Portolan Capital Management 13F-HR Quarterly holdings report by institutional manager

The Securities and Exchange Commission has not necessarily reviewed the information in this filing and has not determined if it is accurate and complete.
The reader should not assume that the information is accurate and complete.

UNITED STATES SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549
FORM 13F

FORM 13F INFORMATION TABLE

OMB APPROVAL

OMB Number:		3235-0006
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COLUMN 1	COLUMN 2	COLUMN 3	COLUMN 4	COLUMN 5			COLUMN 6	COLUMN 7	COLUMN 8
			VALUE	SHRS OR	SH/	PUT/	INVESTMENT	OTHER	VOTING AUTHORITY
NAME OF ISSUER	TITLE OF CLASS	CUSIP	(x$1000)	PRN AMT	PRN	CALL	DISCRETION	MANAGER	SOLE	SHARED	NONE
ABIOMED, INC.	COM	003654100	2,370	7,154	SH		SOLE		7,154	0	0
ACADEMY SPORTS & OUTDOORS, INC.	COM	00402L107	840	21,332	SH		SOLE		21,332	0	0
AGCO CORPORATION	COM	001084102	9,220	63,135	SH		SOLE		63,135	0	0
ALBEMARLE CORPORATION	COM	012653101	5,791	26,187	SH		SOLE		26,187	0	0
ALPHABET, INC.	COM	02079K107	13,256	4,746	SH		SOLE		4,746	0	0
ALTA EQUIPMENT GROUP, INC.	COM	02128L106	19,465	1,574,848	SH		SOLE		1,574,848	0	0
AMAZON.COM, INC.	COM	023135106	10,839	3,325	SH		SOLE		3,325	0	0
AMEDISYS, INC.	COM	023436108	2,861	16,607	SH		SOLE		16,607	0	0
ANDERSONS, INC.	COM	034164103	15,102	300,482	SH		SOLE		300,482	0	0
APPLIED GENETIC TECHNOLOGIES CORPORATION	COM	03820J100	1,718	1,605,599	SH		SOLE		1,605,599	0	0
ARCBEST CORPORATION	COM	03937C105	3,743	46,501	SH		SOLE		46,501	0	0
ARCHER-DANIELS-MIDLAND COMPANY	COM	039483102	14,035	155,496	SH		SOLE		155,496	0	0
ATLAS AIR WORLDWIDE HOLDINGS	COM	049164205	2,558	29,613	SH		SOLE		29,613	0	0
AVID BIOSERVICES, INC.	COM	05368M106	4,464	219,125	SH		SOLE		219,125	0	0
AVID TECHNOLOGY, INC.	COM	05367P100	25,781	739,345	SH		SOLE		739,345	0	0
BABCOCK & WILCOX ENTERPRISES, INC.	COM	05614L209	9,268	1,135,782	SH		SOLE		1,135,782	0	0
BANDWIDTH, INC.	COM	05988J103	33,021	1,019,493	SH		SOLE		1,019,493	0	0
BIO-RAD LABORATORIES, INC.	COM	090572207	19,320	34,303	SH		SOLE		34,303	0	0
BLUELINX HOLDINGS, INC.	COM	09624H208	19,387	269,711	SH		SOLE		269,711	0	0
BTRS HOLDINGS, INC.	COM	11778X104	1,036	138,559	SH		SOLE		138,559	0	0
BUCKLE, INC.	COM	118440106	3,168	95,897	SH		SOLE		95,897	0	0
BUNGE LIMITED	COM	G16962105	10,341	93,325	SH		SOLE		93,325	0	0
CAPRI HOLDINGS LIMITED	COM	G1890L107	11,720	228,062	SH		SOLE		228,062	0	0
CASELLA WASTE SYSTEMS, INC.	COM	147448104	1,499	17,097	SH		SOLE		17,097	0	0
CATALENT, INC.	COM	148806102	9,644	86,961	SH		SOLE		86,961	0	0
CF INDUSTRIES HOLDINGS, INC.	COM	125269100	3,735	36,239	SH		SOLE		36,239	0	0
CHAMPIONX CORPORATION	COM	15872M104	5,717	233,550	SH		SOLE		233,550	0	0
CHARAH SOLUTIONS, INC.	COM	15957P105	17,924	3,591,972	SH		SOLE		3,591,972	0	0
CHESAPEAKE ENERGY CORPORATION	COM	165167735	2,965	34,077	SH		SOLE		34,077	0	0
CHILDREN'S PLACE, INC. (THE)	COM	168905107	3,319	67,330	SH		SOLE		67,330	0	0
COMMERCIAL VEHICLE GROUP, INC.	COM	202608105	323	38,212	SH		SOLE		38,212	0	0
CROCS, INC.	COM	227046109	26,110	341,755	SH		SOLE		341,755	0	0
CUSTOM TRUCK ONE SOURCE, INC.	COM	23204X103	12,153	1,448,509	SH		SOLE		1,448,509	0	0
CUSTOMERS BANCORP, INC.	COM	23204G100	25,492	488,908	SH		SOLE		488,908	0	0
CYBEROPTICS CORPORATION	COM	232517102	6,243	153,844	SH		SOLE		153,844	0	0
DANAOS CORPORATION	COM	Y1968P121	1,426	13,905	SH		SOLE		13,905	0	0
DASEKE, INC.	COM	23753F107	26,511	2,632,670	SH		SOLE		2,632,670	0	0
DESTINATION XL GROUP, INC.	COM	25065K104	1,307	267,843	SH		SOLE		267,843	0	0
DHT HOLDINGS, INC.	COM	Y2065G121	2,994	516,213	SH		SOLE		516,213	0	0
DIGITAL TURBINE, INC.	COM	25400W102	28,467	649,789	SH		SOLE		649,789	0	0
DOMO, INC.	COM	257554105	37,610	743,725	SH		SOLE		743,725	0	0
E.L.F. BEAUTY, INC.	COM	26856L103	14,372	556,404	SH		SOLE		556,404	0	0
ELANCO ANIMAL HEALTH INCORPORATED	COM	28414H103	2,682	102,813	SH		SOLE		102,813	0	0
ENCORE CAPITAL GROUP, INC.	COM	292554102	1,877	29,923	SH		SOLE		29,923	0	0
ENOVA INTERNATIONAL, INC.	COM	29357K103	4,580	120,614	SH		SOLE		120,614	0	0
EURONET WORLDWIDE, INC.	COM	298736109	6,176	47,453	SH		SOLE		47,453	0	0
F45 TRAINING HOLDINGS, INC.	COM	30322L101	380	35,547	SH		SOLE		35,547	0	0
F5, INC.	COM	315616102	10,225	48,937	SH		SOLE		48,937	0	0
FERROGLOBE PLC	COM	G33856108	5,182	673,015	SH		SOLE		673,015	0	0
FIDELITY NATIONAL INFORMATION SERVICES, INC.	COM	31620M106	6,287	62,606	SH		SOLE		62,606	0	0
FULL HOUSE RESORTS	COM	359678109	11,286	1,174,411	SH		SOLE		1,174,411	0	0
GENESCO, INC.	COM	371532102	10,372	163,059	SH		SOLE		163,059	0	0
GFL ENVIRONMENTAL, INC.	COM	36168Q104	10,649	327,257	SH		SOLE		327,257	0	0
GOLAR LNG LIMITED	COM	G9456A100	8,305	335,159	SH		SOLE		335,159	0	0
GREEN PLAINS, INC.	COM	393222104	11,551	372,502	SH		SOLE		372,502	0	0
GTY TECHNOLOGY HOLDINGS, INC.	COM	362409104	4,512	1,397,039	SH		SOLE		1,397,039	0	0
GULFPORT ENERGY CORPORATION	COM	402635502	1,332	14,833	SH		SOLE		14,833	0	0
H&E EQUIPMENT SERVICES, INC.	COM	404030108	7,753	178,139	SH		SOLE		178,139	0	0
HARVARD BIOSCIENCE, INC.	COM	416906105	15,691	2,526,789	SH		SOLE		2,526,789	0	0
HCI GROUP, INC.	COM	40416E103	5,815	85,283	SH		SOLE		85,283	0	0
HEALTH CATALYST, INC.	COM	42225T107	7,249	277,407	SH		SOLE		277,407	0	0
HEALTHEQUITY, INC.	COM	42226A107	8,650	128,261	SH		SOLE		128,261	0	0
HIMAX TECHNOLOGIES, INC.	COM	43289P106	3,014	277,528	SH		SOLE		277,528	0	0
HUDSON PACIFIC PROPERTIES, INC.	COM	444097109	6,763	243,702	SH		SOLE		243,702	0	0
I3 VERTICALS, INC.	COM	46571Y107	7,573	271,833	SH		SOLE		271,833	0	0
IAC/INTERACTIVECORP.	COM	44891N208	723	7,206	SH		SOLE		7,206	0	0
ICAD, INC.	COM	44934S206	4,041	905,984	SH		SOLE		905,984	0	0
IDENTIV, INC.	COM	45170X205	32,820	2,029,694	SH		SOLE		2,029,694	0	0
IMMERSION CORPORATION	COM	452521107	7,029	1,264,245	SH		SOLE		1,264,245	0	0
INOTIV, INC.	COM	45783Q100	10,236	390,988	SH		SOLE		390,988	0	0
INSPIRED ENTERTAINMENT, INC.	COM	45782N108	6,024	489,720	SH		SOLE		489,720	0	0
KALEYRA, INC.	COM	483379103	9,915	1,658,066	SH		SOLE		1,658,066	0	0
KURA SUSHI USA, INC.	COM	501270102	18,482	335,127	SH		SOLE		335,127	0	0
LANTRONIX, INC.	COM	516548203	334	50,000	SH		SOLE		50,000	0	0
LAREDO PETROLEUM, INC.	COM	516806205	16,869	213,150	SH		SOLE		213,150	0	0
LEIDOS HLDGS, INC.	COM	525327102	10,342	95,745	SH		SOLE		95,745	0	0
LESLIE'S, INC.	COM	527064109	3,829	197,787	SH		SOLE		197,787	0	0
LIVEPERSON, INC.	COM	538146101	13,705	561,227	SH		SOLE		561,227	0	0
MAGNACHIP SEMICONDUCTOR CORPORATION	COM	55933J203	2,875	170,020	SH		SOLE		170,020	0	0
MATCH GROUP, INC.	COM	57667L107	805	7,406	SH		SOLE		7,406	0	0
MATERION CORPORATION	COM	576690101	3,337	38,922	SH		SOLE		38,922	0	0
MAXLINEAR, INC.	COM	57776J100	2,808	48,121	SH		SOLE		48,121	0	0
META PLATFORMS, INC.	COM	30303M102	8,715	39,191	SH		SOLE		39,191	0	0
MGIC INVESTMENT CORPORATION	COM	552848103	3,279	241,990	SH		SOLE		241,990	0	0
MGP INGREDIENTS, INC.	COM	55303J106	7,369	86,101	SH		SOLE		86,101	0	0
MICROSOFT CORPORATION	COM	594918104	4,210	13,656	SH		SOLE		13,656	0	0
NMI HOLDINGS, INC.	COM	629209305	4,400	213,405	SH		SOLE		213,405	0	0
NOODLES & COMPANY	COM	65540B105	1,659	277,845	SH		SOLE		277,845	0	0
NORTHERN OIL AND GAS, INC.	COM	665531307	13,469	477,799	SH		SOLE		477,799	0	0
NUTRIEN LTD.	COM	67077M108	3,620	34,815	SH		SOLE		34,815	0	0
OLIN CORPORATION	COM	680665205	7,872	150,583	SH		SOLE		150,583	0	0
PARTY CITY HOLDCO, INC.	COM	702149105	1,790	500,000	SH		SOLE		500,000	0	0
PDC ENERGY, INC.	COM	69327R101	3,469	47,723	SH		SOLE		47,723	0	0
PETIQ, INC.	COM	71639T106	9,476	388,358	SH		SOLE		388,358	0	0
PFSWEB, INC.	COM	717098206	313	27,363	SH		SOLE		27,363	0	0
PORCH GROUP, INC.	COM	733245104	5,344	769,499	SH		SOLE		769,499	0	0
QUALYS, INC.	COM	74758T303	5,580	39,186	SH		SOLE		39,186	0	0
QUINSTREET, INC.	COM	74874Q100	9,994	861,538	SH		SOLE		861,538	0	0
R1 RCM, INC.	COM	749397105	8,828	329,901	SH		SOLE		329,901	0	0
RESOLUTE FOREST PRODUCTS, INC.	COM	76117W109	6,852	530,772	SH		SOLE		530,772	0	0
REX AMERICAN RESOURCES CORPORATION	COM	761624105	10,017	100,570	SH		SOLE		100,570	0	0
RH	COM	74967X103	652	1,998	SH		SOLE		1,998	0	0
SANDRIDGE ENERGY, INC.	COM	80007P869	5,787	361,211	SH		SOLE		361,211	0	0
SCIENCE APPLICATIONS INTERNATIONAL CORPORATION	COM	808625107	4,734	51,363	SH		SOLE		51,363	0	0
SHYFT GROUP, INC. (THE)	COM	825698103	918	25,416	SH		SOLE		25,416	0	0
SIBANYE STILLWATER LIMITED	COM	82575P107	7,413	456,761	SH		SOLE		456,761	0	0
SIGNIFY HEALTH, INC.	COM	82671G100	6,129	337,693	SH		SOLE		337,693	0	0
SKECHERS USA, INC.	COM	830566105	10,567	259,241	SH		SOLE		259,241	0	0
SKYLINE CHAMPION CORPORATION	COM	830830105	3,026	55,139	SH		SOLE		55,139	0	0
SONY GROUP CORPORATION	COM	835699307	13,267	129,172	SH		SOLE		129,172	0	0
STAAR SURGICAL COMPANY	COM	852312305	8,934	111,798	SH		SOLE		111,798	0	0
STEWART INFORMATION SERVICES CORPORATION	COM	860372101	4,314	71,184	SH		SOLE		71,184	0	0
STRATASYS LTD	COM	M85548101	26,940	1,061,067	SH		SOLE		1,061,067	0	0
SUNOPTA, INC.	COM	8676EP108	7,349	1,463,962	SH		SOLE		1,463,962	0	0
SVB FINANCIAL GROUP	COM	78486Q101	4,036	7,214	SH		SOLE		7,214	0	0
SYNOVUS FINANCIAL CORP.	COM	87161C501	7,195	146,838	SH		SOLE		146,838	0	0
TAKE TWO INTERACTIVE SOFTWARE, INC.	COM	874054109	21,110	137,311	SH		SOLE		137,311	0	0
TALOS ENERGY, INC.	COM	87484T108	9,172	580,868	SH		SOLE		580,868	0	0
TAYLOR MORRISON HOME CORP.	COM	87724P106	2,971	109,154	SH		SOLE		109,154	0	0
TECHNIPFMC PLC	COM	G87110105	5,165	666,424	SH		SOLE		666,424	0	0
TECK RESOURCES LIMITED	COM	878742204	10,244	253,627	SH		SOLE		253,627	0	0
TECNOGLASS, INC.	COM	G87264100	22,852	905,373	SH		SOLE		905,373	0	0
TENABLE HOLDINGS, INC.	COM	88025T102	11,321	195,897	SH		SOLE		195,897	0	0
TEXTAINER GROUP HOLDINGS	COM	G8766E109	19,851	521,432	SH		SOLE		521,432	0	0
THERMO FISHER SCIENTIFIC, INC.	COM	883556102	12,282	20,794	SH		SOLE		20,794	0	0
TRANSMEDICS GROUP, INC.	COM	89377M109	10,233	379,859	SH		SOLE		379,859	0	0
TRAVELCENTERS OF AMERICA, INC.	COM	89421B109	25,113	584,556	SH		SOLE		584,556	0	0
TRIPADVISOR, INC.	COM	896945201	4,094	150,955	SH		SOLE		150,955	0	0
TRITON INTERNATIONAL LIMITED	COM	G9078F107	15,199	216,566	SH		SOLE		216,566	0	0
TWILIO, INC.	COM	90138F102	8,359	50,721	SH		SOLE		50,721	0	0
TWITTER, INC.	COM	90184L102	14,440	373,234	SH		SOLE		373,234	0	0
VALERO ENERGY CORPORATION	COM	91913Y100	2,962	29,168	SH		SOLE		29,168	0	0
VISTRA CORP.	COM	92840M102	29,054	1,249,640	SH		SOLE		1,249,640	0	0
WEST FRASER TIMBER CO. LTD.	COM	952845105	13,537	164,225	SH		SOLE		164,225	0	0
WORKDAY, INC.	COM	98138H101	4,047	16,899	SH		SOLE		16,899	0	0
ZIM INTEGRATED SHIPPING SERVICES LTD.	COM	M9T951109	15,917	218,911	SH		SOLE		218,911	0	0