Portolan Capital Management 13F-HR Quarterly holdings report by institutional manager

The Securities and Exchange Commission has not necessarily reviewed the information in this filing and has not determined if it is accurate and complete.
The reader should not assume that the information is accurate and complete.

UNITED STATES SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549
FORM 13F

FORM 13F INFORMATION TABLE

OMB APPROVAL

OMB Number:		3235-0006
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COLUMN 1	COLUMN 2	COLUMN 3	COLUMN 4	COLUMN 5			COLUMN 6	COLUMN 7	COLUMN 8
			VALUE	SHRS OR	SH/	PUT/	INVESTMENT	OTHER	VOTING AUTHORITY
NAME OF ISSUER	TITLE OF CLASS	CUSIP	(x$1000)	PRN AMT	PRN	CALL	DISCRETION	MANAGER	SOLE	SHARED	NONE
ABIOMED, INC.	COM	003654100	2,545	7,085	SH		SOLE		7,085	0	0
ACADEMY SPORTS & OUTDOORS, INC.	COM	00402L107	4,431	100,928	SH		SOLE		100,928	0	0
AEMETIS, INC.	COM	00770K202	1,305	106,090	SH		SOLE		106,090	0	0
ALPHABET, INC.	COM	02079K107	23,479	8,114	SH		SOLE		8,114	0	0
ALTA EQUIPMENT GROUP, INC.	COM	02128L106	23,600	1,611,989	SH		SOLE		1,611,989	0	0
AMAZON.COM, INC.	COM	023135106	18,682	5,603	SH		SOLE		5,603	0	0
AMBARELLA, INC.	COM	G037AX101	1,441	7,103	SH		SOLE		7,103	0	0
AMERICAN INTERNATIONAL GROUP, INC.	COM	026874784	3,715	65,342	SH		SOLE		65,342	0	0
AMERICAS CAR MART, INC.	COM	03062T105	6,911	67,490	SH		SOLE		67,490	0	0
APPLIED GENETIC TECHNOLOGIES CORPORATION	COM	03820J100	2,585	1,360,704	SH		SOLE		1,360,704	0	0
ARCBEST CORPORATION	COM	03937C105	15,585	130,034	SH		SOLE		130,034	0	0
ATLAS AIR WORLDWIDE HOLDINGS	COM	049164205	22,197	235,837	SH		SOLE		235,837	0	0
AVID BIOSERVICES, INC.	COM	05368M106	6,303	216,015	SH		SOLE		216,015	0	0
AVID TECHNOLOGY, INC.	COM	05367P100	14,115	433,382	SH		SOLE		433,382	0	0
AZENTA, INC.	COM	114340102	1,265	12,270	SH		SOLE		12,270	0	0
B. RILEY FINANCIAL, INC.	COM	05580M108	8,619	96,997	SH		SOLE		96,997	0	0
BABCOCK & WILCOX ENTERPRISES, INC.	COM	05614L209	11,932	1,322,808	SH		SOLE		1,322,808	0	0
BAIDU, INC.	COM	056752108	2,561	17,213	SH		SOLE		17,213	0	0
BANDWIDTH, INC.	COM	05988J103	32,641	454,862	SH		SOLE		454,862	0	0
BIO-RAD LABORATORIES, INC.	COM	090572207	14,901	19,721	SH		SOLE		19,721	0	0
BLUELINX HOLDINGS, INC.	COM	09624H208	43,264	451,793	SH		SOLE		451,793	0	0
BTRS HOLDINGS, INC.	COM	11778X104	2,580	329,933	SH		SOLE		329,933	0	0
BUNGE LIMITED	COM	G16962105	3,112	33,334	SH		SOLE		33,334	0	0
CALERES, INC.	COM	129500104	8,018	353,538	SH		SOLE		353,538	0	0
CALLAWAY GOLF	COM	131193104	9,131	332,756	SH		SOLE		332,756	0	0
CAPRI HOLDINGS LIMITED	COM	G1890L107	19,870	306,120	SH		SOLE		306,120	0	0
CARPARTS.COM, INC.	COM	14427M107	5,194	463,780	SH		SOLE		463,780	0	0
CASELLA WASTE SYSTEMS, INC.	COM	147448104	1,460	17,097	SH		SOLE		17,097	0	0
CENTURY COMMUNITIES, INC.	COM	156504300	9,764	119,379	SH		SOLE		119,379	0	0
CF INDUSTRIES HOLDINGS, INC.	COM	125269100	13,202	186,520	SH		SOLE		186,520	0	0
CHARAH SOLUTIONS, INC.	COM	15957P105	16,597	3,508,947	SH		SOLE		3,508,947	0	0
CHEGG, INC.	COM	163092109	6,180	201,306	SH		SOLE		201,306	0	0
COMMERCIAL VEHICLE GROUP, INC.	COM	202608105	594	73,707	SH		SOLE		73,707	0	0
CROCS, INC.	COM	227046109	33,091	258,083	SH		SOLE		258,083	0	0
CUE HEALTH, INC.	COM	229790100	10,607	791,013	SH		SOLE		791,013	0	0
CUSTOM TRUCK ONE SOURCE, INC.	COM	23204X103	11,392	1,423,966	SH		SOLE		1,423,966	0	0
CUSTOMERS BANCORP, INC.	COM	23204G100	41,499	634,836	SH		SOLE		634,836	0	0
CYBEROPTICS CORPORATION	COM	232517102	5,182	111,437	SH		SOLE		111,437	0	0
DANAOS CORPORATION	COM	Y1968P121	1,038	13,910	SH		SOLE		13,910	0	0
DARLING INGREDIENTS, INC.	COM	237266101	1,630	23,520	SH		SOLE		23,520	0	0
DASEKE, INC.	COM	23753F107	29,463	2,934,590	SH		SOLE		2,934,590	0	0
DATTO HOLDING CORP.	COM	23821D100	3,414	129,552	SH		SOLE		129,552	0	0
DECKERS OUTDOOR CORPORATION	COM	243537107	11,932	32,573	SH		SOLE		32,573	0	0
DENBURY, INC.	COM	24790A101	1,563	20,409	SH		SOLE		20,409	0	0
DESTINATION XL GROUP, INC.	COM	25065K104	6,598	1,161,668	SH		SOLE		1,161,668	0	0
DIGITAL TURBINE, INC.	COM	25400W102	16,199	265,600	SH		SOLE		265,600	0	0
DOMO, INC.	COM	257554105	23,254	468,821	SH		SOLE		468,821	0	0
E.L.F. BEAUTY, INC.	COM	26856L103	36,231	1,090,975	SH		SOLE		1,090,975	0	0
ENCORE CAPITAL GROUP, INC.	COM	292554102	7,832	126,091	SH		SOLE		126,091	0	0
ENERGY RECOVERY, INC.	COM	29270J100	1,486	69,150	SH		SOLE		69,150	0	0
ENOVA INTERNATIONAL, INC.	COM	29357K103	18,707	456,706	SH		SOLE		456,706	0	0
EURONET WORLDWIDE, INC.	COM	298736109	11,979	100,523	SH		SOLE		100,523	0	0
FIDELITY NATIONAL INFORMATION SERVICES, INC.	COM	31620M106	3,584	32,837	SH		SOLE		32,837	0	0
FORD MOTOR COMPANY	COM	345370860	9,862	474,814	SH		SOLE		474,814	0	0
FULL HOUSE RESORTS, INC.	COM	359678109	22,267	1,838,714	SH		SOLE		1,838,714	0	0
GENESCO, INC.	COM	371532102	20,987	327,057	SH		SOLE		327,057	0	0
GOLAR LNG LIMITED	COM	G9456A100	3,862	311,711	SH		SOLE		311,711	0	0
GREEN PLAINS, INC.	COM	393222104	29,760	856,170	SH		SOLE		856,170	0	0
GTY TECHNOLOGY HOLDINGS, INC.	COM	362409104	6,794	1,014,003	SH		SOLE		1,014,003	0	0
H & R BLOCK, INC.	COM	093671105	2,613	110,894	SH		SOLE		110,894	0	0
H&E EQUIPMENT SERVICES, INC.	COM	404030108	6,370	143,891	SH		SOLE		143,891	0	0
HARVARD BIOSCIENCE, INC.	COM	416906105	17,899	2,538,799	SH		SOLE		2,538,799	0	0
HIMAX TECHNOLOGIES, INC.	COM	43289P106	7,894	493,710	SH		SOLE		493,710	0	0
HUDSON TECHNOLOGIES, INC.	COM	444144109	73	16,523	SH		SOLE		16,523	0	0
IAC/INTERACTIVECORP.	COM	44891N208	1,541	11,788	SH		SOLE		11,788	0	0
IDENTIV, INC.	COM	45170X205	49,062	1,743,498	SH		SOLE		1,743,498	0	0
IMMERSION CORPORATION	COM	452521107	4,315	755,693	SH		SOLE		755,693	0	0
IMPINJ, INC.	COM	453204109	11,609	130,882	SH		SOLE		130,882	0	0
INOTIV, INC.	COM	45783Q100	30,644	728,395	SH		SOLE		728,395	0	0
INSPIRED ENTERTAINMENT, INC.	COM	45782N108	3,109	239,926	SH		SOLE		239,926	0	0
KALEYRA, INC.	COM	483379103	8,873	883,768	SH		SOLE		883,768	0	0
KURA SUSHI USA, INC.	COM	501270102	19,083	236,064	SH		SOLE		236,064	0	0
LAREDO PETROLEUM, INC.	COM	516806205	4,247	70,636	SH		SOLE		70,636	0	0
LIVEPERSON, INC.	COM	538146101	7,587	212,403	SH		SOLE		212,403	0	0
MAGNA INTERNATIONAL, INC.	COM	559222401	886	10,951	SH		SOLE		10,951	0	0
MAGNACHIP SEMICONDUCTOR CORPORATION	COM	55933J203	2,804	133,694	SH		SOLE		133,694	0	0
MARINEMAX, INC.	COM	567908108	13,702	232,076	SH		SOLE		232,076	0	0
MATCH GROUP, INC.	COM	57667L107	4,938	37,339	SH		SOLE		37,339	0	0
MAXLINEAR, INC.	COM	57776J100	5,846	77,547	SH		SOLE		77,547	0	0
META PLATFORMS, INC.	COM	30303M102	21,894	65,094	SH		SOLE		65,094	0	0
MGIC INVESTMENT CORPORATION	COM	552848103	1,386	96,142	SH		SOLE		96,142	0	0
MICROSOFT CORPORATION	COM	594918104	9,195	27,340	SH		SOLE		27,340	0	0
MKS INSTRUMENTS, INC.	COM	55306N104	6,263	35,958	SH		SOLE		35,958	0	0
MONEYGRAM INTERNATIONAL, INC.	COM	60935Y208	9,855	1,249,004	SH		SOLE		1,249,004	0	0
NAVIENT CORPORATION	COM	63938C108	9,025	425,294	SH		SOLE		425,294	0	0
NOODLES & COMPANY	COM	65540B105	2,740	302,067	SH		SOLE		302,067	0	0
NORTHERN OIL AND GAS, INC.	COM	665531307	1,214	58,972	SH		SOLE		58,972	0	0
NU SKIN ENTERPRISES, INC.	COM	67018T105	678	13,351	SH		SOLE		13,351	0	0
ONEMAIN HOLDINGS, INC.	COM	68268W103	8,228	164,434	SH		SOLE		164,434	0	0
OPEN LENDING CORPORATION	COM	68373J104	4,175	185,712	SH		SOLE		185,712	0	0
OVERSTOCK.COM, INC.	COM	690370101	2,636	44,670	SH		SOLE		44,670	0	0
PENN NATIONAL GAMING, INC.	COM	707569109	4,502	86,826	SH		SOLE		86,826	0	0
PFSWEB, INC.	COM	717098206	367	28,504	SH		SOLE		28,504	0	0
PORCH GROUP, INC.	COM	733245104	15,753	1,010,450	SH		SOLE		1,010,450	0	0
PROG HOLDINGS, INC.	COM	74319R101	9,479	210,126	SH		SOLE		210,126	0	0
PURETECH HEALTH PLC	COM	746237106	213	5,749	SH		SOLE		5,749	0	0
QUINSTREET, INC.	COM	74874Q100	28,278	1,554,614	SH		SOLE		1,554,614	0	0
RANPAK HOLDINGS CORPORATION	COM	75321W103	4,741	126,164	SH		SOLE		126,164	0	0
RENT-A-CENTER, INC.	COM	76009N100	4,809	100,108	SH		SOLE		100,108	0	0
REX AMERICAN RESOURCES CORPORATION	COM	761624105	14,216	148,086	SH		SOLE		148,086	0	0
RH	COM	74967X103	3,473	6,480	SH		SOLE		6,480	0	0
ROCKY BRANDS, INC.	COM	774515100	2,500	62,819	SH		SOLE		62,819	0	0
SANDRIDGE ENERGY, INC.	COM	80007P869	13,879	1,326,836	SH		SOLE		1,326,836	0	0
SEMLER SCIENTIFIC, INC.	COM	81684M104	4,407	48,088	SH		SOLE		48,088	0	0
SHYFT GROUP, INC. (THE)	COM	825698103	5,574	113,444	SH		SOLE		113,444	0	0
SKECHERS USA, INC.	COM	830566105	19,772	455,571	SH		SOLE		455,571	0	0
SKYLINE CHAMPION CORPORATION	COM	830830105	25,094	317,732	SH		SOLE		317,732	0	0
SONOS, INC.	COM	83570H108	8,865	297,468	SH		SOLE		297,468	0	0
SONY GROUP CORPORATION	COM	835699307	15,450	122,234	SH		SOLE		122,234	0	0
SPORTSMAN'S WAREHOUSE HOLDINGS, INC.	COM	84920Y106	4,375	370,733	SH		SOLE		370,733	0	0
STEWART INFORMATION SERVICES CORPORATION	COM	860372101	14,078	176,569	SH		SOLE		176,569	0	0
STRATASYS LTD	COM	M85548101	20,092	820,429	SH		SOLE		820,429	0	0
SUNOPTA, INC.	COM	8676EP108	15,468	2,225,556	SH		SOLE		2,225,556	0	0
SVB FINANCIAL GROUP	COM	78486Q101	7,358	10,849	SH		SOLE		10,849	0	0
SYNCHRONY FINANCIAL	COM	87165B103	24,853	535,733	SH		SOLE		535,733	0	0
SYNOVUS FINANCIAL CORP.	COM	87161C501	3,986	83,269	SH		SOLE		83,269	0	0
TAYLOR MORRISON HOME CORPORATION	COM	87724P106	18,894	540,450	SH		SOLE		540,450	0	0
TECNOGLASS, INC.	COM	G87264100	35,115	1,340,794	SH		SOLE		1,340,794	0	0
TEXTAINER GROUP HOLDINGS LIMITED	COM	G8766E109	18,191	509,404	SH		SOLE		509,404	0	0
TRAVELCENTERS OF AMERICA, INC.	COM	89421B109	31,770	615,462	SH		SOLE		615,462	0	0
TRITON INTERNATIONAL LIMITED	COM	G9078F107	15,450	256,516	SH		SOLE		256,516	0	0
TRONOX HOLDINGS PLC	COM	G9087Q102	12,452	518,168	SH		SOLE		518,168	0	0
TWITTER, INC.	COM	90184L102	7,918	183,191	SH		SOLE		183,191	0	0
VISTRA CORP.	COM	92840M102	9,356	410,911	SH		SOLE		410,911	0	0
VONAGE HOLDINGS CORP.	COM	92886T201	770	37,016	SH		SOLE		37,016	0	0
WEIBO CORPORATION	COM	948596101	2,322	74,938	SH		SOLE		74,938	0	0
WEST FRASER TIMBER CO. LTD.	COM	952845105	16,232	170,222	SH		SOLE		170,222	0	0
WTS/BM TECHNOLOGIES, INC.	WTS	05591L115	494	264,025	SH		SOLE		264,025	0	0
YELLOW CORPORATION	COM	985510106	15,755	1,251,403	SH		SOLE		1,251,403	0	0
ZIM INTEGRATED SHIPPING SERVICES LTD.	COM	M9T951109	17,857	303,387	SH		SOLE		303,387	0	0
ZUMIEZ, INC.	COM	989817101	4,340	90,434	SH		SOLE		90,434	0	0