Portolan Capital Management 13F-HR Quarterly holdings report by institutional manager

The Securities and Exchange Commission has not necessarily reviewed the information in this filing and has not determined if it is accurate and complete.
The reader should not assume that the information is accurate and complete.

UNITED STATES SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549
FORM 13F

FORM 13F INFORMATION TABLE

OMB APPROVAL

OMB Number:		3235-0006
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COLUMN 1	COLUMN 2	COLUMN 3	COLUMN 4	COLUMN 5			COLUMN 6	COLUMN 7	COLUMN 8
			VALUE	SHRS OR	SH/	PUT/	INVESTMENT	OTHER	VOTING AUTHORITY
NAME OF ISSUER	TITLE OF CLASS	CUSIP	(x$1000)	PRN AMT	PRN	CALL	DISCRETION	MANAGER	SOLE	SHARED	NONE
A10 NETWORKS, INC.	COM	002121101	682,362	41,032	SH		SOLE		41,032	0	0
ACACIA RESEARCH CORPORATION	COM	003881307	2,681,732	636,991	SH		SOLE		636,991	0	0
ADOBE, INC.	COM	00724F101	8,318,685	24,719	SH		SOLE		24,719	0	0
AEHR TEST SYSTEMS	COM	00760J108	1,370,639	68,191	SH		SOLE		68,191	0	0
ALPHABET, INC.	COM	02079K107	2,362,791	26,629	SH		SOLE		26,629	0	0
ALTA EQUIPMENT GROUP, INC.	COM	02128L106	7,547,925	572,246	SH		SOLE		572,246	0	0
AMERICAN PUBLIC EDUCATION, INC.	COM	02913V103	4,299,103	349,805	SH		SOLE		349,805	0	0
AMPHASTAR PHARMACEUTICALS, INC.	COM	03209R103	7,587,564	270,791	SH		SOLE		270,791	0	0
ANDERSONS, INC.	COM	034164103	3,877,452	110,816	SH		SOLE		110,816	0	0
ANIKA THERAPEUTICS, INC.	COM	035255108	6,153,840	207,900	SH		SOLE		207,900	0	0
ARCHER-DANIELS-MIDLAND COMPANY	COM	039483102	1,324,134	14,261	SH		SOLE		14,261	0	0
ASSOCIATED BANC-CORP	COM	045487105	3,224,934	139,668	SH		SOLE		139,668	0	0
AVID BIOSERVICES, INC.	COM	05368M106	483,437	35,108	SH		SOLE		35,108	0	0
AVID TECHNOLOGY, INC.	COM	05367P100	20,656,096	776,837	SH		SOLE		776,837	0	0
BABCOCK & WILCOX ENTERPRISES, INC.	COM	05614L209	2,165,129	375,239	SH		SOLE		375,239	0	0
BANDWIDTH, INC.	COM	05988J103	3,604,734	157,069	SH		SOLE		157,069	0	0
BEL FUSE, INC.	COM	077347300	16,014,790	486,476	SH		SOLE		486,476	0	0
BIO-RAD LABORATORIES, INC.	COM	090572207	26,092,666	62,053	SH		SOLE		62,053	0	0
BLUCORA, INC.	COM	095229100	9,602,420	376,123	SH		SOLE		376,123	0	0
BLUELINX HOLDINGS, INC.	COM	09624H208	14,280,310	200,820	SH		SOLE		200,820	0	0
BOX, INC.	COM	10316T104	9,660,417	310,325	SH		SOLE		310,325	0	0
CAPRI HOLDINGS LIMITED	COM	G1890L107	10,461,015	182,502	SH		SOLE		182,502	0	0
CARPARTS.COM, INC.	COM	14427M107	15,089,148	2,410,407	SH		SOLE		2,410,407	0	0
CASELLA WASTE SYSTEMS, INC.	COM	147448104	1,355,963	17,097	SH		SOLE		17,097	0	0
CASTLE BIOSCIENCES, INC.	COM	14843C105	13,740,675	583,716	SH		SOLE		583,716	0	0
CHARAH SOLUTIONS, INC.	COM	15957P303	900,823	166,819	SH		SOLE		166,819	0	0
CLEARFIELD, INC.	COM	18482P103	3,667,224	38,955	SH		SOLE		38,955	0	0
COUCHBASE, INC.	COM	22207T101	12,463,710	939,948	SH		SOLE		939,948	0	0
CROCS, INC.	COM	227046109	40,881,797	377,034	SH		SOLE		377,034	0	0
CUSTOM TRUCK ONE SOURCE, INC.	COM	23204X103	6,584,896	1,041,914	SH		SOLE		1,041,914	0	0
DASEKE, INC.	COM	23753F107	3,446,331	605,682	SH		SOLE		605,682	0	0
DECKERS OUTDOORS CORPORATION	COM	243537107	11,815,934	29,602	SH		SOLE		29,602	0	0
DESTINATION XL GROUP, INC.	COM	25065K104	3,146,546	466,155	SH		SOLE		466,155	0	0
DHT HOLDINGS, INC.	COM	Y2065G121	1,225,928	138,055	SH		SOLE		138,055	0	0
DIGI INTERNATIONAL, INC.	COM	253798102	5,937,511	162,449	SH		SOLE		162,449	0	0
DIGITAL TURBINE, INC.	COM	25400W102	8,615,142	565,298	SH		SOLE		565,298	0	0
DOCGO, INC.	COM	256086109	12,488,441	1,766,399	SH		SOLE		1,766,399	0	0
DOMO, INC.	COM	257554105	2,810,976	197,400	SH		SOLE		197,400	0	0
E.L.F. BEAUTY, INC.	COM	26856L103	34,948,660	631,983	SH		SOLE		631,983	0	0
E2OPEN PARENT HOLDINGS, INC.	COM	29788T103	14,691,037	2,502,732	SH		SOLE		2,502,732	0	0
EHEALTH, INC.	COM	28238P109	2,748,147	567,799	SH		SOLE		567,799	0	0
ELECTRONIC ARTS	COM	285512109	1,947,060	15,936	SH		SOLE		15,936	0	0
ENCORE CAPITAL GROUP, INC.	COM	292554102	223,832	4,669	SH		SOLE		4,669	0	0
ENOVA INTERNATIONAL, INC.	COM	29357K103	7,146,835	186,261	SH		SOLE		186,261	0	0
EURONET WORLDWIDE, INC.	COM	298736109	15,722,764	166,590	SH		SOLE		166,590	0	0
EXPEDIA GROUP, INC.	COM	30212P303	4,434,750	50,625	SH		SOLE		50,625	0	0
EXTREME NETWORKS, INC.	COM	30226D106	9,578,730	523,142	SH		SOLE		523,142	0	0
F5, INC.	COM	315616102	4,488,562	31,277	SH		SOLE		31,277	0	0
FARFETCH LIMITED	COM	30744W107	1,829,124	386,707	SH		SOLE		386,707	0	0
FIDELITY NATIONAL INFORMATION SERVICES, INC.	COM	31620M106	4,586,660	67,600	SH		SOLE		67,600	0	0
FLEX LTD.	COM	Y2573F102	12,807,135	596,791	SH		SOLE		596,791	0	0
FULL HOUSE RESORTS, INC.	COM	359678109	3,064,039	407,452	SH		SOLE		407,452	0	0
GENIUS SPORTS LTD	COM	G3934V109	5,606,674	1,570,497	SH		SOLE		1,570,497	0	0
GREEN PLAINS, INC.	COM	393222104	2,869,684	94,088	SH		SOLE		94,088	0	0
GRIFFON CORPORATION	COM	398433102	5,594,872	156,325	SH		SOLE		156,325	0	0
H&E EQUIPMENT SERVICES, INC.	COM	404030108	5,512,377	121,418	SH		SOLE		121,418	0	0
HARROW HEALTH, INC.	COM	415858109	14,354,277	972,512	SH		SOLE		972,512	0	0
HARVARD BIOSCIENCE, INC.	COM	416906105	2,920,411	1,054,300	SH		SOLE		1,054,300	0	0
HELIX ENERGY SOLUTNS GROUP, INC.	COM	42330P107	1,922,158	260,455	SH		SOLE		260,455	0	0
HONEST COMPANY, INC.	COM	438333106	9,360,378	3,109,760	SH		SOLE		3,109,760	0	0
HUDSON PACIFIC PROPERTIES, INC.	COM	444097109	2,124,682	218,364	SH		SOLE		218,364	0	0
I3 VERTICALS, INC.	COM	46571Y107	19,804,290	813,652	SH		SOLE		813,652	0	0
ICAD, INC.	COM	44934S206	600,560	328,175	SH		SOLE		328,175	0	0
IDENTIV, INC.	COM	45170X205	15,393,768	2,126,211	SH		SOLE		2,126,211	0	0
IMMERSION CORPORATION	COM	452521107	7,530,873	1,071,248	SH		SOLE		1,071,248	0	0
INTERNATIONAL MONEY EXPRESS, INC.	COM	46005L101	22,308,054	915,390	SH		SOLE		915,390	0	0
INTERNATIONAL SEAWAYS, INC.	COM	Y41053102	5,223,374	141,096	SH		SOLE		141,096	0	0
JAKKS PACIFIC, INC.	COM	47012E403	2,007,939	114,805	SH		SOLE		114,805	0	0
KURA SUSHI USA, INC.	COM	501270102	15,894,796	333,364	SH		SOLE		333,364	0	0
L3HARRIS TECHNOLOGIES, INC.	COM	502431109	4,888,354	23,478	SH		SOLE		23,478	0	0
LANTRONIX, INC.	COM	516548203	2,985,630	691,118	SH		SOLE		691,118	0	0
LAREDO PETROLEUM, INC.	COM	516806205	6,793,353	132,115	SH		SOLE		132,115	0	0
LEIDOS HOLDINGS, INC.	COM	525327102	4,528,219	43,048	SH		SOLE		43,048	0	0
LSB INDUSTRIES, INC.	COM	502160104	9,034,278	679,269	SH		SOLE		679,269	0	0
MARKFORGED HOLDING CORPORATION	COM	57064N102	2,282,205	1,967,418	SH		SOLE		1,967,418	0	0
MATCH GROUP, INC.	COM	57667L107	295,533	7,123	SH		SOLE		7,123	0	0
MERITAGE HOMES CORPORATION	COM	59001A102	3,654,624	39,638	SH		SOLE		39,638	0	0
META PLATFORMS, INC.	COM	30303M102	15,997,879	132,939	SH		SOLE		132,939	0	0
MGP INGREDIENTS, INC.	COM	55303J106	1,395,918	13,122	SH		SOLE		13,122	0	0
MICROSOFT CORPORATION	COM	594918104	2,452,160	10,225	SH		SOLE		10,225	0	0
MODINE MANUFACTURING COMPANY	COM	607828100	6,048,284	304,546	SH		SOLE		304,546	0	0
NAVIGATOR HOLDINGS LTD.	COM	Y62132108	6,849,624	572,711	SH		SOLE		572,711	0	0
NAVIOS MARITIME PARTNERS L.P.	COM	Y62267409	4,316,473	166,274	SH		SOLE		166,274	0	0
NOBLE CORPORATION PLC	COM	G65431127	8,450,736	224,098	SH		SOLE		224,098	0	0
NOMAD FOODS LTD	COM	G6564A105	4,568,600	265,000	SH		SOLE		265,000	0	0
NORTHERN OIL AND GAS, INC.	COM	665531307	11,851,923	384,553	SH		SOLE		384,553	0	0
OKTA, INC.	COM	679295105	3,556,372	52,047	SH		SOLE		52,047	0	0
OLIN CORPORATION	COM	680665205	6,566,836	124,043	SH		SOLE		124,043	0	0
ORACLE CORPORATION	COM	68389X105	9,091,940	111,230	SH		SOLE		111,230	0	0
OVERSEAS SHIPHOLDING GROUP, INC.	COM	69036R863	2,437,319	843,363	SH		SOLE		843,363	0	0
OXFORD INDUSTRIES, INC.	COM	691497309	4,144,087	44,474	SH		SOLE		44,474	0	0
PAR PACIFIC HOLDINGS, INC.	COM	69888T207	2,272,292	97,733	SH		SOLE		97,733	0	0
PAYONEER GLOBAL, INC.	COM	70451X104	4,330,965	791,767	SH		SOLE		791,767	0	0
PBF ENERGY, INC.	COM	69318G106	7,661,379	187,871	SH		SOLE		187,871	0	0
PORCH GROUP, INC.	COM	733245104	11,921,264	6,341,098	SH		SOLE		6,341,098	0	0
PROG HOLDINGS, INC.	COM	74319R101	3,482,650	206,196	SH		SOLE		206,196	0	0
QUALYS, INC.	COM	74758T303	4,120,524	36,715	SH		SOLE		36,715	0	0
QUANTERIX CORPORATION	COM	74766Q101	11,941,816	862,225	SH		SOLE		862,225	0	0
QUINSTREET, INC.	COM	74874Q100	6,139,231	427,821	SH		SOLE		427,821	0	0
REGIONS FINANCIAL CORPORATION	COM	7591EP100	7,190,195	333,497	SH		SOLE		333,497	0	0
REPAY HOLDINGS CORPORATION	COM	76029L100	19,837,108	2,464,237	SH		SOLE		2,464,237	0	0
RICHARDSON ELECTRONICS LTD	COM	763165107	10,337,201	484,632	SH		SOLE		484,632	0	0
RIMINI STREET, INC.	COM	76674Q107	2,664,188	699,262	SH		SOLE		699,262	0	0
SANDRIDGE ENERGY, INC.	COM	80007P869	427,095	25,079	SH		SOLE		25,079	0	0
SCIENCE APPLICATIONS INTERNATIONAL CORPORATION	COM	808625107	7,012,440	63,215	SH		SOLE		63,215	0	0
SIMILARWEB LTD	COM	M84137104	5,711,023	888,184	SH		SOLE		888,184	0	0
SITIME CORPORATION	COM	82982T106	2,261,045	22,250	SH		SOLE		22,250	0	0
SKECHERS USA, INC.	COM	830566105	14,927,404	355,838	SH		SOLE		355,838	0	0
SKYLINE CHAMPION CORPORATION	COM	830830105	2,807,501	54,504	SH		SOLE		54,504	0	0
SONY GROUP CORPORATION	COM	835699307	6,149,083	80,612	SH		SOLE		80,612	0	0
STRIDE, INC.	COM	86333M108	2,431,050	77,719	SH		SOLE		77,719	0	0
SUNOPTA, INC.	COM	8676EP108	15,040,207	1,782,015	SH		SOLE		1,782,015	0	0
SUPER MICRO COMPUTER, INC.	COM	86800U104	14,644,013	178,368	SH		SOLE		178,368	0	0
SVB FINANCIAL GROUP	COM	78486Q101	6,824,572	29,654	SH		SOLE		29,654	0	0
TAKE TWO INTERACTIVE SOFTWARE, INC.	COM	874054109	9,319,323	89,497	SH		SOLE		89,497	0	0
TECNOGLASS, INC.	COM	G87264100	24,981,363	811,874	SH		SOLE		811,874	0	0
TEEKAY TANKERS LTD.	COM	Y8565N300	2,689,929	87,307	SH		SOLE		87,307	0	0
TITAN MACHINERY, INC.	COM	88830R101	3,721,390	93,667	SH		SOLE		93,667	0	0
TRAVELCENTERS OF AMERICA, INC.	COM	89421B109	17,849,666	398,608	SH		SOLE		398,608	0	0
TRUECAR, INC.	COM	89785L107	6,007,291	2,393,343	SH		SOLE		2,393,343	0	0
TWILIO, INC.	COM	90138F102	16,700,599	341,107	SH		SOLE		341,107	0	0
UBS GROUP AG	COM	H42097107	4,910,154	262,997	SH		SOLE		262,997	0	0
UNITED THERAPEUTICS CORPORATION	COM	91307C102	21,918,220	78,817	SH		SOLE		78,817	0	0
V2X, INC.	COM	92242T101	4,835,472	117,110	SH		SOLE		117,110	0	0
VELO3D, INC.	COM	92259N104	1,501,976	839,093	SH		SOLE		839,093	0	0
VISTRA CORP.	COM	92840M102	13,131,223	566,001	SH		SOLE		566,001	0	0
WALKME LTD.	COM	M97628107	8,880,028	794,278	SH		SOLE		794,278	0	0
WALLBOX NV	COM	N94209108	1,825,299	509,860	SH		SOLE		509,860	0	0
WEST FRASER TIMBER CO. LTD.	COM	952845105	5,199,241	71,922	SH		SOLE		71,922	0	0
WM TECHNOLOGY, INC.	COM	92971A109	1,522,409	1,507,336	SH		SOLE		1,507,336	0	0
WORKDAY, INC.	COM	98138H101	6,781,048	40,525	SH		SOLE		40,525	0	0