EXHIBIT A
Joint Filing Statement
We, the undersigned, hereby express our agreement that the attached Schedule 13D/A is filed on behalf of each of us.
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Dated as September 22, 2016 | | |
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THE COLUMN GROUP, LP | | | | THE COLUMN GROUP GP, LP |
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By: | | The Column Group GP, LP | | | | | | |
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By: | | /s/ Peter Svennilson | | | | By: | | /s/ Peter Svennilson |
Name: | | Peter Svennilson | | | | Name: | | Peter Svennilson |
Title: | | Managing Partner | | | | Title: | | Managing Partner |
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THE COLUMN GROUP II, LP | | | | THE COLUMN GROUP II GP, LP |
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By: | | The Column Group II GP, LP | | | | | | |
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By: | | /s/ Peter Svennilson | | | | By: | | /s/ Peter Svennilson |
Name: | | Peter Svennilson | | | | Name: | | Peter Svennilson |
Title: | | Managing Partner | | | | Title: | | Managing Partner |
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By: | | /s/ Peter Svennilson | | | | | | |
| | Peter Svennilson | | | | | | |
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By: | | /s/ David V. Goeddel | | | | | | |
| | David V. Goeddel | | | | | | |