,680	893,090	SH		SOLE		893,090	0	0
GILEAD SCIENCES, INC.	COM	375558103	14,977	200,338	SH		SOLE		200,338	0	0
GLU MOBILE, INC.	COM	379890106	5,994	952,979	SH		SOLE		952,979	0	0
GODADDY, INC.	COM	380237107	9,751	170,734	SH		SOLE		170,734	0	0
GRUBHUB, INC.	COM	400110102	2,227	54,674	SH		SOLE		54,674	0	0
GW PHARMACEUTICALS PLC	COM	36197T103	3,626	41,410	SH		SOLE		41,410	0	0
HAEMONETICS CORPORATION	COM	405024100	2,603	26,122	SH		SOLE		26,122	0	0
HALOZYME THERAPEUTICS, INC.	COM	40637H109	5,564	309,270	SH		SOLE		309,270	0	0
HILL-ROM HOLDINGS, INC.	COM	431475102	7,466	74,212	SH		SOLE		74,212	0	0
HYRECAR, INC.	COM	44916T107	792	582,714	SH		SOLE		582,714	0	0
IAC/INTERACTIVECORP	COM	44919P508	3,052	17,028	SH		SOLE		17,028	0	0
ICAD, INC.	COM	44934S206	1,181	160,876	SH		SOLE		160,876	0	0
ICU MEDICAL, INC.	COM	44930G107	3,938	19,517	SH		SOLE		19,517	0	0
J2 GLOBAL, INC.	COM	48123V102	9,898	132,239	SH		SOLE		132,239	0	0
JOYY, INC.	COM	46591M109	1,684	31,620	SH		SOLE		31,620	0	0
K12, INC.	COM	48273U102	9,419	499,398	SH		SOLE		499,398	0	0
KROGER COMPANY	COM	501044101	5,427	180,163	SH		SOLE		180,163	0	0
KVH INDUSTRIES, INC.	COM	482738101	3,909	414,565	SH		SOLE		414,565	0	0
LENDINGTREE, INC.	COM	52603B107	1,196	6,519	SH		SOLE		6,519	0	0
LIVEPERSON, INC.	COM	538146101	18,785	825,710	SH		SOLE		825,710	0	0
LYFT, INC.	COM	55087P104	1,787	66,546	SH		SOLE		66,546	0	0
MATCH GROUP, INC.	COM	57665R106	7,600	115,084	SH		SOLE		115,084	0	0
MICROSOFT CORPORATION	COM	594918104	7,672	48,649	SH		SOLE		48,649	0	0
MIMECAST LTD.	COM	G14838109	14,754	417,963	SH		SOLE		417,963	0	0
MODEL N, INC.	COM	607525102	5,732	258,079	SH		SOLE		258,079	0	0
NOKIA CORPORATION	COM	654902204	1,859	599,805	SH		SOLE		599,805	0	0
NOMAD FOODS LTD.	COM	G6564A105	15,863	854,686	SH		SOLE		854,686	0	0
OLLIE'S BARGAIN OUTLET HOLDINGS, INC.	COM	681116109	1,605	34,636	SH		SOLE		34,636	0	0
ONEMAIN HOLDINGS, INC.	COM	68268W103	2,199	115,002	SH		SOLE		115,002	0	0
OOMA, INC.	COM	683416101	1,571	131,676	SH		SOLE		131,676	0	0
PFIZER, INC.	COM	717081103	8,122	248,839	SH		SOLE		248,839	0	0
PREMIER, INC.	COM	74051N102	5,491	167,831	SH		SOLE		167,831	0	0
PRIMO WATER CORPORATION	COM	74167P108	2,437	268,990	SH		SOLE		268,990	0	0
PROLOGIS, INC.	COM	74340W103	2,441	30,367	SH		SOLE		30,367	0	0
QUALYS, INC.	COM	74758T303	17,057	196,080	SH		SOLE		196,080	0	0
QUANTERIX CORPORATION	COM	74766Q101	6,338	345,000	SH		SOLE		345,000	0	0
QUANTUM CORPORATION	COM	747906501	1,851	627,402	SH		SOLE		627,402	0	0
QUIDEL CORPORATION	COM	74838J101	3,777	38,614	SH		SOLE		38,614	0	0
QUINSTREET, INC.	COM	74874Q100	7,543	937,078	SH		SOLE		937,078	0	0
QUOTIENT TECHNOLOGY, INC.	COM	749119103	3,294	506,734	SH		SOLE		506,734	0	0
RAPID7, INC.	COM	753422104	11,774	271,733	SH		SOLE		271,733	0	0
REALPAGE, INC.	COM	75606N109	14,901	281,527	SH		SOLE		281,527	0	0
REX AMERICAN RESOURCES CORPORATION	COM	761624105	479	10,295	SH		SOLE		10,295	0	0
RUBICON PROJECT, INC. (THE)	COM	78112V102	1,568	282,542	SH		SOLE		282,542	0	0
SCIPLAY CORPORATION	COM	809087109	3,755	394,278	SH		SOLE		394,278	0	0
SKECHERS USA, INC.	COM	830566105	3,720	156,694	SH		SOLE		156,694	0	0
SKYLINE CHAMPION CORPORATION	COM	830830105	2,519	160,639	SH		SOLE		160,639	0	0
SLM CORPORATION	COM	78442P106	3,229	449,031	SH		SOLE		449,031	0	0
SPARTAN MOTORS, INC.	COM	846819100	5,274	408,511	SH		SOLE		408,511	0	0
SPORTSMANS WAREHOUSE HOLDINGS, INC.	COM	84920Y106	1,597	259,219	SH		SOLE		259,219	0	0
SPROUT SOCIAL, INC.	COM	85209W109	3,465	217,101	SH		SOLE		217,101	0	0
SYROS PHARMACEUTICALS, INC.	COM	87184Q107	3,040	512,570	SH		SOLE		512,570	0	0
TARGET CORPORATION	COM	87612E106	3,881	41,740	SH		SOLE		41,740	0	0
TECHTARGET, INC.	COM	87874R100	2,534	122,964	SH		SOLE		122,964	0	0
TREEHOUSE FOODS, INC.	COM	89469A104	3,205	72,598	SH		SOLE		72,598	0	0
TWILIO, INC.	COM	90138F102	9,597	107,244	SH		SOLE		107,244	0	0
TWITTER, INC.	COM	90184L102	4,012	163,367	SH		SOLE		163,367	0	0
UBER TECHNOLOGIES, INC.	COM	90353T100	3,250	116,406	SH		SOLE		116,406	0	0
ULTA BEAUTY, INC.	COM	90384S303	634	3,606	SH		SOLE		3,606	0	0
UNISYS CORPORATION	COM	909214306	4,449	360,222	SH		SOLE		360,222	0	0
UNITED THERAPEUTICS CORPORATION	COM	91307C102	10,884	114,777	SH		SOLE		114,777	0	0
UPLAND SOFTWARE, INC.	COM	91544A109	26,865	1,001,666	SH		SOLE		1,001,666	0	0
VICOR CORPORATION	COM	925815102	1,530	34,358	SH		SOLE		34,358	0	0
VONAGE HOLDINGS CORP.	COM	92886T201	4,877	674,502	SH		SOLE		674,502	0	0
WESTERN DIGITAL CORPORATION	COM	958102105	4,778	114,809	SH		SOLE		114,809	0	0
ZIX CORPORATION	COM	98974P100	8,780	2,037,118	SH		SOLE		2,037,118	0	0
ZYNERBA PHARMACEUTICALS, INC.	COM	98986X109	1,665	434,768	SH		SOLE		434,768	0	0
ZYNGA, INC.	COM	98986T108	5,453	796,089	SH		SOLE		796,089	0	0