Exhibit 99.1
Joint Filing Agreement
In accordance with Rule 13d-1(k)(1)(iii) under the Securities Exchange Act of 1934, as amended, the persons named below agree to the joint filing on behalf of each of them of a Statement on Schedule 13G dated January 3, 2019 (including amendments thereto) with respect to the Common Stock, $0.01 par value, of Miragen Therapeutics, Inc. This Joint Filing Agreement shall be filed as an Exhibit to such Statement.
| Dated: January 3, 2019 | |||
| 683 CAPITAL MANAGEMENT, LLC | |||
| By: | /s/ Ari Zweiman | ||
| Name: | Ari Zweiman | ||
| Title: | Authorized Person | ||
| 683 CAPITAL PARTNERS, LP | |||
| By: | /s/ Ari Zweiman | ||
| Name: | Ari Zweiman | ||
| Title: | Authorized Person | ||
| By: | /s/ Ari Zweiman | ||
| Name: | Ari Zweiman | ||
