Portolan Capital Management 13F-HR Quarterly holdings report by institutional manager

The Securities and Exchange Commission has not necessarily reviewed the information in this filing and has not determined if it is accurate and complete.
The reader should not assume that the information is accurate and complete.

UNITED STATES SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549
FORM 13F

FORM 13F INFORMATION TABLE

OMB APPROVAL

OMB Number:		3235-0006
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COLUMN 1	COLUMN 2	COLUMN 3	COLUMN 4	COLUMN 5			COLUMN 6	COLUMN 7	COLUMN 8
			VALUE	SHRS OR	SH/	PUT/	INVESTMENT	OTHER	VOTING AUTHORITY
NAME OF ISSUER	TITLE OF CLASS	CUSIP	(x$1000)	PRN AMT	PRN	CALL	DISCRETION	MANAGER	SOLE	SHARED	NONE
ABIOMED, INC.	COM	003654100	2,301	7,070	SH		SOLE		7,070	0	0
ACADEMY SPORTS & OUTDOORS, INC.	COM	00402L107	8,389	209,615	SH		SOLE		209,615	0	0
AEMETIS, INC.	COM	00770K202	1,939	106,090	SH		SOLE		106,090	0	0
ALPHABET, INC.	COM	02079K107	21,546	8,084	SH		SOLE		8,084	0	0
ALTA EQUIPMENT GROUP, INC.	COM	02128L106	21,220	1,545,551	SH		SOLE		1,545,551	0	0
AMAZON.COM, INC.	COM	023135106	23,527	7,162	SH		SOLE		7,162	0	0
AMBARELLA, INC.	COM	G037AX101	1,053	6,763	SH		SOLE		6,763	0	0
AMERICAS CAR MART, INC.	COM	03062T105	11,077	94,855	SH		SOLE		94,855	0	0
APPLIED GENETIC TECHNOLOGIES CORPORATION	COM	03820J100	3,803	1,263,421	SH		SOLE		1,263,421	0	0
ARCBEST CORPORATION	COM	03937C105	10,079	123,255	SH		SOLE		123,255	0	0
ATLAS AIR WORLDWIDE HOLDINGS, INC.	COM	049164205	4,266	52,224	SH		SOLE		52,224	0	0
AVID BIOSERVICES, INC.	COM	05368M106	2,079	96,404	SH		SOLE		96,404	0	0
AVID TECHNOLOGY, INC.	COM	05367P100	14,593	504,595	SH		SOLE		504,595	0	0
B. RILEY FINANCIAL, INC.	COM	05580M108	848	14,371	SH		SOLE		14,371	0	0
BABCOCK & WILCOX ENTERPRISES, INC.	COM	05614L209	5,031	784,903	SH		SOLE		784,903	0	0
BANDWIDTH, INC.	COM	05988J103	17,863	197,860	SH		SOLE		197,860	0	0
BARNES & NOBLE EDUCATION, INC.	COM	06777U101	3,646	364,938	SH		SOLE		364,938	0	0
BIO-RAD LABORATORIES, INC.	COM	090572207	18,606	24,943	SH		SOLE		24,943	0	0
BLUELINX HOLDINGS, INC.	COM	09624H208	17,579	359,631	SH		SOLE		359,631	0	0
BRINK'S COMPANY (THE)	COM	109696104	1,827	28,857	SH		SOLE		28,857	0	0
BROOKS AUTOMATION, INC.	COM	114340102	14,092	137,684	SH		SOLE		137,684	0	0
BUNGE LIMITED	COM	G16962105	2,812	34,579	SH		SOLE		34,579	0	0
CAESARSTONE LTD	COM	M20598104	2,353	189,459	SH		SOLE		189,459	0	0
CAI INTERNATIONAL, INC.	COM	12477X106	2,614	46,755	SH		SOLE		46,755	0	0
CALERES, INC.	COM	129500104	13,286	597,952	SH		SOLE		597,952	0	0
CALLAWAY GOLF	COM	131193104	10,542	381,537	SH		SOLE		381,537	0	0
CAPRI HOLDINGS LIMITED	COM	G1890L107	16,606	343,034	SH		SOLE		343,034	0	0
CARPARTS.COM, INC.	COM	14427M107	10,271	658,004	SH		SOLE		658,004	0	0
CASELLA WASTE SYSTEMS, INC.	COM	147448104	1,308	17,227	SH		SOLE		17,227	0	0
CENTURY COMMUNITIES, INC.	COM	156504300	5,601	91,145	SH		SOLE		91,145	0	0
CHARAH SOLUTIONS, INC.	COM	15957P105	14,662	3,187,475	SH		SOLE		3,187,475	0	0
CHILDREN'S PLACE, INC. (THE)	COM	168905107	3,367	44,739	SH		SOLE		44,739	0	0
CITI TRENDS, INC. CMN	COM	17306X102	5,529	75,778	SH		SOLE		75,778	0	0
CLEVELAND-CLIFFS, INC.	COM	185899101	4,170	210,508	SH		SOLE		210,508	0	0
COMMERCIAL VEHICLE GROUP, INC.	COM	202608105	8,553	904,160	SH		SOLE		904,160	0	0
COOPER-STANDARD HOLDINGS, INC.	COM	21676P103	674	30,751	SH		SOLE		30,751	0	0
CROCS, INC.	COM	227046109	33,775	235,402	SH		SOLE		235,402	0	0
CUSTOM TRUCK ONE SOURCE, INC.	COM	23204X103	8,831	946,495	SH		SOLE		946,495	0	0
CUSTOMERS BANCORP, INC.	COM	23204G100	6,978	162,204	SH		SOLE		162,204	0	0
CYBEROPTICS CORPORATION	COM	232517102	6,529	183,489	SH		SOLE		183,489	0	0
DANAOS CORPORATION	COM	Y1968P121	1,079	13,172	SH		SOLE		13,172	0	0
DARLING INGREDIENTS, INC.	COM	237266101	3,288	45,732	SH		SOLE		45,732	0	0
DASEKE, INC.	COM	23753F107	23,862	2,590,930	SH		SOLE		2,590,930	0	0
DECKERS OUTDOOR CORPORATION	COM	243537107	2,461	6,831	SH		SOLE		6,831	0	0
DENBURY, INC.	COM	24790A101	2,212	31,482	SH		SOLE		31,482	0	0
DESTINATION XL GROUP, INC.	COM	25065K104	11,053	1,805,987	SH		SOLE		1,805,987	0	0
DOMO, INC.	COM	257554105	31,120	368,548	SH		SOLE		368,548	0	0
DYADIC INTERNATIONAL, INC.	COM	26745T101	308	55,422	SH		SOLE		55,422	0	0
DZS, INC.	COM	268211109	3,952	322,370	SH		SOLE		322,370	0	0
E.L.F. BEAUTY, INC.	COM	26856L103	29,318	1,009,226	SH		SOLE		1,009,226	0	0
EASTERN BANKSHARES, INC.	COM	27627N105	1,941	95,624	SH		SOLE		95,624	0	0
ELECTRONIC ARTS	COM	285512109	4,666	32,799	SH		SOLE		32,799	0	0
ENCORE CAPITAL GROUP, INC.	COM	292554102	7,752	157,347	SH		SOLE		157,347	0	0
ENERGY RECOVERY, INC.	COM	29270J100	1,613	84,771	SH		SOLE		84,771	0	0
ENOVA INTERNATIONAL, INC.	COM	29357K103	19,763	572,026	SH		SOLE		572,026	0	0
EURONET WORLDWIDE, INC.	COM	298736109	6,713	52,745	SH		SOLE		52,745	0	0
FACEBOOK, INC.	COM	30303M102	24,213	71,343	SH		SOLE		71,343	0	0
FULL HOUSE RESORTS, INC.	COM	359678109	19,713	1,857,937	SH		SOLE		1,857,937	0	0
GENESCO, INC.	COM	371532102	19,167	332,017	SH		SOLE		332,017	0	0
GREEN PLAINS, INC.	COM	393222104	23,882	731,454	SH		SOLE		731,454	0	0
GTY TECHNOLOGY HOLDINGS, INC.	COM	362409104	11,200	1,489,367	SH		SOLE		1,489,367	0	0
GULFPORT ENERGY CORPORATION	COM	402635502	7,171	87,228	SH		SOLE		87,228	0	0
H & R BLOCK, INC.	COM	093671105	8,964	358,555	SH		SOLE		358,555	0	0
H&E EQUIPMENT SERVICES, INC.	COM	404030108	3,656	105,321	SH		SOLE		105,321	0	0
HARVARD BIOSCIENCE, INC.	COM	416906105	16,459	2,358,010	SH		SOLE		2,358,010	0	0
HCI GROUP, INC.	COM	40416E103	11,284	101,871	SH		SOLE		101,871	0	0
HIMAX TECHNOLOGIES, INC.	COM	43289P106	7,380	691,636	SH		SOLE		691,636	0	0
HUDSON TECHNOLOGIES, INC.	COM	444144109	65	18,305	SH		SOLE		18,305	0	0
HUT 8 MINING CORP.	COM	44812T102	2,008	239,051	SH		SOLE		239,051	0	0
HYRECAR, INC.	COM	44916T107	1,111	130,683	SH		SOLE		130,683	0	0
IAC/INTERACTIVECORP.	COM	44891N208	6,155	47,237	SH		SOLE		47,237	0	0
ICAD, INC.	COM	44934S206	5,376	500,119	SH		SOLE		500,119	0	0
IDENTIV, INC.	COM	45170X205	28,026	1,487,584	SH		SOLE		1,487,584	0	0
IMPINJ, INC.	COM	453204109	5,424	94,941	SH		SOLE		94,941	0	0
INOTIV, INC.	COM	45783Q100	21,809	745,869	SH		SOLE		745,869	0	0
KALEYRA, INC.	COM	483379103	10,313	936,733	SH		SOLE		936,733	0	0
KURA SUSHI USA, INC.	COM	501270102	10,193	233,345	SH		SOLE		233,345	0	0
LAREDO PETROLEUM, INC.	COM	516806205	19,311	238,203	SH		SOLE		238,203	0	0
LGI HOMES, INC.	COM	50187T106	1,744	12,290	SH		SOLE		12,290	0	0
LIVEPERSON, INC.	COM	538146101	13,061	221,564	SH		SOLE		221,564	0	0
MAGNA INTERNATIONAL, INC.	COM	559222401	1,006	13,367	SH		SOLE		13,367	0	0
MATCH GROUP, INC.	COM	57667L107	10,641	67,783	SH		SOLE		67,783	0	0
MAXLINEAR, INC.	COM	57776J100	5,366	108,964	SH		SOLE		108,964	0	0
MGIC INVESTMENT CORPORATION	COM	552848103	7,808	521,908	SH		SOLE		521,908	0	0
MICROSOFT CORPORATION	COM	594918104	12,715	45,102	SH		SOLE		45,102	0	0
MIMECAST LIMITED	COM	G14838109	5,522	86,825	SH		SOLE		86,825	0	0
MKS INSTRUMENTS, INC.	COM	55306N104	2,749	18,215	SH		SOLE		18,215	0	0
MONEYGRAM INTERNATIONAL, INC.	COM	60935Y208	25,827	3,220,273	SH		SOLE		3,220,273	0	0
NAVIENT CORPORATION	COM	63938C108	20,213	1,024,467	SH		SOLE		1,024,467	0	0
NORTHERN OIL AND GAS, INC.	COM	665531307	9,812	458,504	SH		SOLE		458,504	0	0
NU SKIN ENTERPRISES, INC.	COM	67018T105	4,884	120,692	SH		SOLE		120,692	0	0
ONEMAIN HOLDINGS, INC.	COM	68268W103	10,100	182,534	SH		SOLE		182,534	0	0
OPEN LENDING CORPORATION	COM	68373J104	7,153	198,316	SH		SOLE		198,316	0	0
OVERSTOCK.COM, INC.	COM	690370101	7,836	100,561	SH		SOLE		100,561	0	0
PFSWEB, INC.	COM	717098206	18,080	1,401,541	SH		SOLE		1,401,541	0	0
PORCH GROUP, INC.	COM	733245104	24,583	1,390,467	SH		SOLE		1,390,467	0	0
PROG HOLDINGS, INC.	COM	74319R101	19,896	473,590	SH		SOLE		473,590	0	0
PURETECH HEALTH PLC	COM	746237106	2,516	52,421	SH		SOLE		52,421	0	0
QUALYS, INC.	COM	74758T303	2,582	23,205	SH		SOLE		23,205	0	0
QUINSTREET, INC.	COM	74874Q100	20,647	1,175,804	SH		SOLE		1,175,804	0	0
RAFAEL HOLDINGS, INC.	COM	75062E106	3,810	123,980	SH		SOLE		123,980	0	0
RANPAK HOLDINGS CORPORATION	COM	75321W103	3,502	130,557	SH		SOLE		130,557	0	0
RECRO PHARMA, INC.	COM	75629F109	332	161,253	SH		SOLE		161,253	0	0
RENT-A-CENTER, INC.	COM	76009N100	12,888	229,289	SH		SOLE		229,289	0	0
REX AMERICAN RESOURCES CORPORATION	COM	761624105	4,667	58,435	SH		SOLE		58,435	0	0
RH	COM	74967X103	4,404	6,604	SH		SOLE		6,604	0	0
RIMINI STREET, INC.	COM	76674Q107	7,232	749,478	SH		SOLE		749,478	0	0
ROCKY BRANDS, INC.	COM	774515100	4,512	94,770	SH		SOLE		94,770	0	0
SANDRIDGE ENERGY, INC.	COM	80007P869	22,994	1,767,394	SH		SOLE		1,767,394	0	0
SEMLER SCIENTIFIC, INC.	COM	81684M104	10,279	82,212	SH		SOLE		82,212	0	0
SHYFT GROUP, INC. (THE)	COM	825698103	2,874	75,602	SH		SOLE		75,602	0	0
SKECHERS USA, INC.	COM	830566105	16,636	394,959	SH		SOLE		394,959	0	0
SKYLINE CHAMPION CORPORATION	COM	830830105	18,809	313,167	SH		SOLE		313,167	0	0
SONOS, INC.	COM	83570H108	18,697	577,788	SH		SOLE		577,788	0	0
SONY GROUP CORPORATION	COM	835699307	5,441	49,205	SH		SOLE		49,205	0	0
STEWART INFORMATION SERVICES CORPORATION	COM	860372101	7,764	122,739	SH		SOLE		122,739	0	0
STRIDE, INC.	COM	86333M108	4,148	115,423	SH		SOLE		115,423	0	0
SUNOPTA, INC.	COM	8676EP108	21,963	2,459,433	SH		SOLE		2,459,433	0	0
SYNCHRONY FINANCIAL	COM	87165B103	28,746	588,098	SH		SOLE		588,098	0	0
TAYLOR MORRISON HOME CORPORATION	COM	87724P106	10,538	408,766	SH		SOLE		408,766	0	0
TECK RESOURCES LIMITED	COM	878742204	4,576	183,705	SH		SOLE		183,705	0	0
TECNOGLASS, INC.	COM	G87264100	30,757	1,415,438	SH		SOLE		1,415,438	0	0
TEXTAINER GROUP HOLDINGS LIMITED	COM	G8766E109	5,181	148,417	SH		SOLE		148,417	0	0
TRANSMEDICS GROUP, INC.	COM	89377M109	2,585	78,106	SH		SOLE		78,106	0	0
TRAVELCENTERS OF AMERICA, INC.	COM	89421B109	33,380	670,411	SH		SOLE		670,411	0	0
TRITON INTERNATIONAL LIMITED	COM	G9078F107	13,278	255,154	SH		SOLE		255,154	0	0
TRONOX HOLDINGS PLC	COM	G9087Q102	12,288	498,485	SH		SOLE		498,485	0	0
TWITTER, INC.	COM	90184L102	3,895	64,503	SH		SOLE		64,503	0	0
VONAGE HOLDINGS CORP.	COM	92886T201	5,203	322,786	SH		SOLE		322,786	0	0
WEST FRASER TIMBER CO. LTD.	COM	952845105	13,020	154,572	SH		SOLE		154,572	0	0
WM TECHNOLOGY, INC.	COM	92971A109	8,872	611,885	SH		SOLE		611,885	0	0
WTS/BM TECHNOLOGIES, INC.	WTS	05591L115	409	274,239	SH		SOLE		274,239	0	0
YELLOW CORPORATION	COM	985510106	9,672	1,711,805	SH		SOLE		1,711,805	0	0
ZUMIEZ, INC.	COM	989817101	6,118	153,871	SH		SOLE		153,871	0	0