Exhibit 99.1
JOINT FILING AGREEMENT
We, the undersigned, hereby express our agreement that the attached Schedule 13G is filed on behalf of each of us.
Dated: October 10, 2019
THE COLUMN GROUP, LP | THE COLUMN GROUP GP, LP | |||||
By: | The Column Group GP, LP | |||||
By: | /s/ Peter Svennilson | By: | /s/ Peter Svennilson | |||
Name: | Peter Svennilson | Name: | Peter Svennilson | |||
Title: | Managing Partner | Title: | Managing Partner | |||
PONOI CAPITAL, LP | PONOI MANAGEMENT, LLC | |||||
By: | Ponoi Management, LLC | |||||
By: | /s/ Peter Svennilson | By: | /s/ Peter Svennilson | |||
Name: | Peter Svennilson | Name: | Peter Svennilson | |||
Title: | Managing Partner | Title: | Managing Partner | |||
PONOI CAPITAL II, LP | PONOI II MANAGEMENT, LLC | |||||
By: | Ponoi II Management, LLC | |||||
By: | /s/ Peter Svennilson | By: | /s/ Peter Svennilson | |||
Name: | Peter Svennilson | Name: | Peter Svennilson | |||
Title: | Managing Partner | Title: | Managing Partner | |||
By: | /s/ Peter Svennilson | |||||
Peter Svennilson | ||||||
By: | /s/ David V. Goeddel | |||||
David V. Goeddel | ||||||
By: | /s/ Tim Kutzkey | |||||
Tim Kutzkey |