Portolan Capital Management 13F-HR Quarterly holdings report by institutional manager

The Securities and Exchange Commission has not necessarily reviewed the information in this filing and has not determined if it is accurate and complete.
The reader should not assume that the information is accurate and complete.

UNITED STATES SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549
FORM 13F

FORM 13F INFORMATION TABLE

OMB APPROVAL

OMB Number:		3235-0006
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COLUMN 1	COLUMN 2	COLUMN 3	COLUMN 4	COLUMN 5			COLUMN 6	COLUMN 7	COLUMN 8
			VALUE	SHRS OR	SH/	PUT/	INVESTMENT	OTHER	VOTING AUTHORITY
NAME OF ISSUER	TITLE OF CLASS	CUSIP	(x$1000)	PRN AMT	PRN	CALL	DISCRETION	MANAGER	SOLE	SHARED	NONE
AARON'S, INC.	COM	002535300	11,174	246,133	SH		SOLE		246,133	0	0
ABIOMED, INC.	COM	003654100	2,025	8,385	SH		SOLE		8,385	0	0
ACTIVISION BLIZZARD, INC.	COM	00507V109	6,574	86,609	SH		SOLE		86,609	0	0
AKAMAI TECHNOLOGIES, INC.	COM	00971T101	5,196	48,516	SH		SOLE		48,516	0	0
ALLOT LTD .	COM	M0854Q105	3,255	310,877	SH		SOLE		310,877	0	0
ALPHABET, INC.	COM	02079K107	12,280	8,687	SH		SOLE		8,687	0	0
ALTA EQUIPMENT GROUP, INC.	COM	02128L106	8,783	1,130,410	SH		SOLE		1,130,410	0	0
AMAZON.COM, INC.	COM	023135106	21,519	7,800	SH		SOLE		7,800	0	0
AMERICAN PUBLIC EDUCATION, INC.	COM	02913V103	6,575	222,135	SH		SOLE		222,135	0	0
AMERICAS CAR MART, INC.	COM	03062T105	18,829	214,284	SH		SOLE		214,284	0	0
ANGI HOMESERVICES, INC.	COM	00183L102	12,369	1,018,020	SH		SOLE		1,018,020	0	0
ANIKA THERAPEUTICS, INC.	COM	035255108	370	9,814	SH		SOLE		9,814	0	0
APPLIED GENETIC TECHNOLOGIES CORPORATION	COM	03820J100	4,354	785,987	SH		SOLE		785,987	0	0
AVID BIOSERVICES, INC.	COM	05368M106	2,854	434,712	SH		SOLE		434,712	0	0
AXT, INC.	COM	00246W103	3,471	729,142	SH		SOLE		729,142	0	0
BANDWIDTH, INC.	COM	05988J103	30,086	236,897	SH		SOLE		236,897	0	0
BEYOND AIR, INC.	COM	08862L103	2,073	285,970	SH		SOLE		285,970	0	0
BIO-RAD LABORATORIES, INC.	COM	090572207	16,315	36,135	SH		SOLE		36,135	0	0
CACI INTERNATIONAL, INC.	COM	127190304	5,708	26,321	SH		SOLE		26,321	0	0
CALLAWAY GOLF CO.	COM	131193104	4,440	253,580	SH		SOLE		253,580	0	0
CARROLS RESTAURANT GROUP, INC.	COM	14574X104	4,147	856,873	SH		SOLE		856,873	0	0
CASELLA WASTE SYSTEMS, INC.	COM	147448104	5,375	103,124	SH		SOLE		103,124	0	0
CASTLE BIOSCIENCES, INC.	COM	14843C105	2,707	71,834	SH		SOLE		71,834	0	0
CHARAH SOLUTIONS, INC.	COM	15957P105	1,588	499,511	SH		SOLE		499,511	0	0
CHEGG, INC.	COM	163092109	8,990	133,661	SH		SOLE		133,661	0	0
CHEMOURS COMPANY	COM	163851108	5,974	389,181	SH		SOLE		389,181	0	0
CHINA ONLINE EDUCATION GROUP	COM	16954L105	3,199	124,567	SH		SOLE		124,567	0	0
CISCO SYSTEMS, INC.	COM	17275R102	2,780	59,599	SH		SOLE		59,599	0	0
CLOUDERA, INC.	COM	18914U100	12,322	968,673	SH		SOLE		968,673	0	0
COEUR MINING, INC.	COM	192108504	4,250	836,614	SH		SOLE		836,614	0	0
CONTRAFECT CORPORATION	COM	212326300	1,966	307,723	SH		SOLE		307,723	0	0
CROCS, INC.	COM	227046109	14,041	381,338	SH		SOLE		381,338	0	0
DASAN ZHONE SOLUTIONS, INC.	COM	23305L206	2,456	275,004	SH		SOLE		275,004	0	0
DASEKE, INC.	COM	23753F107	956	243,142	SH		SOLE		243,142	0	0
DICKS SPORTING GOODS, INC.	COM	253393102	13,649	330,797	SH		SOLE		330,797	0	0
DOLLAR GENERAL CORPORATION	COM	256677105	5,582	29,301	SH		SOLE		29,301	0	0
DOLLAR TREE STORES, INC.	COM	256746108	2,579	27,831	SH		SOLE		27,831	0	0
DOMO, INC.	COM	257554105	11,483	356,943	SH		SOLE		356,943	0	0
E.L.F. BEAUTY, INC.	COM	26856L103	17,561	920,888	SH		SOLE		920,888	0	0
EBIX, INC.	COM	278715206	751	33,594	SH		SOLE		33,594	0	0
ELECTRONIC ARTS	COM	285512109	9,507	71,996	SH		SOLE		71,996	0	0
EMERGENT BIOSOLUTIONS, INC.	COM	29089Q105	10,380	131,255	SH		SOLE		131,255	0	0
ENCORE CAPITAL GROUP, INC.	COM	292554102	9,157	267,911	SH		SOLE		267,911	0	0
EQT CORPORATION	COM	26884L109	4,462	374,918	SH		SOLE		374,918	0	0
EROS INTERNATIONAL PLC	COM	G3788M114	6,629	2,097,661	SH		SOLE		2,097,661	0	0
ETSY, INC.	COM	29786A106	1,713	16,123	SH		SOLE		16,123	0	0
EVERBRIDGE, INC.	COM	29978A104	20,822	150,488	SH		SOLE		150,488	0	0
EXELIXIS, INC.	COM	30161Q104	3,913	164,809	SH		SOLE		164,809	0	0
F5 NETWORKS, INC.	COM	315616102	6,706	48,076	SH		SOLE		48,076	0	0
FACEBOOK, INC.	COM	30303M102	7,257	31,960	SH		SOLE		31,960	0	0
FLEXSTEEL INDUSTRIES, INC.	COM	339382103	2,805	222,059	SH		SOLE		222,059	0	0
FLIR SYSTEMS, INC.	COM	302445101	5,624	138,613	SH		SOLE		138,613	0	0
FRONT YARD RESIDENTIAL CORPORATION	COM	35904G107	3,663	421,058	SH		SOLE		421,058	0	0
GENMARK DIAGNOSTICS, INC.	COM	372309104	14,585	991,497	SH		SOLE		991,497	0	0
GILEAD SCIENCES, INC.	COM	375558103	2,969	38,592	SH		SOLE		38,592	0	0
GLU MOBILE, INC.	COM	379890106	3,827	412,875	SH		SOLE		412,875	0	0
GODADDY, INC.	COM	380237107	10,621	144,837	SH		SOLE		144,837	0	0
GOODRICH PETROLEUM CORPORATION	COM	382410843	78	10,824	SH		SOLE		10,824	0	0
GW PHARMACEUTICALS PLC	COM	36197T103	5,446	44,377	SH		SOLE		44,377	0	0
H&E EQUIPMENT SERVICES, INC.	COM	404030108	6,439	348,442	SH		SOLE		348,442	0	0
HAEMONETICS CORPORATION	COM	405024100	2,395	26,747	SH		SOLE		26,747	0	0
HALOZYME THERAPEUTICS, INC.	COM	40637H109	14,298	533,310	SH		SOLE		533,310	0	0
HARVARD BIOSCIENCE, INC.	COM	416906105	1,796	579,294	SH		SOLE		579,294	0	0
HD SUPPLY HOLDINGS, INC.	COM	40416M105	2,424	69,971	SH		SOLE		69,971	0	0
HILL-ROM HOLDINGS, INC.	COM	431475102	934	8,509	SH		SOLE		8,509	0	0
HYRECAR, INC.	COM	44916T107	478	163,144	SH		SOLE		163,144	0	0
IAC/INTERACTIVECORP	COM	44919P508	7,161	22,142	SH		SOLE		22,142	0	0
ICAD, INC.	COM	44934S206	2,523	252,514	SH		SOLE		252,514	0	0
ICF INTERNATIONAL, INC.	COM	44925C103	3,340	51,518	SH		SOLE		51,518	0	0
JOYY, INC.	COM	46591M109	3,226	36,430	SH		SOLE		36,430	0	0
K12, INC.	COM	48273U102	26,983	990,559	SH		SOLE		990,559	0	0
KINROSS GOLD CORPORATION	COM	496902404	6,023	834,146	SH		SOLE		834,146	0	0
KROGER COMPANY	COM	501044101	6,973	206,005	SH		SOLE		206,005	0	0
KVH INDUSTRIES, INC.	COM	482738101	2,946	329,882	SH		SOLE		329,882	0	0
LENDINGTREE, INC.	COM	52603B107	1,750	6,046	SH		SOLE		6,046	0	0
LIVEPERSON, INC.	COM	538146101	10,970	264,787	SH		SOLE		264,787	0	0
MATCH GROUP, INC.	COM	57665R106	5,207	48,644	SH		SOLE		48,644	0	0
MAXLINEAR, INC.	COM	57776J100	5,065	236,029	SH		SOLE		236,029	0	0
MICROSOFT CORPORATION	COM	594918104	9,847	48,386	SH		SOLE		48,386	0	0
MIMECAST LIMITED	COM	G14838109	6,856	164,579	SH		SOLE		164,579	0	0
MOBILEIRON, INC.	COM	60739U204	3,320	673,338	SH		SOLE		673,338	0	0
MODEL N, INC.	COM	607525102	6,851	197,101	SH		SOLE		197,101	0	0
NAVIENT CORPORATION	COM	63938C108	6,644	945,084	SH		SOLE		945,084	0	0
NOKIA CORPORATION	COM	654902204	14,040	3,190,900	SH		SOLE		3,190,900	0	0
NOMAD FOODS LTD.	COM	G6564A105	12,291	573,002	SH		SOLE		573,002	0	0
NU SKIN ENTERPRISES, INC.	COM	67018T105	6,184	161,769	SH		SOLE		161,769	0	0
OLLIE'S BARGAIN OUTLET HOLDINGS, INC.	COM	681116109	2,682	27,464	SH		SOLE		27,464	0	0
ONEMAIN HOLDINGS, INC.	COM	68268W103	3,369	137,300	SH		SOLE		137,300	0	0
OPEN LENDING CORPORATION	COM	68373J104	15,840	1,049,006	SH		SOLE		1,049,006	0	0
ORION ENGINEERED CARBONS S.A	COM	L72967109	2,795	263,965	SH		SOLE		263,965	0	0
OVERSTOCK.COM, INC.	COM	690370101	8,772	308,555	SH		SOLE		308,555	0	0
PAYPAL HOLDINGS, INC.	COM	70450Y103	6,016	34,527	SH		SOLE		34,527	0	0
PFIZER, INC.	COM	717081103	5,111	156,310	SH		SOLE		156,310	0	0
PFSWEB, INC.	COM	717098206	1,887	282,432	SH		SOLE		282,432	0	0
PREMIER, INC.	COM	74051N102	4,261	124,295	SH		SOLE		124,295	0	0
QUALYS, INC.	COM	74758T303	6,861	65,958	SH		SOLE		65,958	0	0
QUANTERIX CORPORATION	COM	74766Q101	4,073	148,719	SH		SOLE		148,719	0	0
QUANTUM CORPORATION	COM	747906501	1,504	389,688	SH		SOLE		389,688	0	0
QUIDEL CORPORATION	COM	74838J101	5,385	24,070	SH		SOLE		24,070	0	0
QUINSTREET, INC.	COM	74874Q100	14,815	1,416,346	SH		SOLE		1,416,346	0	0
RAPID7, INC.	COM	753422104	4,162	81,573	SH		SOLE		81,573	0	0
REALPAGE, INC.	COM	75606N109	6,681	102,770	SH		SOLE		102,770	0	0
RECRO PHARMA, INC.	COM	75629F109	835	183,554	SH		SOLE		183,554	0	0
RH	COM	74967X103	5,378	21,606	SH		SOLE		21,606	0	0
RUBICON PROJECT, INC. (THE)	COM	78112V102	526	78,787	SH		SOLE		78,787	0	0
SHYFT GROUP, INC. (THE)	COM	825698103	5,232	310,701	SH		SOLE		310,701	0	0
SKECHERS USA, INC.	COM	830566105	10,664	339,826	SH		SOLE		339,826	0	0
SKYLINE CHAMPION CORPORATION	COM	830830105	6,923	284,430	SH		SOLE		284,430	0	0
SONOS, INC.	COM	83570H108	7,654	523,204	SH		SOLE		523,204	0	0
SPORTSMAN'S WAREHOUSE HOLDINGS, INC.	COM	84920Y106	20,734	1,455,047	SH		SOLE		1,455,047	0	0
SPROUT SOCIAL, INC.	COM	85209W109	3,006	111,329	SH		SOLE		111,329	0	0
SUNOPTA, INC.	COM	8676EP108	6,344	1,349,825	SH		SOLE		1,349,825	0	0
SYNCHRONY FINANCIAL	COM	87165B103	6,239	281,541	SH		SOLE		281,541	0	0
SYROS PHARMACEUTICALS, INC.	COM	87184Q107	5,316	498,666	SH		SOLE		498,666	0	0
TARGET CORPORATION	COM	87612E106	6,078	50,678	SH		SOLE		50,678	0	0
TREEHOUSE FOODS, INC.	COM	89469A104	11,024	251,685	SH		SOLE		251,685	0	0
TWILIO, INC.	COM	90138F102	11,563	52,699	SH		SOLE		52,699	0	0
TWITTER, INC.	COM	90184L102	1,068	35,866	SH		SOLE		35,866	0	0
UNISYS CORPORATION	COM	909214306	485	44,463	SH		SOLE		44,463	0	0
UNITED THERAPEUTICS CORPORATION	COM	91307C102	9,080	75,039	SH		SOLE		75,039	0	0
UPLAND SOFTWARE, INC.	COM	91544A109	21,636	622,447	SH		SOLE		622,447	0	0
VICOR CORPORATION	COM	925815102	1,451	20,165	SH		SOLE		20,165	0	0
VONAGE HOLDINGS CORP.	COM	92886T201	7,085	704,273	SH		SOLE		704,273	0	0
WAITR HOLDINGS, INC.	COM	930752100	5,393	2,050,522	SH		SOLE		2,050,522	0	0
WESTERN DIGITAL CORPORATION	COM	958102105	2,420	54,810	SH		SOLE		54,810	0	0
WILLIAMS-SONOMA, INC.	COM	969904101	4,004	48,824	SH		SOLE		48,824	0	0
ZIX CORPORATION	COM	98974P100	4,119	597,026	SH		SOLE		597,026	0	0
ZYNGA, INC.	COM	98986T108	9,412	986,552	SH		SOLE		986,552	0	0