Portolan Capital Management 13F-HR Quarterly holdings report by institutional manager

The Securities and Exchange Commission has not necessarily reviewed the information in this filing and has not determined if it is accurate and complete.
The reader should not assume that the information is accurate and complete.

UNITED STATES SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549
FORM 13F

FORM 13F INFORMATION TABLE

OMB APPROVAL

OMB Number:		3235-0006
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COLUMN 1	COLUMN 2	COLUMN 3	COLUMN 4	COLUMN 5			COLUMN 6	COLUMN 7	COLUMN 8
			VALUE	SHRS OR	SH/	PUT/	INVESTMENT	OTHER	VOTING AUTHORITY
NAME OF ISSUER	TITLE OF CLASS	CUSIP	(x$1000)	PRN AMT	PRN	CALL	DISCRETION	MANAGER	SOLE	SHARED	NONE
2U, INC.	COM	90214J101	2,030	59,939	SH		SOLE		59,939	0	0
AARON'S, INC.	COM	002535300	13,519	238,634	SH		SOLE		238,634	0	0
ABIOMED, INC.	COM	003654100	2,127	7,677	SH		SOLE		7,677	0	0
ACTIVISION BLIZZARD, INC.	COM	00507V109	4,687	57,898	SH		SOLE		57,898	0	0
AKAMAI TECHNOLOGIES, INC.	COM	00971T101	1,186	10,725	SH		SOLE		10,725	0	0
ALPHABET, INC.	COM	02079K107	10,264	6,984	SH		SOLE		6,984	0	0
ALTA EQUIPMENT GROUP, INC.	COM	02128L106	8,777	1,120,898	SH		SOLE		1,120,898	0	0
AMAZON.COM, INC.	COM	023135106	23,877	7,583	SH		SOLE		7,583	0	0
AMBARELLA, INC.	COM	G037AX101	6,274	120,235	SH		SOLE		120,235	0	0
AMERICAN PUBLIC EDUCATION, INC.	COM	02913V103	9,991	354,424	SH		SOLE		354,424	0	0
AMERICAS CAR MART, INC.	COM	03062T105	18,644	219,653	SH		SOLE		219,653	0	0
ANGI HOMESERVICES, INC.	COM	00183L102	518	46,674	SH		SOLE		46,674	0	0
ANIKA THERAPEUTICS, INC.	COM	035255108	273	7,708	SH		SOLE		7,708	0	0
APPLIED GENETIC TECHNOLOGIES CORPORATION	COM	03820J100	6,323	1,301,075	SH		SOLE		1,301,075	0	0
AVID BIOSERVICES, INC.	COM	05368M106	7,341	963,374	SH		SOLE		963,374	0	0
AVIS BUDGET GROUP, INC.	COM	053774105	1,517	57,631	SH		SOLE		57,631	0	0
AXT, INC.	COM	00246W103	5,670	926,413	SH		SOLE		926,413	0	0
BANDWIDTH, INC.	COM	05988J103	21,772	124,718	SH		SOLE		124,718	0	0
BIO-RAD LABORATORIES, INC.	COM	090572207	9,205	17,857	SH		SOLE		17,857	0	0
CAI INTERNATIONAL, INC.	COM	12477X106	4,592	166,804	SH		SOLE		166,804	0	0
CALLAWAY GOLF CO.	COM	131193104	2,666	139,312	SH		SOLE		139,312	0	0
CARPARTS.COM, INC.	COM	14427M107	6,720	621,690	SH		SOLE		621,690	0	0
CARROLS RESTAURANT GROUP, INC.	COM	14574X104	11,677	1,810,338	SH		SOLE		1,810,338	0	0
CASELLA WASTE SYSTEMS, INC.	COM	147448104	5,885	105,373	SH		SOLE		105,373	0	0
CASTLE BIOSCIENCES, INC.	COM	14843C105	1,422	27,632	SH		SOLE		27,632	0	0
CBRE GROUP, INC.	COM	12504L109	6,653	141,635	SH		SOLE		141,635	0	0
CHARAH SOLUTIONS, INC.	COM	15957P105	9,150	2,980,339	SH		SOLE		2,980,339	0	0
CHEGG, INC.	COM	163092109	2,086	29,200	SH		SOLE		29,200	0	0
CHEMOURS COMPANY	COM	163851108	6,432	307,625	SH		SOLE		307,625	0	0
CLOUDERA, INC.	COM	18914U100	11,864	1,089,419	SH		SOLE		1,089,419	0	0
COEUR MINING, INC.	COM	192108504	7,229	979,513	SH		SOLE		979,513	0	0
COMMERCIAL VEHICLE GROUP, INC.	COM	202608105	4,884	747,961	SH		SOLE		747,961	0	0
COMSTOCK RESOURCES, INC.	COM	205768302	7,986	1,823,247	SH		SOLE		1,823,247	0	0
CONTRAFECT CORPORATION	COM	212326300	1,441	273,006	SH		SOLE		273,006	0	0
COSTAMARE, INC.	COM	Y1771G102	1,883	310,173	SH		SOLE		310,173	0	0
CROCS, INC.	COM	227046109	13,056	305,542	SH		SOLE		305,542	0	0
DASEKE, INC.	COM	23753F107	5,633	1,049,053	SH		SOLE		1,049,053	0	0
DICKS SPORTING GOODS, INC.	COM	253393102	5,372	92,817	SH		SOLE		92,817	0	0
DOLLAR GENERAL CORPORATION	COM	256677105	5,556	26,505	SH		SOLE		26,505	0	0
DOMO, INC.	COM	257554105	10,651	277,888	SH		SOLE		277,888	0	0
DZS, INC.	COM	268211109	3,056	326,143	SH		SOLE		326,143	0	0
E.L.F. BEAUTY, INC.	COM	26856L103	18,768	1,021,654	SH		SOLE		1,021,654	0	0
EBAY, INC.	COM	278642103	3,037	58,294	SH		SOLE		58,294	0	0
ELECTRONIC ARTS	COM	285512109	4,765	36,535	SH		SOLE		36,535	0	0
EMERGENT BIOSOLUTIONS, INC.	COM	29089Q105	5,911	57,203	SH		SOLE		57,203	0	0
ENCORE CAPITAL GROUP, INC.	COM	292554102	9,998	259,084	SH		SOLE		259,084	0	0
ENOVA INTERNATIONAL, INC.	COM	29357K103	9,819	599,067	SH		SOLE		599,067	0	0
EQT CORPORATION	COM	26884L109	7,482	578,680	SH		SOLE		578,680	0	0
EROS STX GLOBAL CORPORATION	COM	G3788M114	7,969	3,605,881	SH		SOLE		3,605,881	0	0
ETSY, INC.	COM	29786A106	778	6,398	SH		SOLE		6,398	0	0
EVERBRIDGE, INC.	COM	29978A104	7,178	57,089	SH		SOLE		57,089	0	0
EXELIXIS, INC.	COM	30161Q104	3,574	146,168	SH		SOLE		146,168	0	0
FACEBOOK, INC.	COM	30303M102	7,596	29,002	SH		SOLE		29,002	0	0
FLEXION THERAPEUTICS, INC.	COM	33938J106	2,948	283,169	SH		SOLE		283,169	0	0
FLEXSTEEL INDUSTRIES, INC.	COM	339382103	3,635	151,452	SH		SOLE		151,452	0	0
FRANCHISE GROUP, INC.	COM	35180X105	5,605	221,020	SH		SOLE		221,020	0	0
GAMESTOP CORP.	COM	36467W109	6,501	637,318	SH		SOLE		637,318	0	0
GENMARK DIAGNOSTICS, INC.	COM	372309104	3,109	218,924	SH		SOLE		218,924	0	0
GIBRALTAR INDUSTRIES, INC.	COM	374689107	2,752	42,243	SH		SOLE		42,243	0	0
GLU MOBILE, INC.	COM	379890106	2,761	359,785	SH		SOLE		359,785	0	0
GREEN PLAINS, INC.	COM	393222104	4,540	293,301	SH		SOLE		293,301	0	0
GRIFFON CORPORATION	COM	398433102	3,835	196,268	SH		SOLE		196,268	0	0
GW PHARMACEUTICALS PLC	COM	36197T103	4,584	47,091	SH		SOLE		47,091	0	0
H&E EQUIPMENT SERVICES, INC.	COM	404030108	7,978	405,823	SH		SOLE		405,823	0	0
HALOZYME THERAPEUTICS, INC.	COM	40637H109	10,668	405,934	SH		SOLE		405,934	0	0
HARVARD BIOSCIENCE, INC.	COM	416906105	2,947	979,067	SH		SOLE		979,067	0	0
HOUGHTON MIFFLIN HARCOURT COMPANY	COM	44157R109	2,105	1,216,584	SH		SOLE		1,216,584	0	0
HYRECAR, INC.	COM	44916T107	465	147,728	SH		SOLE		147,728	0	0
IAC/INTERACTIVECORP.	COM	44891N109	6,411	53,522	SH		SOLE		53,522	0	0
ICAD, INC.	COM	44934S206	4,299	487,917	SH		SOLE		487,917	0	0
IDENTIV, INC.	COM	45170X205	3,625	579,073	SH		SOLE		579,073	0	0
IMMERSION CORPORATION	COM	452521107	2,959	419,723	SH		SOLE		419,723	0	0
IMPINJ, INC.	COM	453204109	4,521	171,574	SH		SOLE		171,574	0	0
INSURANCE ACQUISITION CORP.	COM	457867109	1,174	99,435	SH		SOLE		99,435	0	0
JOYY, INC.	COM	46591M109	6,760	83,804	SH		SOLE		83,804	0	0
K12, INC.	COM	48273U102	8,710	330,674	SH		SOLE		330,674	0	0
KINROSS GOLD CORPORATION	COM	496902404	5,004	567,387	SH		SOLE		567,387	0	0
LENDINGTREE, INC.	COM	52603B107	1,198	3,903	SH		SOLE		3,903	0	0
LGI HOMES, INC.	COM	50187T106	7,113	61,229	SH		SOLE		61,229	0	0
LIVEPERSON, INC.	COM	538146101	12,525	240,904	SH		SOLE		240,904	0	0
MATCH GROUP, INC.	COM	57667L107	8,996	81,301	SH		SOLE		81,301	0	0
MAXLINEAR, INC.	COM	57776J100	5,641	242,742	SH		SOLE		242,742	0	0
MERCER INTERNATIONAL, INC.	COM	588056101	2,282	345,822	SH		SOLE		345,822	0	0
MGIC INVESTMENT CORP.	COM	552848103	2,411	272,098	SH		SOLE		272,098	0	0
MICROSOFT CORPORATION	COM	594918104	8,238	39,168	SH		SOLE		39,168	0	0
NAVIENT CORPORATION	COM	63938C108	19,077	2,257,635	SH		SOLE		2,257,635	0	0
NEW HOME CO LLC/THE	COM	645370107	2,171	399,110	SH		SOLE		399,110	0	0
NOKIA CORPORATION	COM	654902204	4,216	1,078,202	SH		SOLE		1,078,202	0	0
NOMAD FOODS LTD.	COM	G6564A105	5,483	215,197	SH		SOLE		215,197	0	0
NU SKIN ENTERPRISES, INC.	COM	67018T105	12,489	249,337	SH		SOLE		249,337	0	0
OLLIE'S BARGAIN OUTLET HOLDINGS, INC.	COM	681116109	2,123	24,305	SH		SOLE		24,305	0	0
ONEMAIN HOLDINGS, INC.	COM	68268W103	9,748	311,935	SH		SOLE		311,935	0	0
OPEN LENDING CORPORATION	COM	68373J104	28,830	1,130,588	SH		SOLE		1,130,588	0	0
ORION ENGINEERED CARBONS S.A	COM	L72967109	5,416	432,911	SH		SOLE		432,911	0	0
OVERSTOCK.COM, INC.	COM	690370101	1,862	25,623	SH		SOLE		25,623	0	0
PACIFIC ETHANOL, INC.	COM	69423U305	10,069	1,377,475	SH		SOLE		1,377,475	0	0
PAYPAL HOLDINGS, INC.	COM	70450Y103	2,152	10,922	SH		SOLE		10,922	0	0
PFSWEB, INC.	COM	717098206	7,670	1,146,455	SH		SOLE		1,146,455	0	0
PROPTECH ACQUISITION CORP.	COM	74349F101	7,116	639,372	SH		SOLE		639,372	0	0
PROPTECH ACQUISITION CORP.	WTS	74349F119	1,185	721,440	SH		SOLE		721,440	0	0
QUALYS, INC.	COM	74758T303	8,282	84,502	SH		SOLE		84,502	0	0
QUANTERIX CORPORATION	COM	74766Q101	1,616	47,900	SH		SOLE		47,900	0	0
QUANTUM CORPORATION	COM	747906501	552	119,953	SH		SOLE		119,953	0	0
QUIDEL CORPORATION	COM	74838J101	1,447	6,598	SH		SOLE		6,598	0	0
QUINSTREET, INC.	COM	74874Q100	24,123	1,522,916	SH		SOLE		1,522,916	0	0
RANPAK HOLDINGS CORP.	COM	75321W103	2,077	218,194	SH		SOLE		218,194	0	0
RAPID7, INC.	COM	753422104	1,501	24,512	SH		SOLE		24,512	0	0
REALPAGE, INC.	COM	75606N109	5,886	102,108	SH		SOLE		102,108	0	0
RECRO PHARMA, INC.	COM	75629F109	481	229,210	SH		SOLE		229,210	0	0
REX AMERICAN RESOURCES CORP.	COM	761624105	4,755	72,480	SH		SOLE		72,480	0	0
RH	COM	74967X103	7,375	19,275	SH		SOLE		19,275	0	0
SHARPSPRING, INC.	COM	820054104	2,602	233,402	SH		SOLE		233,402	0	0
SHYFT GROUP, INC. (THE)	COM	825698103	11,812	625,620	SH		SOLE		625,620	0	0
SKECHERS USA, INC.	COM	830566105	4,847	160,383	SH		SOLE		160,383	0	0
SKYLINE CHAMPION CORPORATION	COM	830830105	8,623	322,120	SH		SOLE		322,120	0	0
SOFTWARE ACQUISITION GROUP, INC.	COM	83406B101	762	76,536	SH		SOLE		76,536	0	0
SONOS, INC.	COM	83570H108	7,437	489,947	SH		SOLE		489,947	0	0
SPORTSMAN'S WAREHOUSE HOLDINGS, INC.	COM	84920Y106	14,701	1,027,303	SH		SOLE		1,027,303	0	0
SPROUT SOCIAL, INC.	COM	85209W109	4,706	122,232	SH		SOLE		122,232	0	0
STAMPS.COM, INC.	COM	852857200	9,837	40,826	SH		SOLE		40,826	0	0
SUNOPTA, INC.	COM	8676EP108	19,221	2,590,407	SH		SOLE		2,590,407	0	0
SYNCHRONY FINANCIAL	COM	87165B103	9,765	373,141	SH		SOLE		373,141	0	0
SYROS PHARMACEUTICALS, INC.	COM	87184Q107	3,986	450,861	SH		SOLE		450,861	0	0
TARGET CORPORATION	COM	87612E106	7,909	50,240	SH		SOLE		50,240	0	0
TAYLOR MORRISON HOME CORP.	COM	87724P106	7,904	321,427	SH		SOLE		321,427	0	0
TPI COMPOSITES, INC.	COM	87266J104	1,921	66,326	SH		SOLE		66,326	0	0
TREEHOUSE FOODS, INC.	COM	89469A104	11,882	293,176	SH		SOLE		293,176	0	0
TRITON INTERNATIONAL LTD.	COM	G9078F107	1,997	49,097	SH		SOLE		49,097	0	0
TWILIO, INC.	COM	90138F102	2,200	8,902	SH		SOLE		8,902	0	0
TWITTER, INC.	COM	90184L102	357	8,019	SH		SOLE		8,019	0	0
UNISYS CORPORATION	COM	909214306	394	36,921	SH		SOLE		36,921	0	0
UPLAND SOFTWARE, INC.	COM	91544A109	17,948	476,077	SH		SOLE		476,077	0	0
VICOR CORPORATION	COM	925815102	521	6,709	SH		SOLE		6,709	0	0
VISTRA CORP.	COM	92840M102	1,820	96,514	SH		SOLE		96,514	0	0
VONAGE HOLDINGS CORP.	COM	92886T201	5,698	556,959	SH		SOLE		556,959	0	0
WAITR HOLDINGS, INC.	COM	930752100	1,533	476,216	SH		SOLE		476,216	0	0
WHOLE EARTH BRANDS, INC.	COM	96684W100	2,766	331,649	SH		SOLE		331,649	0	0
WILLIAMS-SONOMA, INC.	COM	969904101	6,150	68,006	SH		SOLE		68,006	0	0
ZUMIEZ, INC.	COM	989817101	3,730	134,062	SH		SOLE		134,062	0	0