Portolan Capital Management 13F-HR Quarterly holdings report by institutional manager

The Securities and Exchange Commission has not necessarily reviewed the information in this filing and has not determined if it is accurate and complete.
The reader should not assume that the information is accurate and complete.

UNITED STATES SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549
FORM 13F

FORM 13F INFORMATION TABLE

OMB APPROVAL

OMB Number:		3235-0006
Expires:	Oct 31, 2018
Estimated average burden
hours per response:		23.8

COLUMN 1	COLUMN 2	COLUMN 3	COLUMN 4	COLUMN 5			COLUMN 6	COLUMN 7	COLUMN 8
			VALUE	SHRS OR	SH/	PUT/	INVESTMENT	OTHER	VOTING AUTHORITY
NAME OF ISSUER	TITLE OF CLASS	CUSIP	(x$1000)	PRN AMT	PRN	CALL	DISCRETION	MANAGER	SOLE	SHARED	NONE
A10 NETWORKS, INC.	COM	002121101	3,828	288,460	SH		SOLE		288,460	0	0
ABIOMED, INC.	COM	003654100	1,754	7,139	SH		SOLE		7,139	0	0
AGCO CORPORATION	COM	001084102	2,262	23,521	SH		SOLE		23,521	0	0
AGILON HEALTH, INC.	COM	00857U107	4,351	185,797	SH		SOLE		185,797	0	0
ALBEMARLE CORPORATION	COM	012653101	6,681	25,265	SH		SOLE		25,265	0	0
ALPHABET, INC.	COM	02079K107	1,670	17,365	SH		SOLE		17,365	0	0
ALTA EQUIPMENT GROUP, INC.	COM	02128L106	8,337	757,239	SH		SOLE		757,239	0	0
AMAZON.COM, INC.	COM	023135106	1,656	14,652	SH		SOLE		14,652	0	0
AMEDISYS, INC.	COM	023436108	3,706	38,286	SH		SOLE		38,286	0	0
AMPHASTAR PHARMACEUTICALS, INC.	COM	03209R103	7,929	282,187	SH		SOLE		282,187	0	0
ANDERSONS, INC.	COM	034164103	8,215	264,745	SH		SOLE		264,745	0	0
ANIKA THERAPEUTICS, INC.	COM	035255108	5,584	234,603	SH		SOLE		234,603	0	0
APPLIED GENETIC TECHNOLOGIES CORPORATION	COM	03820J100	435	1,600,902	SH		SOLE		1,600,902	0	0
ARCHAEA ENERGY, INC.	COM	03940F103	3,654	202,909	SH		SOLE		202,909	0	0
ARCHER-DANIELS-MIDLAND COMPANY	COM	039483102	9,899	123,050	SH		SOLE		123,050	0	0
AVID BIOSERVICES, INC.	COM	05368M106	1,230	64,309	SH		SOLE		64,309	0	0
AVID TECHNOLOGY, INC.	COM	05367P100	20,291	872,374	SH		SOLE		872,374	0	0
BANDWIDTH, INC.	COM	05988J103	921	77,409	SH		SOLE		77,409	0	0
THE BEAUTY HEALTH COMPANY	COM	88331L108	4,605	390,599	SH		SOLE		390,599	0	0
BEL FUSE, INC.	COM	077347300	7,971	315,702	SH		SOLE		315,702	0	0
BIO-RAD LABORATORIES, INC.	COM	090572207	18,741	44,927	SH		SOLE		44,927	0	0
BLUCORA, INC.	COM	095229100	7,729	399,648	SH		SOLE		399,648	0	0
BLUELINX HOLDINGS, INC.	COM	09624H208	9,797	157,762	SH		SOLE		157,762	0	0
BOOT BARN HOLDINGS, INC.	COM	099406100	4,403	75,320	SH		SOLE		75,320	0	0
BOX, INC.	COM	10316T104	8,022	328,916	SH		SOLE		328,916	0	0
CALAMP CORPORATION	COM	128126109	1,104	287,390	SH		SOLE		287,390	0	0
CAPRI HOLDINGS LIMITED	COM	G1890L107	3,961	103,035	SH		SOLE		103,035	0	0
CARPARTS.COM, INC.	COM	14427M107	7,663	1,482,290	SH		SOLE		1,482,290	0	0
CASELLA WASTE SYSTEMS, INC.	COM	147448104	1,306	17,097	SH		SOLE		17,097	0	0
CASTLE BIOSCIENCES, INC.	COM	14843C105	7,839	300,590	SH		SOLE		300,590	0	0
CATALENT, INC.	COM	148806102	5,416	74,842	SH		SOLE		74,842	0	0
CF INDUSTRIES HOLDINGS, INC.	COM	125269100	19,906	206,820	SH		SOLE		206,820	0	0
CHARAH SOLUTIONS, INC.	COM	15957P105	4,523	2,471,443	SH		SOLE		2,471,443	0	0
CHECK POINT SOFTWARE TECHNOLOGIES LTD.	COM	M22465104	6,443	57,518	SH		SOLE		57,518	0	0
CHEGG, INC.	COM	163092109	4,354	206,627	SH		SOLE		206,627	0	0
COMPASS MINERALS INTERNATIONAL, INC.	COM	20451N101	7,225	187,518	SH		SOLE		187,518	0	0
COUCHBASE, INC.	COM	22207T101	4,421	309,785	SH		SOLE		309,785	0	0
CROCS, INC.	COM	227046109	27,722	403,754	SH		SOLE		403,754	0	0
CUSTOM TRUCK ONE SOURCE, INC.	COM	23204X103	6,903	1,184,059	SH		SOLE		1,184,059	0	0
DARLING INGREDIENTS, INC.	COM	237266101	4,756	71,893	SH		SOLE		71,893	0	0
DASEKE, INC.	COM	23753F107	5,210	962,941	SH		SOLE		962,941	0	0
DECKERS OUTDOORS CORPORATION	COM	243537107	5,160	16,506	SH		SOLE		16,506	0	0
DESTINATION XL GROUP, INC.	COM	25065K104	2,532	467,097	SH		SOLE		467,097	0	0
DHT HOLDINGS, INC.	COM	Y2065G121	9,286	1,228,327	SH		SOLE		1,228,327	0	0
DIGI INTERNATIONAL, INC.	COM	253798102	4,964	143,607	SH		SOLE		143,607	0	0
DOCGO, INC.	COM	256086109	6,233	628,312	SH		SOLE		628,312	0	0
DOMO, INC.	COM	257554105	15,576	865,814	SH		SOLE		865,814	0	0
E.L.F. BEAUTY, INC.	COM	26856L103	28,282	751,791	SH		SOLE		751,791	0	0
E2OPEN PARENT HOLDINGS, INC.	COM	29788T103	2,566	422,758	SH		SOLE		422,758	0	0
ELECTRONIC ARTS CM	COM	285512109	1,849	15,980	SH		SOLE		15,980	0	0
ENCORE CAPITAL GROUP, INC.	COM	292554102	1,348	29,630	SH		SOLE		29,630	0	0
ENOVA INTERNATIONAL, INC.	COM	29357K103	2,671	91,240	SH		SOLE		91,240	0	0
EXPEDIA GROUP, INC.	COM	30212P303	3,448	36,801	SH		SOLE		36,801	0	0
EXTREME NETWORKS, INC.	COM	30226D106	8,034	614,653	SH		SOLE		614,653	0	0
F5, INC.	COM	315616102	7,092	49,002	SH		SOLE		49,002	0	0
FLEX LTD.	COM	Y2573F102	5,555	333,437	SH		SOLE		333,437	0	0
FULL HOUSE RESORTS, INC.	COM	359678109	2,661	473,502	SH		SOLE		473,502	0	0
GREEN PLAINS, INC.	COM	393222104	4,452	153,148	SH		SOLE		153,148	0	0
H&E EQUIPMENT SERVICES, INC.	COM	404030108	3,310	116,795	SH		SOLE		116,795	0	0
HARVARD BIOSCIENCE, INC.	COM	416906105	3,436	1,342,091	SH		SOLE		1,342,091	0	0
HEALTH CATALYST, INC.	COM	42225T107	2,945	303,602	SH		SOLE		303,602	0	0
HEALTHEQUITY, INC.	COM	42226A107	3,910	58,206	SH		SOLE		58,206	0	0
HONEST COMPANY, INC.	COM	438333106	4,175	1,192,794	SH		SOLE		1,192,794	0	0
HUDSON PACIFIC PROPERTIES, INC.	COM	444097109	1,718	156,918	SH		SOLE		156,918	0	0
I3 VERTICALS, INC.	COM	46571Y107	13,026	650,322	SH		SOLE		650,322	0	0
ICAD, INC.	COM	44934S206	865	411,724	SH		SOLE		411,724	0	0
IDENTIV, INC.	COM	45170X205	25,733	2,052,095	SH		SOLE		2,052,095	0	0
IMMERSION CORPORATION	COM	452521107	6,249	1,138,191	SH		SOLE		1,138,191	0	0
INTERNATIONAL MONEY EXPRESS, INC.	COM	46005L101	9,662	423,962	SH		SOLE		423,962	0	0
INTERNATIONAL SEAWAYS, INC.	COM	Y41053102	6,079	173,033	SH		SOLE		173,033	0	0
JAKKS PACIFIC, INC.	COM	47012E403	2,223	114,987	SH		SOLE		114,987	0	0
KOSMOS ENERGY LTD.	COM	500688106	4,031	779,637	SH		SOLE		779,637	0	0
KROGER COMPANY	COM	501044101	4,844	110,727	SH		SOLE		110,727	0	0
KURA SUSHI USA, INC.	COM	501270102	21,825	296,622	SH		SOLE		296,622	0	0
L3HARRIS TECHNOLOGIES, INC.	COM	502431109	7,570	36,423	SH		SOLE		36,423	0	0
LANTRONIX, INC.	COM	516548203	12,444	2,587,131	SH		SOLE		2,587,131	0	0
LAREDO PETROLEUM, INC.	COM	516806205	9,768	155,419	SH		SOLE		155,419	0	0
LEIDOS HOLDINGS, INC.	COM	525327102	6,699	76,590	SH		SOLE		76,590	0	0
LSB INDUSTRIES, INC.	COM	502160104	10,147	712,068	SH		SOLE		712,068	0	0
MARINEMAX, INC.	COM	567908108	2,246	75,390	SH		SOLE		75,390	0	0
MATCH GROUP, INC.	COM	57667L107	340	7,123	SH		SOLE		7,123	0	0
META PLATFORMS, INC.	COM	30303M102	5,453	40,188	SH		SOLE		40,188	0	0
MGP INGREDIENTS, INC.	COM	55303J106	5,386	50,733	SH		SOLE		50,733	0	0
MICROSOFT CORPORATION	COM	594918104	2,381	10,225	SH		SOLE		10,225	0	0
NATERA, INC.	COM	632307104	13,650	311,497	SH		SOLE		311,497	0	0
NAVIGATOR HOLDINGS LTD.	COM	Y62132108	3,515	307,014	SH		SOLE		307,014	0	0
NOBLE CORPORATION PLC	COM	G65431127	3,323	112,340	SH		SOLE		112,340	0	0
NOMAD FOODS LTD.	COM	G6564A105	8,401	591,585	SH		SOLE		591,585	0	0
NORTHERN OIL AND GAS, INC.	COM	665531307	13,331	486,355	SH		SOLE		486,355	0	0
NUTRIEN LTD.	COM	67077M108	6,313	75,709	SH		SOLE		75,709	0	0
OCWEN FINANCIAL CORPORATION	COM	675746606	762	32,734	SH		SOLE		32,734	0	0
OLIN CORPORATION	COM	680665205	2,406	56,113	SH		SOLE		56,113	0	0
ORACLE CORPORATION	COM	68389X105	17,334	283,838	SH		SOLE		283,838	0	0
PAYONEER GLOBAL, INC.	COM	70451X104	3,488	576,598	SH		SOLE		576,598	0	0
PBF ENERGY, INC.	COM	69318G106	16,600	472,136	SH		SOLE		472,136	0	0
PORCH GROUP, INC.	COM	733245104	1,641	729,449	SH		SOLE		729,449	0	0
PREMIER, INC.	COM	74051N102	12,954	381,673	SH		SOLE		381,673	0	0
PRIVIA HEALTH GROUP, INC.	COM	74276R102	7,112	208,807	SH		SOLE		208,807	0	0
QUALYS, INC.	COM	74758T303	3,351	24,043	SH		SOLE		24,043	0	0
QUANTERIX CORPORATION	COM	74766Q101	8,363	758,853	SH		SOLE		758,853	0	0
QUINSTREET, INC.	COM	74874Q100	2,601	247,703	SH		SOLE		247,703	0	0
R1 RCM, INC.	COM	77634L105	15,130	816,515	SH		SOLE		816,515	0	0
REPAY HOLDINGS CORPORATION	COM	76029L100	2,415	342,081	SH		SOLE		342,081	0	0
RICHARDSON ELECTRONICS LTD.	COM	763165107	10,664	706,240	SH		SOLE		706,240	0	0
SANDRIDGE ENERGY, INC.	COM	80007P869	409	25,079	SH		SOLE		25,079	0	0
SCIENCE APPLICATIONS INTERNATIONAL CORPORATION	COM	808625107	6,955	78,653	SH		SOLE		78,653	0	0
SFL CORPORATION LTD.	COM	G7738W106	3,529	387,371	SH		SOLE		387,371	0	0
SKECHERS USA, INC.	COM	830566105	4,130	130,203	SH		SOLE		130,203	0	0
SKYLINE CHAMPION CORPORATION	COM	830830105	2,386	45,124	SH		SOLE		45,124	0	0
SOCIEDAD QUIMICA MINERAL DE CHILE S A	COM	833635105	12,616	139,019	SH		SOLE		139,019	0	0
SONY GROUP CORPORATION	COM	835699307	4,687	73,185	SH		SOLE		73,185	0	0
STAAR SURGICAL COMPANY	COM	852312305	4,345	61,583	SH		SOLE		61,583	0	0
STRATASYS LTD.	COM	M85548101	2,103	145,936	SH		SOLE		145,936	0	0
STRIDE, INC.	COM	86333M108	27,825	662,027	SH		SOLE		662,027	0	0
SUNOPTA, INC.	COM	8676EP108	18,816	2,067,700	SH		SOLE		2,067,700	0	0
TAKE TWO INTERACTIVE SOFTWARE, INC.	COM	874054109	6,048	55,483	SH		SOLE		55,483	0	0
TECNOGLASS, INC.	COM	G87264100	23,080	1,099,591	SH		SOLE		1,099,591	0	0
THERMO FISHER SCIENTIFIC, INC.	COM	883556102	6,499	12,814	SH		SOLE		12,814	0	0
TITAN MACHINERY, INC.	COM	88830R101	4,271	151,119	SH		SOLE		151,119	0	0
TRAVELCENTERS OF AMERICA, INC.	COM	89421B109	33,005	612,005	SH		SOLE		612,005	0	0
TRIPADVISOR, INC.	COM	896945201	7,659	346,858	SH		SOLE		346,858	0	0
TWILIO, INC.	COM	90138F102	10,009	144,758	SH		SOLE		144,758	0	0
UNITED THERAPEUTICS CORPORATION	COM	91307C102	17,021	81,290	SH		SOLE		81,290	0	0
VERMILION ENERGY, INC.	COM	923725105	4,543	212,091	SH		SOLE		212,091	0	0
VISTRA CORP CMN	COM	92840M102	9,943	473,494	SH		SOLE		473,494	0	0
WALKME LTD.	COM	M97628107	7,276	856,020	SH		SOLE		856,020	0	0
WEST FRASER TIMBER CO. LTD.	COM	952845105	10,257	141,496	SH		SOLE		141,496	0	0
WORKDAY, INC.	COM	98138H101	4,830	31,728	SH		SOLE		31,728	0	0