Quanterix (QTRX) 10-K/A 23 Dec 24 2023 FY Annual report (amended) Financial data

Document and Entity Information

Document and Entity Information - USD ($) $ in Millions	12 Months Ended
	Dec. 31, 2023	Feb. 26, 2024	Jun. 30, 2023
Cover [Abstract]
Document Type	10-K/A
Document Annual Report	true
Document Period End Date	Dec. 31, 2023
Document Transition Report	false
Entity File Number	001-38319
Entity Registrant Name	Quanterix Corp
Entity Incorporation, State or Country Code	DE
Entity Tax Identification Number	20-8957988
Entity Address, Address Line One	900 Middlesex Turnpike
Entity Address, City or Town	Billerica
Entity Address, State or Province	MA
Entity Address, Postal Zip Code	01821
City Area Code	617
Local Phone Number	301-9400
Title of 12(g) Security	Common Stock, $0.001 par value per share
Trading Symbol	QTRX
Security Exchange Name	NASDAQ
Entity Well-known Seasoned Issuer	Yes
Entity Voluntary Filers	No
Entity Current Reporting Status	No
Entity Interactive Data Current	Yes
Entity Filer Category	Large Accelerated Filer
Entity Small Business	false
Entity Emerging Growth Company	false
Entity Shell Company	false
ICFR Auditor Attestation Flag	true
Document Financial Statement Error Correction [Flag]	false
Entity Public Float			$ 796
Entity Common Stock, Shares Outstanding		38,063,618
Entity Central Index Key	0001503274
Current Fiscal Year End Date	--12-31
Document Fiscal Year Focus	2023
Document Fiscal Period Focus	FY
Amendment Flag	true
Auditor Name	Ernst & Young LLP
Auditor Firm ID	42
Auditor Location	Boston, Massachusetts

CONSOLIDATED BALANCE SHEETS

CONSOLIDATED BALANCE SHEETS - USD ($) $ in Thousands	Dec. 31, 2023	Sep. 30, 2023	Jun. 30, 2023	Mar. 31, 2023	Dec. 31, 2022	Sep. 30, 2022	Jun. 30, 2022	Mar. 31, 2022	Dec. 31, 2021	Dec. 31, 2020
Current assets:
Cash and cash equivalents	$ 174,422	$ 201,261	$ 329,525	$ 329,354	$ 338,740	$ 343,743	$ 361,551	$ 374,543
Marketable securities	146,902	126,449			0
Accounts receivable, net of allowance for expected credit losses	25,414	24,083	24,369	22,546	19,017	18,330	19,683	22,616
Inventory	26,123	20,857	18,333	16,629	16,964	18,044	22,685	24,426
Prepaid expenses and other current assets	9,234	9,216	6,897	6,945	6,803	6,075	9,979	13,878
Total current assets	382,095	381,866	379,124	375,474	381,524	386,192	413,898	435,463
Restricted cash	2,604	2,602	2,601	2,598	2,597	2,596	2,594	2,577
Property and equipment, net	17,926	17,287	18,048	18,690	19,811	21,195	21,976	19,639
Intangible assets, net	6,034	6,003	6,476	7,129	7,516	7,511	8,527	9,692
Operating lease right-of-use assets	18,251	19,860	20,380	20,891	21,223	27,165	32,935	29,915
Other non-current assets	1,657	1,603	1,637	938	911	910	77	78
Total assets	428,567	429,221	428,266	425,720	433,582	445,569	488,682	506,687
Current liabilities:
Accounts payable	5,048	4,786	3,238	2,585	3,836	2,106	5,908	4,187
Accrued compensation and benefits	14,170	10,505	8,175	5,411	11,035	10,760	10,777	8,196
Accrued expenses and other current liabilities	6,055	6,686	6,791		5,147
Deferred revenue	9,468	9,827	10,421	10,682	8,644	8,976	12,157	9,194
Operating lease liabilities	4,241	4,093	3,986	3,875	2,687	767	2,696	1,886
Total current liabilities	38,982	35,897	32,611	27,482	31,349	28,918	36,733	31,755
Deferred revenue, net of current portion	1,227	1,126	1,304	1,419	1,415	1,591	1,531	1,222
Operating lease liabilities, net of current portion	37,223	38,306	39,378	40,409	41,417	42,196	43,135	43,563
Other non-current liabilities	1,177	1,105	1,225	1,216	1,469	1,570	1,810	1,691
Total liabilities	78,609	76,434	74,518	70,526	75,650	74,275	83,209	78,231
Commitments and contingencies (Note 16)
Stockholders’ equity:
Common stock	38	38	37	37	37	37	37	37
Additional paid-in capital	783,142	778,615	772,472	768,123	763,629	759,254	756,087	750,717
Accumulated other comprehensive loss	(1,672)	(3,129)	(2,740)	(2,496)	(2,538)	(2,999)	(2,203)	(756)
Accumulated deficit	(431,550)	(422,737)	(416,021)	(410,470)	(403,196)	(384,998)	(348,448)	(321,542)
Total stockholders’ equity	349,958	352,787	353,748	355,194	357,932	371,294	405,473	428,456	$ 442,792	$ 205,825
Total liabilities and stockholders’ equity	$ 428,567	$ 429,221	$ 428,266	$ 425,720	$ 433,582	$ 445,569	$ 488,682	$ 506,687

CONSOLIDATED BALANCE SHEETS (Pa

CONSOLIDATED BALANCE SHEETS (Parenthetical) - USD ($) $ in Thousands	Dec. 31, 2023	Sep. 30, 2023	Jun. 30, 2023	Mar. 31, 2023	Dec. 31, 2022	Sep. 30, 2022	Jun. 30, 2022	Mar. 31, 2022
Statement of Financial Position [Abstract]
Common stock, par value	$ 0.001	$ 0.001	$ 0.001	$ 0.001	$ 0.001	$ 0.001	$ 0.001	$ 0.001
Common stock, authorized shares	120,000,000	120,000,000	120,000,000	120,000,000	120,000,000	120,000,000	120,000,000	120,000,000
Common stock, shares issued	38,014,000	37,839,000	37,566,000	37,424,000	37,280,000	37,094,000	36,975,000	36,899,000
Common stock, shares outstanding	38,014,000	37,839,000	37,566,000	37,424,000	37,280,000	37,094,000	36,975,000	36,899,000
Marketable securities	$ 146,902	$ 126,449			$ 0

CONSOLIDATED STATEMENTS OF OPER

CONSOLIDATED STATEMENTS OF OPERATIONS - USD ($) $ in Thousands	3 Months Ended								6 Months Ended		9 Months Ended		12 Months Ended
	Dec. 31, 2023	Sep. 30, 2023	Jun. 30, 2023	Mar. 31, 2023	Dec. 31, 2022	Sep. 30, 2022	Jun. 30, 2022	Mar. 31, 2022	Jun. 30, 2023	Jun. 30, 2022	Sep. 30, 2023	Sep. 30, 2022	Dec. 31, 2023	Dec. 31, 2022	Dec. 31, 2021
Revenues:
Total revenues	$ 31,549	$ 31,559	$ 30,804	$ 28,456	$ 25,824	$ 26,646	$ 23,500	$ 29,552	$ 59,260	$ 53,052	$ 90,819	$ 79,698	$ 122,368	$ 105,522	$ 110,556
Costs of goods sold and services:
Total costs of goods sold and services	12,143	12,150	11,388	12,463	13,180	16,341	15,183	15,455	23,851	30,638	36,001	46,979	48,144	60,159	45,784
Gross profit	19,406	19,409	19,416	15,993	12,644	10,305	8,317	14,097	35,409	22,414	54,818	32,719	74,224	45,363	64,772
Operating expenses:
Research and development	7,210	7,739	6,128	4,987	5,843	6,829	7,257	6,880	11,115	14,137	18,854	20,966	26,064	26,809	28,383
Selling, general and administrative	23,436	23,550	21,308	20,817	18,385	20,479	27,816	25,171	42,125	52,987	65,675	73,466	89,111	91,851	92,910
Other lease costs	1,016	928	968	800	802	609			1,768		2,696	609	3,712	1,411	0
Impairment and restructuring	1,361	0	0	(33)	9,006	20,343	207		(33)	207	(33)	20,550	1,328	29,556	0
Total operating expenses	33,023	32,217	28,404	26,571	34,036	48,260	35,280	32,051	54,975	67,331	87,192	115,591	120,215	149,627	121,293
Loss from operations	(13,617)	(12,808)	(8,988)	(10,578)	(21,392)	(37,955)	(26,963)	(17,954)	(19,566)	(44,917)	(32,374)	(82,872)	(45,991)	(104,264)	(56,521)
Interest income (expense), net	4,319	4,185	3,886	3,449	2,815	1,712	552	52	7,335	604	11,520	2,316	15,839	5,131	(403)
Other income (expense), net	626	2,110	(214)	(5)	533	(235)	(358)	(217)	(219)	(575)	1,891	(810)	2,517	(277)	1,340
Loss before income taxes	(8,672)	(6,513)	(5,316)	(7,134)	(18,044)	(36,478)	(26,769)	(18,119)	(12,450)	(44,888)	(18,963)	(81,366)	(27,635)	(99,410)	(55,584)
Income tax (expense) benefit	(141)	(203)	(235)	(140)	(154)	(72)	(137)	199	(375)	62	(578)	(10)	(719)	(164)	36
Net loss	$ (8,813)	$ (6,716)	$ (5,551)	$ (7,274)	$ (18,198)	$ (36,550)	$ (26,906)	$ (17,920)	$ (12,825)	$ (44,826)	$ (19,541)	$ (81,376)	$ (28,354)	$ (99,574)	$ (55,548)
Net loss per common share, basic (in dollars per share)	$ (0.23)	$ (0.18)	$ (0.15)	$ (0.19)	$ (0.49)	$ (0.99)	$ (0.73)	$ (0.49)	$ (0.34)	$ (1.22)	$ (0.52)	$ (2.20)	$ (0.75)	$ (2.69)	$ (1.54)
Net loss per common share, diluted (in dollars per share)	$ (0.23)	$ (0.18)	$ (0.15)	$ (0.19)	$ (0.49)	$ (0.99)	$ (0.73)	$ (0.49)	$ (0.34)	$ (1.22)	$ (0.52)	$ (2.20)	$ (0.75)	$ (2.69)	$ (1.54)
Weighted-average common shares outstanding, basic (in shares)	37,890,000	37,657,000	37,494,000	37,327,000	37,160,000	37,005,000	36,922,000	36,851,000	37,411,000	36,887,000	37,494,000	36,927,000	37,594,000	36,991,000	35,997,000
Weighted-average common shares outstanding, diluted (in shares)	37,890,000	37,657,000	37,494,000	37,327,000	37,160,000	37,005,000	36,922,000	36,851,000	37,411,000	36,887,000	37,494,000	36,927,000	37,594,000	36,991,000	35,997,000
Product revenue
Revenues:
Total revenues	$ 20,821	$ 19,940	$ 19,549	$ 19,360	$ 16,674	$ 17,693	$ 14,785	$ 20,656	$ 38,909	$ 35,441	$ 58,849	$ 53,134	$ 79,670	$ 69,808	$ 81,062
Costs of goods sold and services:
Total costs of goods sold and services	7,314	7,209	6,701	7,879	9,491	11,347	10,546	11,457	14,580	22,003	21,789	33,350	29,103	42,841	32,114
Service and other revenue
Revenues:
Total revenues	10,230	10,883	10,470	8,506	8,767	8,370	8,548	8,810	18,976	17,358	29,859	25,728	40,089	34,495	23,629
Costs of goods sold and services:
Total costs of goods sold and services	4,829	4,941	4,687	4,584	3,689	4,994	4,637	3,998	9,271	8,635	14,212	13,629	19,041	17,318	13,670
Collaboration and license revenue
Revenues:
Total revenues	146	237	629	368	170	301	92	$ 86	997	178	1,234	479	1,380	649	648
Grant revenue
Revenues:
Total revenues	$ 352	$ 499	$ 156	$ 222	$ 213	$ 282	$ 75		$ 378	$ 75	$ 877	$ 357	$ 1,229	$ 570	$ 5,217

CONSOLIDATED STATEMENTS OF COMP

CONSOLIDATED STATEMENTS OF COMPREHENSIVE LOSS - USD ($) $ in Thousands	3 Months Ended								6 Months Ended		9 Months Ended		12 Months Ended
	Dec. 31, 2023	Sep. 30, 2023	Jun. 30, 2023	Mar. 31, 2023	Dec. 31, 2022	Sep. 30, 2022	Jun. 30, 2022	Mar. 31, 2022	Jun. 30, 2023	Jun. 30, 2022	Sep. 30, 2023	Sep. 30, 2022	Dec. 31, 2023	Dec. 31, 2022	Dec. 31, 2021
Statement of Comprehensive Income [Abstract]
Net loss	$ (8,813)	$ (6,716)	$ (5,551)	$ (7,274)	$ (18,198)	$ (36,550)	$ (26,906)	$ (17,920)	$ (12,825)	$ (44,826)	$ (19,541)	$ (81,376)	$ (28,354)	$ (99,574)	$ (55,548)
Other comprehensive income (loss), net of tax:
Unrealized gains (losses) on marketable securities		(241)									(241)		325	0	0
Foreign currency translation adjustment		(148)	(244)	42		(796)	(1,447)	(1,197)	(202)	(2,644)	(350)	(3,440)	541	(2,979)	(1,993)
Total other comprehensive income (loss)		(389)	(244)	42		(796)	(1,447)	(1,197)	(202)	(2,644)	(591)	(3,440)	866	(2,979)	(1,993)
Comprehensive loss		$ (7,105)	$ (5,795)	$ (7,232)		$ (37,346)	$ (28,353)	$ (19,117)	$ (13,027)	$ (47,470)	$ (20,132)	$ (84,816)	$ (27,488)	$ (102,553)	$ (57,541)

CONSOLIDATED STATEMENTS OF STOC

CONSOLIDATED STATEMENTS OF STOCKHOLDERS' EQUITY - USD ($) shares in Thousands, $ in Thousands	Total	Common Stock	Additional paid-in capital	Accumulated other comprehensive income (loss)	Accumulated deficit
Beginning balance (in shares) at Dec. 31, 2020		31,797
Beginning Balance at Dec. 31, 2020	$ 205,825	$ 32	$ 451,433	$ 2,434	$ (248,074)
Increase (Decrease) in Stockholders' Equity
Issuance of common stock under stock plans, including tax effects (in shares)		864
Issuance of common stock under stock plans, including tax effects	8,815	$ 1	8,814
Sale of common stock in underwritten public offering, net (in shares)		4,107
Sale of common stock in underwritten public offering, net	269,718	$ 4	269,714
Stock-based compensation expense	15,975		15,975
Unrealized gains on marketable securities, net of tax	0
Foreign currency translation adjustment	(1,993)			(1,993)
Net loss, as restated	(55,548)				(55,548)
Ending Balance (in shares) at Dec. 31, 2021		36,768
Ending Balance at Dec. 31, 2021	442,792	$ 37	745,936	441	(303,622)
Increase (Decrease) in Stockholders' Equity
Issuance of common stock under stock plans, including tax effects (in shares)		131
Issuance of common stock under stock plans, including tax effects	979		979
Stock-based compensation expense	3,802		3,802
Foreign currency translation adjustment	(1,197)			(1,197)
Net loss, as restated	(17,920)				(17,920)
Ending Balance (in shares) at Mar. 31, 2022		36,899
Ending Balance at Mar. 31, 2022	428,456	$ 37	750,717	(756)	(321,542)
Beginning balance (in shares) at Dec. 31, 2021		36,768
Beginning Balance at Dec. 31, 2021	442,792	$ 37	745,936	441	(303,622)
Increase (Decrease) in Stockholders' Equity
Issuance of common stock under stock plans, including tax effects (in shares)		207
Issuance of common stock under stock plans, including tax effects	1,190		1,190
Stock-based compensation expense	8,961		8,961
Foreign currency translation adjustment	(2,644)			(2,644)
Net loss, as restated	(44,826)				(44,826)
Ending Balance (in shares) at Jun. 30, 2022		36,975
Ending Balance at Jun. 30, 2022	405,473	$ 37	756,087	(2,203)	(348,448)
Beginning balance (in shares) at Dec. 31, 2021		36,768
Beginning Balance at Dec. 31, 2021	442,792	$ 37	745,936	441	(303,622)
Increase (Decrease) in Stockholders' Equity
Issuance of common stock under stock plans, including tax effects (in shares)		326
Issuance of common stock under stock plans, including tax effects	1,597		1,597
Stock-based compensation expense	11,721		11,721
Foreign currency translation adjustment	(3,440)			(3,440)
Net loss, as restated	(81,376)				(81,376)
Ending Balance (in shares) at Sep. 30, 2022		37,094
Ending Balance at Sep. 30, 2022	371,294	$ 37	759,254	(2,999)	(384,998)
Beginning balance (in shares) at Dec. 31, 2021		36,768
Beginning Balance at Dec. 31, 2021	442,792	$ 37	745,936	441	(303,622)
Increase (Decrease) in Stockholders' Equity
Issuance of common stock under stock plans, including tax effects (in shares)		512
Issuance of common stock under stock plans, including tax effects	2,310		2,310
Stock-based compensation expense	15,383		15,383
Unrealized gains on marketable securities, net of tax	0
Foreign currency translation adjustment	(2,979)			(2,979)
Net loss, as restated	(99,574)				(99,574)
Ending Balance (in shares) at Dec. 31, 2022		37,280
Ending Balance at Dec. 31, 2022	357,932	$ 37	763,629	(2,538)	(403,196)
Beginning balance (in shares) at Mar. 31, 2022		36,899
Beginning Balance at Mar. 31, 2022	428,456	$ 37	750,717	(756)	(321,542)
Increase (Decrease) in Stockholders' Equity
Issuance of common stock under stock plans, including tax effects (in shares)		76
Issuance of common stock under stock plans, including tax effects	211		211
Stock-based compensation expense	5,159		5,159
Foreign currency translation adjustment	(1,447)			(1,447)
Net loss, as restated	(26,906)				(26,906)
Ending Balance (in shares) at Jun. 30, 2022		36,975
Ending Balance at Jun. 30, 2022	405,473	$ 37	756,087	(2,203)	(348,448)
Increase (Decrease) in Stockholders' Equity
Issuance of common stock under stock plans, including tax effects (in shares)		119
Issuance of common stock under stock plans, including tax effects	407		407
Stock-based compensation expense	2,760		2,760
Foreign currency translation adjustment	(796)			(796)
Net loss, as restated	(36,550)				(36,550)
Ending Balance (in shares) at Sep. 30, 2022		37,094
Ending Balance at Sep. 30, 2022	371,294	$ 37	759,254	(2,999)	(384,998)
Increase (Decrease) in Stockholders' Equity
Net loss, as restated	(18,198)
Ending Balance (in shares) at Dec. 31, 2022		37,280
Ending Balance at Dec. 31, 2022	357,932	$ 37	763,629	(2,538)	(403,196)
Increase (Decrease) in Stockholders' Equity
Issuance of common stock under stock plans, including tax effects (in shares)		144
Issuance of common stock under stock plans, including tax effects	551		551
Stock-based compensation expense	3,943		3,943
Foreign currency translation adjustment	42			42
Net loss, as restated	(7,274)				(7,274)
Ending Balance (in shares) at Mar. 31, 2023		37,424
Ending Balance at Mar. 31, 2023	355,194	$ 37	768,123	(2,496)	(410,470)
Beginning balance (in shares) at Dec. 31, 2022		37,280
Beginning Balance at Dec. 31, 2022	357,932	$ 37	763,629	(2,538)	(403,196)
Increase (Decrease) in Stockholders' Equity
Issuance of common stock under stock plans, including tax effects (in shares)		286
Issuance of common stock under stock plans, including tax effects	690		690
Stock-based compensation expense	8,153		8,153
Foreign currency translation adjustment	(202)			(202)
Net loss, as restated	(12,825)				(12,825)
Ending Balance (in shares) at Jun. 30, 2023		37,566
Ending Balance at Jun. 30, 2023	353,748	$ 37	772,472	(2,740)	(416,021)
Beginning balance (in shares) at Dec. 31, 2022		37,280
Beginning Balance at Dec. 31, 2022	357,932	$ 37	763,629	(2,538)	(403,196)
Increase (Decrease) in Stockholders' Equity
Issuance of common stock under stock plans, including tax effects (in shares)		559
Issuance of common stock under stock plans, including tax effects	2,490	$ 1	2,489
Stock-based compensation expense	12,497		12,497
Unrealized gains on marketable securities, net of tax	(241)			(241)
Foreign currency translation adjustment	(350)			(350)
Net loss, as restated	(19,541)				(19,541)
Ending Balance (in shares) at Sep. 30, 2023		37,839
Ending Balance at Sep. 30, 2023	352,787	$ 38	778,615	(3,129)	(422,737)
Beginning balance (in shares) at Dec. 31, 2022		37,280
Beginning Balance at Dec. 31, 2022	357,932	$ 37	763,629	(2,538)	(403,196)
Increase (Decrease) in Stockholders' Equity
Issuance of common stock under stock plans, including tax effects (in shares)		734
Issuance of common stock under stock plans, including tax effects	2,691	$ 1	2,690
Stock-based compensation expense	16,823		16,823
Unrealized gains on marketable securities, net of tax	325			325
Foreign currency translation adjustment	541			541
Net loss, as restated	(28,354)				(28,354)
Ending Balance (in shares) at Dec. 31, 2023		38,014
Ending Balance at Dec. 31, 2023	349,958	$ 38	783,142	(1,672)	(431,550)
Beginning balance (in shares) at Mar. 31, 2023		37,424
Beginning Balance at Mar. 31, 2023	355,194	$ 37	768,123	(2,496)	(410,470)
Increase (Decrease) in Stockholders' Equity
Issuance of common stock under stock plans, including tax effects (in shares)		142
Issuance of common stock under stock plans, including tax effects	139		139
Stock-based compensation expense	4,210		4,210
Foreign currency translation adjustment	(244)			(244)
Net loss, as restated	(5,551)				(5,551)
Ending Balance (in shares) at Jun. 30, 2023		37,566
Ending Balance at Jun. 30, 2023	353,748	$ 37	772,472	(2,740)	(416,021)
Increase (Decrease) in Stockholders' Equity
Issuance of common stock under stock plans, including tax effects (in shares)		273
Issuance of common stock under stock plans, including tax effects	1,800	$ 1	1,799
Stock-based compensation expense	4,344		4,344
Unrealized gains on marketable securities, net of tax	(241)			(241)
Foreign currency translation adjustment	(148)			(148)
Net loss, as restated	(6,716)				(6,716)
Ending Balance (in shares) at Sep. 30, 2023		37,839
Ending Balance at Sep. 30, 2023	352,787	$ 38	778,615	(3,129)	(422,737)
Increase (Decrease) in Stockholders' Equity
Net loss, as restated	(8,813)
Ending Balance (in shares) at Dec. 31, 2023		38,014
Ending Balance at Dec. 31, 2023	$ 349,958	$ 38	$ 783,142	$ (1,672)	$ (431,550)

CONSOLIDATED STATEMENT OF CASH

CONSOLIDATED STATEMENT OF CASH FLOWS - USD ($) $ in Thousands	3 Months Ended		6 Months Ended		9 Months Ended		12 Months Ended
	Mar. 31, 2023	Mar. 31, 2022	Jun. 30, 2023	Jun. 30, 2022	Sep. 30, 2023	Sep. 30, 2022	Dec. 31, 2023	Dec. 31, 2022	Dec. 31, 2021
Cash flows from operating activities:
Net loss	$ (7,274)	$ (17,920)	$ (12,825)	$ (44,826)	$ (19,541)	$ (81,376)	$ (28,354)	$ (99,574)	$ (55,548)
Adjustments to reconcile net loss to net cash used in operating activities:
Depreciation and amortization expense	1,454	1,373	2,883	2,820	4,843	4,231	6,275	5,409	4,880
Credit losses on accounts receivable	178	(171)	324	581	311	102	336	(301)	213
Accretion of marketable securities					(1,249)		(1,964)	0	0
Operating lease right-of-use asset amortization			1,002		1,518		2,015	715	499
Stock-based compensation expense	3,943	3,802	8,153	8,961	12,497	11,721	16,823	15,383	15,975
Impairment			54	207	130		1,361	25,801	0
Other operating activity			602				(150)	(340)	692
Changes in assets and liabilities:
Accounts receivable	(3,741)	1,319	(5,750)	3,479	(5,615)	5,045	(6,695)	5,156	(6,853)
Inventory	530	(206)	(1,180)	1,515	(3,700)	6,146	(8,944)	7,243	(10,125)
Prepaid expenses and other current assets	(422)	(2,071)	(527)	961	(2,829)	(282)	(2,371)	(738)	(166)
Other non-current assets	(33)	1	(707)	2	(702)	(857)	(717)	(822)	(2)
Accounts payable	(1,271)	(5,306)	(631)	(3,220)	948	(7,010)	1,189	(5,287)	2,339
Accrued compensation and benefits, accrued expenses, and other current liabilities	(5,809)	(4,864)	(1,053)	(4,248)	1,229	(2,914)	4,410	(3,599)	606
Deferred revenue	2,041	2,956	1,666	6,228	894	3,108	635	2,599	1,462
Operating lease liabilities	179	(87)	(730)	(3,128)	(1,690)	(1,156)	(2,645)	(266)	(1,230)
Other non-current liabilities	(203)	(271)	(72)	(9)	(107)	128	(53)	10	(363)
Net cash used in operating activities	(9,824)	(21,755)	(8,791)	(29,716)	(12,657)	(44,521)	(18,849)	(48,611)	(47,621)
Cash flows from investing activities:
Purchases of marketable debt securities					(125,200)		(175,613)	0	0
Proceeds from marketable debt securities							31,000	0	0
Purchases of property and equipment	(136)	(1,335)	(947)	(6,342)	(1,878)	(10,019)	(3,841)	(11,614)	(13,675)
Proceeds from RADx grant on assets purchased		520		520		520	0	520	7,278
Net cash used in investing activities	(136)	(815)	(947)	(5,822)	(127,078)	(9,499)	(148,454)	(11,094)	(6,397)
Cash flows from financing activities:
Proceeds from common stock issued under stock plans	551	594	777	594	2,632	881	2,889	2,311	8,815
Payments for employee taxes withheld on stock-based compensation awards	(13)		(87)		(142)		(198)	0	0
Payments on notes payable							0	0	(7,738)
Net cash provided by financing activities	551	979	690	1,190	2,490	1,597	2,691	2,311	270,795
Net increase (decrease) in cash, cash equivalents, and restricted cash	(9,409)	(21,591)	(9,048)	(34,348)	(137,245)	(52,423)	(164,612)	(57,394)	216,777
Effect of exchange rate changes on cash, cash equivalents, and restricted cash	24	(558)	(163)	(776)	(229)	(507)	301	(538)	(92)
Cash, cash equivalents, and restricted cash at beginning of period	341,337	399,269	341,337	399,269	341,337	399,269	341,337	399,269	182,584
Cash, cash equivalents, and restricted cash at end of period	331,952	377,120	332,126	$ 364,145	203,863	346,339	177,026	341,337	399,269
Supplemental disclosure of cash flow information:
Cash paid for taxes	246		$ 502		$ 719	263	808	684	0
Cash paid for interest							0	0	389
Purchases of property and equipment in accounts payable and accruals	$ 147					288	419	152	229
Operating lease right-of-use assets obtained in exchange for lease liabilities		$ 18,156				$ 22,239	0	22,494	0
Shares received as consideration under product sales agreement (Note 4, 7)							775	0	0
Underwritten public offering
Cash flows from financing activities:
Sale of common stock in underwritten public offering, net							$ 0	$ 0	$ 269,718

Restatement of Financial Statem

Restatement of Financial Statements - Balance Sheet (Parenthetical) - $ / shares	Dec. 31, 2023	Sep. 30, 2023	Jun. 30, 2023	Mar. 31, 2023	Dec. 31, 2022	Sep. 30, 2022	Jun. 30, 2022	Mar. 31, 2022
Statement of Financial Position [Abstract]
Common stock, par value	$ 0.001	$ 0.001	$ 0.001	$ 0.001	$ 0.001	$ 0.001	$ 0.001	$ 0.001
Common stock, authorized shares	120,000,000	120,000,000	120,000,000	120,000,000	120,000,000	120,000,000	120,000,000	120,000,000
Common stock, shares issued	38,014,000	37,839,000	37,566,000	37,424,000	37,280,000	37,094,000	36,975,000	36,899,000
Common stock, shares outstanding	38,014,000	37,839,000	37,566,000	37,424,000	37,280,000	37,094,000	36,975,000	36,899,000

Restatement of Financial Stat_2

Restatement of Financial Statements	12 Months Ended
	Dec. 31, 2023
Accounting Changes and Error Corrections [Abstract]
Restatement of Financial Statements	Note 1. Restatement of Financial Statements Subsequent to the issuance of Quanterix Corporation's (“Quanterix” or the “Company”) Annual Report on Form 10-K for the fiscal year ended December 31, 2023, originally filed with the U.S. Securities and Exchange Commission (the “SEC”) on February 29, 2024 (the “Original Report"), the Company identified an error related to the capitalization of labor and overhead costs in the Company's inventory balances (the "Misstatement"), which impacted the previously issued audited Consolidated Financial Statements as of December 31, 2023 and 2022 and for each of the three years in the period ended December, 31 2023 and the unaudited Consolidated Financial Statements for the quarterly and year-to-date (as applicable) periods ended March 31, 2022, June 30, 2022, September 30, 2022, March 31, 2023, June 30, 2023, and September 30, 2023 (the "Restatement Periods"). In accordance with ASC 250 - Accounting Changes and Error Corrections , SEC Staff Accounting Bulletin ("SAB") No. 99 - Materiality , and SAB No. 108 - Considering the Effects of Prior Year Misstatements when Quantifying Misstatements in Current Year Financial Statements , management concluded the error was material to the Company’s financial statements and required restatement of the financial statements for the Restatement Periods (the "Restatement"). In connection with the Restatement, the Company is also correcting unrelated errors identified by the Company in prior periods. Restatement Background In connection with the Company's efforts to remediate a material weakness in its internal control over financial reporting relating to the operating effectiveness of internal controls associated with the accounting for inventory valuation, and while performing closing procedures for the third quarter of 2024, the Company identified the Misstatement. The correction of the Misstatement impacts the previously reported amounts of inventory, cost of product revenue, net loss per common share, and all related financial statement subtotals and totals. In addition, the correction of unrelated errors identified by the Company in prior periods includes, but is not limited to, adjustments to property and equipment, accrued compensation and benefits, and operating expenses (together with the Misstatement, the "Combined Misstatements"). Impact of Restatement The following tables present the impact of the Restatement to the specific line items presented in the previously reported audited Consolidated Financial Statements in all Restatement Periods. The amounts labeled "As Previously Reported" were derived from the Original Report. The amounts labeled “Adjustments” represent the impact of correcting the Combined Misstatements identified by the Company. The effects of the Restatement have been corrected in all impacted tables and footnotes throughout the audited Consolidated Financial Statements. QUANTERIX CORPORATION RESTATED CONSOLIDATED BALANCE SHEETS (amounts in thousands, except per share data) As of December 31, 2023 As Previously Reported Adjustments As Restated ASSETS Current assets: Cash and cash equivalents $ 174,422 $ — $ 174,422 Marketable securities 146,902 — 146,902 Accounts receivable, net of allowance for expected credit losses 25,414 — 25,414 Inventory 22,365 3,758 26,123 Prepaid expenses and other current assets 9,291 (57) 9,234 Total current assets 378,394 3,701 382,095 Restricted cash 2,604 — 2,604 Property and equipment, net 17,926 — 17,926 Intangible assets, net 6,034 — 6,034 Operating lease right-of-use assets 18,251 — 18,251 Other non-current assets 1,802 (145) 1,657 Total assets $ 425,011 $ 3,556 $ 428,567 LIABILITIES AND STOCKHOLDERS' EQUITY Current liabilities: Accounts payable $ 5,048 $ — $ 5,048 Accrued compensation and benefits 13,659 511 14,170 Accrued expenses and other current liabilities 6,041 14 6,055 Deferred revenue 9,468 — 9,468 Operating lease liabilities 4,241 — 4,241 Total current liabilities 38,457 525 38,982 Deferred revenue, net of current portion 1,227 — 1,227 Operating lease liabilities, net of current portion 37,223 — 37,223 Other non-current liabilities 1,177 — 1,177 Total liabilities 78,084 525 78,609 Commitments and contingencies (Note 16) Stockholders’ equity: Common stock, $0.001 par value per share: Authorized: 120,000 shares; Issued and outstanding: 38,014 shares at December 31, 2023 38 — 38 Additional paid-in capital 783,142 — 783,142 Accumulated other comprehensive loss (1,757) 85 (1,672) Accumulated deficit (434,496) 2,946 (431,550) Total stockholders’ equity 346,927 3,031 349,958 Total liabilities and stockholders’ equity $ 425,011 $ 3,556 $ 428,567 As of December 31, 2022 As Previously Reported Adjustments As Restated ASSETS Current assets: Cash and cash equivalents $ 338,740 $ — $ 338,740 Accounts receivable, net of allowance for expected credit losses 19,017 — 19,017 Inventory 16,786 178 16,964 Prepaid expenses and other current assets 6,860 (57) 6,803 Total current assets 381,403 121 381,524 Restricted cash 2,597 — 2,597 Property and equipment, net 20,162 (351) 19,811 Intangible assets, net 7,516 — 7,516 Operating lease right-of-use assets 21,223 — 21,223 Other non-current assets 1,298 (387) 911 Total assets $ 434,199 $ (617) $ 433,582 LIABILITIES AND STOCKHOLDERS' EQUITY Current liabilities: Accounts payable $ 3,836 $ — $ 3,836 Accrued compensation and benefits 10,658 377 11,035 Accrued expenses and other current liabilities 5,133 14 5,147 Deferred revenue 8,644 — 8,644 Operating lease liabilities 2,687 — 2,687 Total current liabilities 30,958 391 31,349 Deferred revenue, net of current portion 1,415 — 1,415 Operating lease liabilities, net of current portion 41,417 — 41,417 Other non-current liabilities 1,469 — 1,469 Total liabilities 75,259 391 75,650 Commitments and contingencies (Note 16) Stockholders’ equity: Common stock, $0.001 par value per share: Authorized: 120,000 shares; Issued and outstanding: 37,280 shares at December 31, 2022 37 — 37 Additional paid-in capital 763,688 (59) 763,629 Accumulated other comprehensive loss (2,623) 85 (2,538) Accumulated deficit (402,162) (1,034) (403,196) Total stockholders’ equity 358,940 (1,008) 357,932 Total liabilities and stockholders’ equity $ 434,199 $ (617) $ 433,582 QUANTERIX CORPORATION RESTATED CONSOLIDATED STATEMENTS OF OPERATIONS (amounts in thousands, except per share data) Year Ended December 31, 2023 As Previously Reported Adjustments As Restated Revenues: Product revenue $ 79,460 $ 210 $ 79,670 Service and other revenue 40,299 (210) 40,089 Collaboration and license revenue 1,380 — 1,380 Grant revenue 1,229 — 1,229 Total revenues 122,368 — 122,368 Costs of goods sold and services: Cost of product revenue 32,636 (3,533) 29,103 Cost of service and other revenue 19,086 (45) 19,041 Total costs of goods sold and services 51,722 (3,578) 48,144 Gross profit 70,646 3,578 74,224 Operating expenses: Research and development 24,857 1,207 26,064 Selling, general and administrative 90,241 (1,130) 89,111 Other lease costs 3,712 — 3,712 Impairment and restructuring 1,537 (209) 1,328 Total operating expenses 120,347 (132) 120,215 Loss from operations (49,701) 3,710 (45,991) Interest income 15,839 — 15,839 Other income 2,247 270 2,517 Loss before income taxes (31,615) 3,980 (27,635) Income tax expense (719) — (719) Net loss $ (32,334) $ 3,980 $ (28,354) Net loss per common share, basic and diluted $ (0.86) $ 0.11 $ (0.75) Weighted-average common shares outstanding, basic and diluted 37,594 — 37,594 Year Ended December 31, 2022 As Previously Reported Adjustments As Restated Revenues: Product revenue $ 69,808 $ — $ 69,808 Service and other revenue 34,495 — 34,495 Collaboration and license revenue 649 — 649 Grant revenue 570 — 570 Total revenues 105,522 — 105,522 Costs of goods sold and services: Cost of product revenue 40,809 2,032 42,841 Cost of service and other revenue 17,907 (589) 17,318 Total costs of goods sold and services 58,716 1,443 60,159 Gross profit 46,806 (1,443) 45,363 Operating expenses: Research and development 25,890 919 26,809 Selling, general and administrative 91,995 (144) 91,851 Other lease costs 1,278 133 1,411 Impairment and restructuring 29,347 209 29,556 Total operating expenses 148,510 1,117 149,627 Loss from operations (101,704) (2,560) (104,264) Interest income 5,131 — 5,131 Other expense (62) (215) (277) Loss before income taxes (96,635) (2,775) (99,410) Income tax expense (65) (99) (164) Net loss $ (96,700) $ (2,874) $ (99,574) Net loss per common share, basic and diluted $ (2.61) $ (0.08) $ (2.69) Weighted-average common shares outstanding, basic and diluted 36,991 — 36,991 Year Ended December 31, 2021 As Previously Reported Adjustments As Restated Revenues: Product revenue $ 81,062 $ — $ 81,062 Service and other revenue 23,629 — 23,629 Collaboration and license revenue 648 — 648 Grant revenue 5,217 — 5,217 Total revenues 110,556 — 110,556 Costs of goods sold and services: Cost of product revenue 34,149 (2,035) 32,114 Cost of service and other revenue 14,679 (1,009) 13,670 Total costs of goods sold and services 48,828 (3,044) 45,784 Gross profit 61,728 3,044 64,772 Operating expenses: Research and development 27,978 405 28,383 Selling, general and administrative 92,336 574 92,910 Total operating expenses 120,314 979 121,293 Loss from operations (58,586) 2,065 (56,521) Interest expense (403) — (403) Other income 1,265 75 1,340 Loss before income taxes (57,724) 2,140 (55,584) Income tax benefit 36 — 36 Net loss $ (57,688) $ 2,140 $ (55,548) Net loss per common share, basic and diluted $ (1.60) $ 0.06 $ (1.54) Weighted-average common shares outstanding, basic and diluted 35,997 — 35,997 QUANTERIX CORPORATION RESTATED CONSOLIDATED STATEMENTS OF COMPREHENSIVE LOSS (amounts in thousands) Year Ended December 31, 2023 As Previously Reported Adjustments As Restated Net loss $ (32,334) $ 3,980 $ (28,354) Other comprehensive income (loss), net of tax: Unrealized gains on marketable securities 325 — 325 Foreign currency translation adjustment 541 — 541 Total other comprehensive income 866 — 866 Comprehensive loss $ (31,468) $ 3,980 $ (27,488) Year Ended December 31, 2022 As Previously Reported Adjustments As Restated Net loss $ (96,700) $ (2,874) $ (99,574) Other comprehensive income (loss), net of tax: Foreign currency translation adjustment (3,064) 85 (2,979) Total other comprehensive income (loss) (3,064) 85 (2,979) Comprehensive loss $ (99,764) $ (2,789) $ (102,553) Year Ended December 31, 2021 As Previously Reported Adjustments As Restated Net loss $ (57,688) $ 2,140 $ (55,548) Other comprehensive income (loss), net of tax: Foreign currency translation adjustment (1,993) — (1,993) Total other comprehensive loss (1,993) — (1,993) Comprehensive loss $ (59,681) $ 2,140 $ (57,541) QUANTERIX CORPORATION RESTATED CONSOLIDATED STATEMENTS OF STOCKHOLDERS’ EQUITY (amounts in thousands) Year Ended December 31, 2023 Common Stock As Previously Reported Shares Amount Additional paid-in capital Accumulated other comprehensive income (loss) Accumulated Total stockholders' equity Balance at December 31, 2022 37,280 $ 37 $ 763,688 $ (2,623) $ (402,162) $ 358,940 Issuance of common stock under stock plans, including tax effects 734 1 2,690 — — 2,691 Stock-based compensation expense — — 16,764 — — 16,764 Unrealized gains on marketable securities, net of tax — — — 325 — 325 Foreign currency translation adjustment — — — 541 — 541 Net loss — — — — (32,334) (32,334) Balance at December 31, 2023 38,014 $ 38 $ 783,142 $ (1,757) $ (434,496) $ 346,927 Adjustments Balance at December 31, 2022 — $ — $ (59) $ 85 $ (1,034) $ (1,008) Issuance of common stock under stock plans, including tax effects — — — — — — Stock-based compensation expense — — 59 — — 59 Unrealized gains on marketable securities, net of tax — — — — — — Foreign currency translation adjustment — — — — — — Net loss — — — — 3,980 3,980 Balance at December 31, 2023 — $ — $ — $ 85 $ 2,946 $ 3,031 As Restated Balance at December 31, 2022 37,280 $ 37 $ 763,629 $ (2,538) $ (403,196) $ 357,932 Issuance of common stock under stock plans, including tax effects 734 1 2,690 — — 2,691 Stock-based compensation expense — — 16,823 — — 16,823 Unrealized gains on marketable securities, net of tax — — — 325 — 325 Foreign currency translation adjustment — — — 541 — 541 Net loss — — — — (28,354) (28,354) Balance at December 31, 2023 38,014 $ 38 $ 783,142 $ (1,672) $ (431,550) $ 349,958 Year Ended December 31, 2022 Common Stock As Previously Reported Shares Amount Additional paid-in capital Accumulated other comprehensive income (loss) Accumulated Total stockholders' equity Balance at December 31, 2021 36,768 $ 37 $ 745,936 $ 441 $ (305,462) $ 440,952 Issuance of common stock under stock plans, including tax effects 512 — 2,310 — — 2,310 Stock-based compensation expense — — 15,442 — — 15,442 Foreign currency translation adjustment — — — (3,064) — (3,064) Net loss — — — — (96,700) (96,700) Balance at December 31, 2022 37,280 $ 37 $ 763,688 $ (2,623) $ (402,162) $ 358,940 Adjustments Balance at December 31, 2021 — $ — $ — $ — $ 1,840 $ 1,840 Issuance of common stock under stock plans, including tax effects — — — — — — Stock-based compensation expense — — (59) — — (59) Foreign currency translation adjustment — — — 85 — 85 Net loss — — — — (2,874) (2,874) Balance at December 31, 2022 — $ — $ (59) $ 85 $ (1,034) $ (1,008) As Restated Balance at December 31, 2021 36,768 $ 37 $ 745,936 $ 441 $ (303,622) $ 442,792 Issuance of common stock under stock plans, including tax effects 512 — 2,310 — — 2,310 Stock-based compensation expense — — 15,383 — — 15,383 Foreign currency translation adjustment — — — (2,979) — (2,979) Net loss — — — — (99,574) (99,574) Balance at December 31, 2022 37,280 $ 37 $ 763,629 $ (2,538) $ (403,196) $ 357,932 Year Ended December 31, 2021 Common Stock As Previously Reported Shares Amount Additional paid-in capital Accumulated other comprehensive income (loss) Accumulated Total stockholders' equity Balance at December 31, 2020 31,797 $ 32 $ 451,433 $ 2,434 $ (247,774) $ 206,125 Issuance of common stock under stock plans, including tax effects 864 1 8,814 — — 8,815 Sale of common stock in underwritten public offering, net 4,107 4 269,714 — — 269,718 Stock-based compensation expense — — 15,975 — — 15,975 Foreign currency translation adjustment — — — (1,993) — (1,993) Net loss — — — — (57,688) (57,688) Balance at December 31, 2021 36,768 $ 37 $ 745,936 $ 441 $ (305,462) $ 440,952 Adjustments Balance at December 31, 2020 — $ — $ — $ — $ (300) $ (300) Issuance of common stock under stock plans, including tax effects — — — — — — Sale of common stock in underwritten public offering, net — — — — — — Stock-based compensation expense — — — — — — Foreign currency translation adjustment — — — — — — Net loss — — — — 2,140 2,140 Balance at December 31, 2021 — $ — $ — $ — $ 1,840 $ 1,840 As Restated Balance at December 31, 2020 31,797 $ 32 $ 451,433 $ 2,434 $ (248,074) $ 205,825 Issuance of common stock under stock plans, including tax effects 864 1 8,814 — — 8,815 Sale of common stock in underwritten public offering, net 4,107 4 269,714 — — 269,718 Stock-based compensation expense — — 15,975 — — 15,975 Foreign currency translation adjustment — — — (1,993) — (1,993) Net loss — — — — (55,548) (55,548) Balance at December 31, 2021 36,768 $ 37 $ 745,936 $ 441 $ (303,622) $ 442,792 QUANTERIX CORPORATION RESTATED CONSOLIDATED STATEMENTS OF CASH FLOWS (amounts in thousands) Year Ended December 31, 2023 As Previously Reported Adjustments As Restated Cash flows from operating activities: Net loss $ (32,334) $ 3,980 $ (28,354) Adjustments to reconcile net loss to net cash used in operating activities: Depreciation and amortization expense 6,364 (89) 6,275 Credit losses on accounts receivable 336 — 336 Accretion of marketable securities (1,964) — (1,964) Operating lease right-of-use asset amortization 2,015 — 2,015 Stock-based compensation expense 16,764 59 16,823 Impairment 1,570 (209) 1,361 Other operating activity 150 (300) (150) Changes in assets and liabilities: Accounts receivable (6,695) — (6,695) Inventory (5,364) (3,580) (8,944) Prepaid expenses and other current assets (2,371) — (2,371) Other non-current assets (775) 58 (717) Accounts payable 1,189 — 1,189 Accrued compensation and benefits, accrued expenses, and other current liabilities 4,276 134 4,410 Deferred revenue 635 — 635 Operating lease liabilities (2,645) — (2,645) Other non-current liabilities (53) — (53) Net cash used in operating activities (18,902) 53 (18,849) Cash flows from investing activities: Purchases of marketable debt securities (175,613) — (175,613) Proceeds from marketable debt securities 31,000 — 31,000 Purchases of property and equipment (3,788) (53) (3,841) Net cash used in investing activities (148,401) (53) (148,454) Cash flows from financing activities: Proceeds from common stock issued under stock plans 2,889 — 2,889 Payments for employee taxes withheld on stock-based compensation awards (198) — (198) Net cash provided by financing activities 2,691 — 2,691 Net increase (decrease) in cash, cash equivalents, and restricted cash (164,612) — (164,612) Effect of exchange rate changes on cash, cash equivalents, and restricted cash 301 — 301 Cash, cash equivalents, and restricted cash at beginning of period 341,337 — 341,337 Cash, cash equivalents, and restricted cash at end of period $ 177,026 $ — $ 177,026 Supplemental disclosure of cash flow information: Cash paid for taxes $ 808 $ — $ 808 Purchases of property and equipment in accounts payable and accruals $ 419 $ — $ 419 Shares received as consideration under product sales agreement (Note 4, 7) $ 775 $ — $ 775 Year Ended December 31, 2022 As Previously Reported Adjustments As Restated Cash flows from operating activities: Net loss $ (96,700) $ (2,874) $ (99,574) Adjustments to reconcile net loss to net cash used in operating activities: Depreciation and amortization expense 5,349 60 5,409 Credit losses on accounts receivable (301) — (301) Operating lease right-of-use asset amortization 715 — 715 Stock-based compensation expense 15,442 (59) 15,383 Impairment 25,592 209 25,801 Other operating activity (439) 99 (340) Changes in assets and liabilities: Accounts receivable 5,156 — 5,156 Inventory 5,386 1,857 7,243 Prepaid expenses and other current assets (568) (170) (738) Other non-current assets (909) 87 (822) Accounts payable (5,362) 75 (5,287) Accrued compensation and benefits, accrued expenses, and other current liabilities (3,976) 377 (3,599) Deferred revenue 2,599 — 2,599 Operating lease liabilities (266) — (266) Other non-current liabilities 10 — 10 Net cash used in operating activities (48,272) (339) (48,611) Cash flows from investing activities: Purchases of property and equipment (11,726) 112 (11,614) Proceeds from RADx grant on assets purchased 520 — 520 Net cash used in investing activities (11,206) 112 (11,094) Cash flows from financing activities: Proceeds from common stock issued under stock plans 2,311 — 2,311 Net cash provided by financing activities 2,311 — 2,311 Net increase (decrease) in cash, cash equivalents, and restricted cash (57,167) (227) (57,394) Effect of exchange rate changes on cash, cash equivalents, and restricted cash (538) — (538) Cash, cash equivalents, and restricted cash at beginning of period 399,042 227 399,269 Cash, cash equivalents, and restricted cash at end of period $ 341,337 $ — $ 341,337 Supplemental disclosure of cash flow information: Cash paid for taxes $ 684 $ — $ 684 Purchases of property and equipment in accounts payable and accruals $ 152 $ — $ 152 Operating lease right-of-use assets obtained in exchange for lease liabilities $ 22,494 $ — $ 22,494 Year Ended December 31, 2021 As Previously Reported Adjustments As Restated Cash flows from operating activities: Net loss $ (57,688) $ 2,140 $ (55,548) Adjustments to reconcile net loss to net cash used in operating activities: Depreciation and amortization expense 4,851 29 4,880 Credit losses on accounts receivable 213 — 213 Operating lease right-of-use asset amortization 499 — 499 Stock-based compensation expense 15,975 — 15,975 Other operating activity 692 — 692 Changes in assets and liabilities: Accounts receivable (6,853) — (6,853) Inventory (8,090) (2,035) (10,125) Prepaid expenses and other current assets (393) 227 (166) Other non-current assets (2) — (2) Accounts payable 2,414 (75) 2,339 Accrued compensation and benefits, accrued expenses, and other current liabilities 606 — 606 Deferred revenue 1,462 — 1,462 Operating lease liabilities (1,230) — (1,230) Other non-current liabilities (363) — (363) Net cash used in operating activities (47,907) 286 (47,621) Cash flows from investing activities: Purchases of marketable debt securities — — — Proceeds from marketable debt securities — — — Purchases of property and equipment (13,616) (59) (13,675) Proceeds from RADx grant on assets purchased 7,278 — 7,278 Net cash used in investing activities (6,338) (59) (6,397) Cash flows from financing activities: Proceeds from common stock issued under stock plans 8,815 — 8,815 Sale of common stock in underwritten public offering, net 269,718 — 269,718 Payments on notes payable (7,738) — (7,738) Net cash provided by financing activities 270,795 — 270,795 Net increase (decrease) in cash, cash equivalents, and restricted cash 216,550 227 216,777 Effect of exchange rate changes on cash, cash equivalents, and restricted cash (92) — (92) Cash, cash equivalents, and restricted cash at beginning of period 182,584 — 182,584 Cash, cash equivalents, and restricted cash at end of period $ 399,042 $ 227 $ 399,269 Supplemental disclosure of cash flow information: Cash paid for interest $ 389 $ — $ 389 Purchases of property and equipment in accounts payable and accruals $ 229 $ — $ 229

Organization and Nature of Busi

Organization and Nature of Business	12 Months Ended
	Dec. 31, 2023
Organization, Consolidation and Presentation of Financial Statements [Abstract]
Organization and Nature of Business	Note 2. Organization and Nature of Business Quanterix is a life sciences company that has developed next-generation, ultra-sensitive digital immunoassay platforms that advance life sciences research and diagnostics. The Company’s platforms are based on its proprietary digital “Simoa” detection technology and enable customers to reliably detect protein biomarkers in ultra- low concentrations in blood, serum, and other fluids that, in many cases, are undetectable using conventional, analog immunoassay technologies. The ability of the Company’s Simoa platforms to detect proteins in the femtomolar range is enabling the development of novel therapies and diagnostics and has the potential to facilitate a paradigm shift in healthcare from an emphasis on treatment to a focus on earlier detection, monitoring, prognosis, and, ultimately, prevention. The Company also provides contract research services for customers and Laboratory Developed Test (“LDT”) services through its CLIA-certified Accelerator Laboratory (the “Accelerator Laboratory”). The Accelerator Laboratory provides customers with access to Simoa technology and supports multiple projects and services, including sample testing, homebrew assay development, custom assay development, and blood-based biomarker testing. To date, the Company has completed over 2,200 projects for more than 480 customers from all over the world using its Simoa platforms. Liquidity (As Restated) The Company has recognized annual losses from operations since inception and has an accumulated deficit of $431.6 million as of December 31, 2023. The Company incurred net losses of $28.4 million, $99.6 million, and $55.5 million for the years ended December 31, 2023, 2022, and 2021, respectively. As of December 31, 2023, the Company had cash and cash equivalents of $174.4 million and marketable securities of $146.9 million. The Company expects that its current cash, cash equivalents, and marketable securities will be sufficient to fund its operations for a period of at least one year from the date the Consolidated Financial Statements are issued.

Significant Accounting Policies

Significant Accounting Policies	12 Months Ended
	Dec. 31, 2023
Accounting Policies [Abstract]
Significant Accounting Policies	Note 3. Significant Accounting Policies Basis of Presentation The accompanying Consolidated Financial Statements and notes herein have been prepared in accordance with generally accepted accounting principles in the United States of America (“U.S. GAAP”) and pursuant to the rules and regulations of the U.S. Securities and Exchange Commission (“SEC”) regarding annual financial reporting on Form 10-K. The Company’s fiscal year is the twelve-month period from January 1 through December 31, and all references to “2023,” “2022,” and “2021,” refer to the fiscal year unless otherwise noted. Certain amounts in the prior years’ Consolidated Financial Statements have been reclassified to conform to the current year’s presentation. Use of Estimates The preparation of the Consolidated Financial Statements and Notes to Consolidated Financial Statements in conformity with U.S. GAAP requires management to make estimates and assumptions that affect the reported amounts of assets and liabilities at the dates of the Consolidated Financial Statements, and the reported amounts of revenues and expenses during each fiscal year. Such estimates include, but are not limited to, revenue recognition, valuation of inventory, leases, valuation and impairment of long-lived assets, recoverability of deferred tax assets, and stock-based compensation expense. The Company bases its estimates on historical experience, known trends, worldwide economic conditions, both general and specific to the life sciences industry, and other relevant factors it believes to be reasonable under the circumstances. On an ongoing basis, management evaluates its estimates and changes in estimates are recorded in the period in which they become known. Actual results could differ from those estimates. Principles of Consolidation The Consolidated Financial Statements include the accounts of Quanterix and its wholly-owned subsidiaries. All intercompany transactions have been eliminated in consolidation. In accordance with Accounting Standards Codification (“ASC”) 810 – Consolidation , the Company assesses the terms of its non-marketable equity investments in entities to determine if any meet the definition of a variable interest entity (“VIE”) and require consolidation into its Consolidated Financial Statements. Refer to Note 20 - Variable Interest Entities for further discussion. Foreign Currency (As Restated) The functional currency of the Company’s subsidiaries is their respective local currencies. These subsidiary financial statements are translated into U.S. dollars using the period-end exchange rates for assets and liabilities, average exchange rates during the corresponding period for revenue and expenses, and historical rates for equity. The effects of foreign currency translation adjustments are recorded in accumulated other comprehensive income (loss), a component of stockholders’ equity on the Consolidated Balance Sheets. Foreign currency transaction gains (losses) are included in other income (expense), net on the Consolidated Statements of Operations. Foreign exchange losses were $0.2 million, $1.0 million and $0.4 million during the years ended December 31, 2023, 2022 and 2021, respectively. Segment Information Operating segments are defined as components of an enterprise about which separate discrete information is available for evaluation by the chief operating decision-maker (“CODM”), in deciding how to allocate resources and assess performance. The Company’s CODM is the chief executive officer, who reviews the Company's operations and manages its business as a single operating segment as of December 31, 2023. Revenue from Contracts with Customers The Company generates the majority of its revenues from contracts with customers and accounts for them pursuant to the Financial Accounting Standards Board (“FASB”) Accounting Standards Codification Topic 606 - Revenue from Contracts with Customers (“ASC 606”). Refer to the section titled “Grant Revenue” below for discussion the Company’s accounting policy for revenue generated from grant awards. For contracts with customers, the Company recognizes revenue when a customer obtains control of promised products or services, for an amount that reflects the consideration expected to be received in exchange for those products or services. The Company follows the five step revenue model prescribed by ASC 606 to determine revenue recognition: (i) identify the contract(s) with a customer; (ii) identify the performance obligations in the contract; (iii) determine the transaction price, including variable consideration, if any; (iv) allocate the transaction price to the performance obligations in the contract; and (v) recognize revenue when (or as) the entity satisfies a performance obligation. Revenues are presented net of any sales, value added, or similar taxes collected from customers and remitted to the government. The Company accounts for a contract when it has approval and commitment from both parties, the fees, payment terms and rights of the parties regarding the products or services to be transferred are identified, the contract has commercial substance, and it is probable that substantially all of the consideration for the products and services expected to be transferred is collectible. The Company applies judgment in determining the customer’s ability and intention to pay for services expected to be transferred, which is based on factors including the customer’s payment history, management’s ability to mitigate exposure to credit risk (for example, requiring payment in advance of the transfer of products or services, or the ability to stop transferring promised products or services in the event a customer fails to pay consideration when due), and experience selling to similarly situated customers. The Company’s contracts may include either a single promise (referred to as a performance obligation) to transfer a product or service, or a combination of multiple performance obligations to transfer products or services. The Company evaluates the existence of multiple performance obligations within its contracts by using judgment to determine if (1) the customer can benefit from each contractual promise on its own or together with readily available resources and (2) the transfer of each contractual promise is separately identifiable from other promises in a contract. When both criteria are met, each promise is accounted for as a separate performance obligation. Additionally, the Company has elected the practical expedient under ASC 606 to account for shipping and handling as an activity to fulfill a promise to transfer a product, and therefore does not evaluate whether shipping and handling activities are promised services to its customers. Contracts that include rights to additional products or services that are exercisable at a customer’s discretion are generally considered options. The Company assesses if these options provide a material right to the customer and if so, the material right is considered a performance obligation. The identification of material rights requires judgment to determine the value of the option to purchase additional products and services in relation to options that may be provided to, and prices paid by, customers in the normal course of business. Material rights are recognized when exercised by a customer or upon expiration of the right. The Company determines the transaction price of its contracts based on the amount of consideration it expects to be entitled to, which is generally equal to the contract amount. In some cases, contracts contain variable consideration which primarily relates to (1) sales- and usage-based royalties related to the license of intellectual property in collaboration and license contracts and (2) contracts with minimum purchase commitments. For sales and usage-based royalties, ASC 606 provides an exception to estimating variable consideration. Under this exception, the Company recognizes revenues from sales- or usage-based royalty revenue at the later of when the sales or usage occurs or the satisfaction (or partial satisfaction) of the performance obligation to which the royalty has been allocated. All other variable amounts are constrained to the minimum guaranteed contract amount so that a reversal of cumulative revenue does not occur in future periods. Once there is no longer uncertainty over a variable amount, any incremental fees the Company is entitled to are allocated to the related performance obligation(s). For contracts that contain multiple performance obligations, the Company allocates the transaction price among the performance obligations on a relative basis according to their standalone selling prices (“SSP”). Determining the SSP for performance obligations requires judgment and is based on factors including prices charged to customers in observable transactions, internal pricing objectives and list prices, and pricing of similar products. The Company uses a range of amounts to estimate SSP and has more than one range for certain products and services based on the geographic location of customers and sales channel. Product Revenue The Company’s product revenues are composed of instruments, assay kits, replacement parts, and other consumables such as reagents and antibodies. Products are sold directly to customers and are also sold through distributors in EMEA and Asia Pacific regions. Direct instrument sales include installation and an initial year of implied service-type warranties. The Company has determined that the instrument and installation are a combined performance obligation as the customer cannot benefit from the instrument without the installation and no other vendors can provide the installation of the Company’s specialized instruments. The implied service-type warranty is considered a separate performance obligation since a customer could benefit from it independently with readily available resources and is capable of being sold on its own. Sales of instruments to distributors include a license to import and resell the instruments. The Company has determined these distributor licenses are part of a combined performance obligation with the instrument as the distributor only benefits from the combination of the instrument and ability to resell it. Instrument sales may also be bundled with assays and other consumables, training, and/or an extended service warranty, each of which is considered a separate performance obligation. Product revenues for direct instrument sales to customers are recognized upon completion of the instrument’s installation. For instrument sales to distributors, revenue is recognized based on the agreed upon shipping terms (either upon shipment or delivery) as that is when title passes to the customer. Service Revenue Service revenues are composed of contract research services, initial year of implied service-type warranties, extended services warranty contracts, repair services, and other services such as training. Contract research services are provided through the Company’s Accelerator Laboratory and generally consist of fixed fee contracts. Revenues from contract research services are recognized at the point in time when the Company completes and delivers its research results on each individually completed study, or over time if the contractual provisions allow for the collection of transaction consideration for costs incurred plus a reasonable margin through the period of performance of the services. For contract research services recognized over time, the Company uses the output method based on the number of completed results. Revenues from other services are recognized at the point in time when the training or other services are delivered as the customer simultaneously receives and benefits from the services. Revenues from service-type warranties are recognized ratably over the contract service period. Collaboration and License Revenue Collaboration and license revenues are composed of revenue associated with licensing to third parties the Company’s technology, intellectual property, and know-how associated with our instruments and for related services. License arrangements consist of sales or usage-based fees and/or future royalties. Revenues are recognized at the point in time the license is delivered as the underlying license is considered functional intellectual property. Royalty revenues that are sales- or usage- based are recognized at the later of when the sales or usage occurs and the satisfaction (or partial satisfaction) of the performance obligation to which the royalty has been allocated. Contract Assets and Liabilities Accounts Receivable and Allowance for Credit Losses The Company is exposed to credit losses primarily through accounts receivable from sales of its products and services. Accounts receivable includes amounts billed and currently due from customers. Since the only condition for payment of the Company’s invoices is the passage of time, the Company records a receivable on the date the invoice is issued. Also included in accounts receivable are unbilled amounts resulting from revenue exceeding the amount billed to the customer, where the right to payment is unconditional. If the right to payment for services performed was conditional on something other than the passage of time, the unbilled amount would be recorded as a separate contract asset. The Company’s expected loss allowance methodology is developed using historical collection experience, current and future economic and market conditions, and a review of the current status of customers’ accounts receivable. Specific allowance amounts are established to record the appropriate provision for customers that have a higher probability of default. The Company monitors changes to the receivables balance on a timely basis, and balances are written off as they are determined to be uncollectable after collection efforts have been exhausted. Generally, the Company’s contracts are non-cancellable. For contracts that are cancellable by the customer, the Company does not record a receivable when it issues an invoice. The Company records accounts receivable on these contracts only up to the amount of revenue recognized but not yet collected. The Company’s payment terms vary by the type and location of the customer and the products or services offered. Payment from customers is generally required 30 to 45 days from date of shipment or satisfaction of the performance obligation. The Company does not provide financing arrangements to its customers. The Company refers to contract liabilities as deferred revenue on the Consolidated Balance Sheets. For deferred revenue, the Company applies the practical expedient under ASC 606 that allows for the exclusion of (1) contracts with original expected length of one year or less and (2) contracts for which revenue is recognized at the amount to which the Company has the right to invoice for services performed from the amount of remaining performance obligations disclosed. Costs to Obtain Contracts The Company capitalizes commissions paid to its sales representatives and related fringe benefit costs that are incremental to obtaining customer contracts. Capitalized commissions are recorded in prepaid expenses and other current assets and other non-current assets on the Consolidated Balance Sheets. These commissions are amortized over the life of the contract and are recorded in cost of goods sold and selling, general and administrative expense on the Consolidated Statements of Operations. The Company has elected the practical expedient allowing commissions with an amortization period of one year or less to be expensed as incurred. Commissions associated with instrument sales are generally earned when installation is complete and title to the instrument has transferred. Commissions associated with consumables sales are earned when title to the product transfers. Commissions associated with warranty and extended service contracts are earned upon booking. Warranties The Company provides an initial year of warranty and maintenance service related to its instruments sold directly to customers and sells extended warranty contracts for additional periods. The Company defers revenue associated with these warranty services and recognizes them ratably over the service period. Grant Revenue Accounting for grants does not fall under ASC 606, as the grantor will not benefit directly from the Company’s expansion or product development, and no products or services are transferred to the grantor. As there is no authoritative guidance under U.S. GAAP on accounting for grants to for profit business entities from government entities, the Company accounts for grants by analogy to International Accounting Standards Topic 20, Accounting for Government Grants and Disclosure of Government Assistance (“IAS 20”) and ASC Topic 958, Not for-Profit Entities (“ASC 958”). The decision to account for a grant under IAS 20 or ASC 958 is based on whether the grantor is a government entity. The Company records grants related to assets as a deduction in calculating the carrying value of the asset, and to record grants related to income separately on the Consolidated Statements of Operations on a gross basis as grant revenue. The related expenses are recorded on a gross basis within operating expenses. These methods are elections an entity can make under both IAS 20 and ASC 958. The Company recognizes grant revenue as the Company performs services under the arrangement when the funding is committed and as each grant’s activities are performed. The timing of revenue recognition and receipt of funding varies by grant and can be independent from performance of the related activities, such as an upfront payment of the award value, or subsequent to the Company’s requests for reimbursement for already performed activities (subject to the approval of the granting organization). Cost of Goods Sold and Services Cost of Product Revenue Cost of product revenue consists of manufacturing and assembly costs for instruments, related reagents, other consumables, contract manufacturer costs, personnel costs, royalties, overhead, and other direct costs related to product sales. Raw material part costs include inbound freight, shipping and handling costs associated with purchased goods, contract manufacturer costs, personnel costs, royalties, overhead and other direct costs related to product sales. Additionally included in cost of product revenue are royalty fees due to third parties from revenue generated by collaboration or license deals. Cost of Service and Other Revenue Cost of services and other revenue consists of direct costs associated with operating the Company’s Accelerator Laboratory on behalf of its customers, including raw materials, personnel costs, royalties, allocated overhead costs that include facility and other related costs, and other direct costs. Additional costs include costs related to warranties, and other costs of servicing equipment at customer sites. Research and Development Expenses Research and development expense consists primarily of personnel costs, research supplies, third-party development costs for new products, materials for prototypes, quality assurance, and allocated overhead costs that include facility and other related costs. The Company accounts for nonrefundable advance payments for products and services that will be used in future research and development activities as expense when the service has been performed or when the products have been received. For arrangements in which the Company receives funding from third parties for research and development activities (excluding the government sponsored arrangements), the Company assesses whether the arrangement is within the scope of ASC 730 – Research and Development . When the Company is entitled to receive reimbursements, does not have an obligation to repay, does not transfer products or services, and is the primary beneficiary of the activities, the Company records the reimbursable amounts as a reduction to research and development expense. Amounts reimbursed in excess of the cost incurred by the Company are recorded within other income (expense), net on the Consolidated Statements of Operations. Reimbursable amounts recorded as a reduction to research and development expenses were not material during the years ended December 31, 2023, 2022, and 2021. Amounts reimbursed in excess of costs incurred by the Company related to activities funded by third parties were $0.4 million during the year ended December 31, 2023. No reimbursable amounts were received during the years ended December 31, 2022, and 2021. Selling, General and Administrative Expenses Selling, general and administrative expense consists primarily of personnel costs for our sales and marketing, finance, legal, human resources, and general management teams, shipping and handling for product sales, other general and administrative costs, as well as professional services costs, such as marketing, advertising, legal and accounting services, and allocated overhead costs that include facility and other related costs. The classification of shipping and handling costs for product sales as selling, general and administrative expenses varies from company to company with some companies recording these as selling, general and administrative expenses and others recording such expenses within costs of goods sold for products. To the extent the classification of its shipping and handling costs differs from the reporting approach used by other companies, the Company’s gross margins may not be comparable with those reported by such other companies. Net Loss Per Share Basic net loss per common share attributable to common stockholders is calculated by dividing the loss attributable to common stockholders by the weighted-average number of common shares outstanding. Diluted net loss per common share attributable to common stockholders is calculated under the treasury stock method by dividing the loss attributable to common stockholders by the diluted weighted-average number of common shares outstanding. Diluted weighted-average shares outstanding reflect the dilutive effect, if any, of potential common shares issued, such as unvested common stock, unvested restricted stock units (“RSUs”), common stock options, and shares estimated to be purchased under the Company’s employee stock purchase plan (“ESPP”). During periods when the Company is in a net loss position, these potential common shares are excluded from the diluted net loss per common share attributable to common stockholders because their effect would be anti-dilutive. Accordingly, basic and diluted net loss per common share attributable to common stockholders were the same for all periods presented. Cash and Cash Equivalents Cash and cash equivalents consist of cash deposits and short-term, highly liquid marketable securities that are readily convertible into cash, with original maturities of three months or less. Cash and cash equivalents consist of the following (in thousands): As of December 31, 2023 2022 Cash $ 12,162 $ 32,643 Money market funds 155,367 306,097 Marketable securities 6,893 — Total cash and cash equivalents $ 174,422 $ 338,740 Restricted Cash The following table summarizes the period ending cash and cash equivalents as presented on the Consolidated Balance Sheets and the total cash, cash equivalents, and restricted cash as presented on the Consolidated Statements of Cash Flows (in thousands): As of December 31, 2023 2022 Cash and cash equivalents $ 174,422 $ 338,740 Restricted cash 2,604 2,597 Cash, cash equivalents, and restricted cash $ 177,026 $ 341,337 Restricted cash consists of collateral for a letter of credit issued as security for two of the Company’s leased facilities and to secure the Company’s corporate credit card program. The short-term or long-term classification is determined in accordance with the expiration of the underlying letter of credit and security. Marketable Securities The Company’s current portfolio of marketable securities is entirely debt securities and may at any time include commercial paper, U.S. Treasuries, corporate notes and bonds, U.S. Government agency bonds, certificates of deposit, and similar types of debt securities. Marketable debt securities with original maturities of three months or less at the time of purchase are recorded in cash equivalents on the Consolidated Balance Sheets as they are considered highly liquid and readily convertible into cash. All other marketable securities, including those with maturities beyond one year, are recorded as current assets on the Consolidated Balance Sheets based on their highly liquid nature and because such securities are available for use in current operations. The Company classifies its marketable securities as either held to maturity, available-for-sale, or trading at the time of purchase and re-evaluates such classification at each balance sheet date. All of the Company’s marketable securities are currently classified as available-for-sale as it may use them in current operations. Available-for-sale securities are recorded at fair value (refer to Note 7 - Fair Value of Financial Instruments ). Unrealized gains and losses (other than impairment or credit related losses) are recorded in accumulated other comprehensive income (loss), a component of stockholders’ equity on the Consolidated Balance Sheets. Realized gains and losses are determined using the specific identification method and are recorded in other income (expense), net on the Consolidated Statements of Operations. Quarterly, or more frequently if circumstances warrant, the Company monitors its marketable securities for impairment. In the event a security’s fair value is less than its amortized cost basis, the Company evaluates whether an impairment exists and if the impairment is a result of credit loss or other factors. For a security in an unrealized loss position, if the Company intends to sell the security in an unrealized loss position, or it is more likely than not that the Company will be required to sell the security before recovery of its amortized cost basis, an impairment loss equal to the difference between the security’s fair value and amortized cost basis is recorded in other income (expense), net. Additionally, the Company determines if a credit loss exists by considering information about the collectability of the security, current market conditions, and the issuer’s financial condition. If a decline in fair value is a result of a credit loss, an allowance for credit losses is recorded in other income (expense), net on the Consolidated Statement of Operations, limited to the portion attributed to the credit loss. The Company has also elected the practical expedient to separately present accrued interest receivable from its marketable securities balance. Such accrued interest is recorded in prepaid expenses and other current assets on the Consolidated Balance Sheets and is not included in the assessment and measurement of impairment of its marketable securities. Inventory Inventory consists of instruments, assays, and the materials required to manufacture instruments and assays. Inventory is stated at the lower of cost or net realizable value on a first-in, first-out (“FIFO”) basis and includes the cost of materials, labor, and manufacturing overhead. The Company analyzes its inventory levels on each reporting date for slow-moving, excess, and obsolete inventory, and inventory expected to expire prior to being used. These analyses require judgment and are based on factors including, but not limited to, recent historical activity, anticipated or forecasted demand for the Company’s products (developed through its planning and sales and marketing inputs, and market conditions). If the Company identifies adverse conditions exist, the carrying value of the inventory is reduced to its estimated net realizable value by providing estimated reserves for excess or obsolete inventory. The Company adjusts the reserves for excess or obsolete inventory and records additional inventory write downs based on unfavorable changes in estimated customer demand or actual market conditions that may differ from its projections. Property and Equipment Property and equipment, including leasehold improvements, are stated at cost, net of accumulated depreciation. These assets are depreciated over their estimated useful lives using the straight-line method. Expenditures for maintenance and repairs are charged to expense as incurred, whereas significant expenditures that extend the useful lives of existing assets are capitalized as additions to property and equipment. Depreciation is calculated based upon the following estimated useful lives: Estimated Useful Life Laboratory and manufacturing equipment 5 years Office furniture and equipment 7 years Computers and software 3 years Leasehold improvements Shorter of asset's life or remaining lease term Leases The Company enters into operating leases for office, laboratory, and manufacturing spaces, as well as office equipment, and determines whether an arrangement is a lease at inception of the arrangement. The Company accounts for a lease when it has the right to control the leased asset for a period of time, while obtaining substantially all of the assets’ economic benefits. Leases are recorded on the Consolidated Balance Sheets as operating lease right-of-use (“ROU”) assets and current or non-current operating lease liabilities, as applicable. All of the Company’s leases are classified as operating leases. Additionally, the Company does not separate lease and non-lease components. Operating lease ROU assets and lease liabilities are recognized at the lease commencement date based on the estimated present value of the future minimum lease payments over the lease term and any initial direct costs incurred. Initial direct costs are incremental costs of a lease that would not have been incurred had the lease not been executed. The discount rate used to determine the present value of the lease payments is the Company’s incremental borrowing rate on a collateralized basis for a similar term and amount, as generally an implicit rate in the lease is not readily determinable. To estimate its incremental borrowing rate, a credit rating applicable to the Company is estimated using a synthetic credit rating analysis since the Company does not currently have an agency-based credit rating. The Company’s lease agreements can contain lease and non-lease components. The Company accounts for the lease and fixed payments for non-lease components as a single lease component under ASC 842– Leases , which increases the amount of the ROU assets and lease liabilities. Most of the Company’s lease agreements also contain variable payments, primarily maintenance, utility, and other-related costs, which are expensed as incurred and not included in the measurement of the ROU assets and lease liabilities. Some of the Company’s leases contain options to extend or terminate the lease. When determining the lease term, these options are included in the measurement and recognition of the Company’s ROU assets and lease liabilities when it is reasonably certain that the Company will exercise the option(s). The Company considers various economic factors when making this determination, including, but not limited to, the significance of leasehold improvements incurred in the leased space, the difficulty in replacing the asset, underlying contractual obligations, and specific characteristics unique to a particular lease. Subsequent to entering into a lease arrangement, the Company reassesses the certainty of exercising options to extend or terminate a lease. When it becomes reasonably certain that the Company will exercise an option that was not included in the lease term, the Company accounts for the change in circumstances as a lease modification, which results in the remeasurement of the ROU asset and lease liability as of the modification date. Leases with a term of 12 months or less upon commencement are not recorded on the Consolidated Balance Sheets and are recorded to expense on a straight-line basis over the lease term. Impairment of Goodwill The Company assesses goodwill for impairment at the reporting unit level annually and whenever events occur or circumstances change that would more likely than not reduce the fair value of a reporting unit below its carrying amount. Such events or circumstances could include the occurrence of operating losses, a significant decline in earnings, or significant changes in or r

Revenue and Related Matters

Revenue and Related Matters	12 Months Ended
	Dec. 31, 2023
Revenue from Contract with Customer [Abstract]
Revenue and Related Matters	Note 4. Revenue and Related Matters Revenue from Contracts with Customers The Company’s customers primarily consist of entities engaged in the life sciences research that pursue the discovery and development of new drugs for a variety of neurologic, oncologic, cardiovascular, infectious disease, and other protein biomarkers associated with diseases. The Company’s customer base includes pharmaceutical, biotechnology, contract research organizations, academic, and government institutions. Disaggregated Revenue When disaggregating revenue, the Company considers all of the economic factors that may affect its revenues. The following tables disaggregate the Company’s revenue from contracts with customers by geography, based on the location products and services are consumed, and revenue type (in thousands): Year Ended December 31, 2023 North America EMEA Asia Pacific Total (As Restated) (As Restated) (As Restated) (As Restated) Product revenue: Instruments $ 6,374 $ 4,384 $ 4,947 $ 15,705 Consumable and other products 35,122 21,216 7,627 63,965 Total $ 41,496 $ 25,600 $ 12,574 $ 79,670 Service revenue: Service-type warranties $ 6,265 $ 3,001 $ 613 $ 9,879 Research services 24,706 2,000 1,122 27,828 Other services 1,436 951 (5) 2,382 Total $ 32,407 $ 5,952 $ 1,730 $ 40,089 Collaboration and license revenue: Total $ 1,380 $ — $ — $ 1,380 Year Ended December 31, 2022 North America EMEA Asia Pacific Total Product revenue: Instruments $ 9,254 $ 8,362 $ 7,388 $ 25,004 Consumable and other products 25,894 14,514 4,396 44,804 Total $ 35,148 $ 22,876 $ 11,784 $ 69,808 Service and other revenues: Service-type warranties $ 5,581 $ 2,779 $ 480 $ 8,840 Research services 22,493 1,013 147 23,653 Other services 1,144 722 136 2,002 Total $ 29,218 $ 4,514 $ 763 $ 34,495 Collaboration and license revenue: Total $ 274 $ 323 $ 52 $ 649 Year Ended December 31, 2021 North America EMEA Asia Pacific Total Product revenue: Instruments $ 12,138 $ 8,178 $ 5,657 $ 25,973 Consumable and other products 34,997 16,122 3,970 55,089 Total $ 47,135 $ 24,300 $ 9,627 $ 81,062 Service and other revenue: Service-type warranties $ 4,334 $ 2,039 $ 255 $ 6,628 Research services 12,101 2,600 124 14,825 Other services 1,372 695 109 2,176 Total $ 17,807 $ 5,334 $ 488 $ 23,629 Collaboration and license revenue: Total $ 360 $ 288 $ — $ 648 UltraDx Product Sales Agreement On May 26, 2022, the Company and UltraDx Limited (“UltraDx”), a company formed by ARCH Venture Partners (“ARCH”), entered into an agreement (the “UltraDx Agreement”). Under the UltraDx Agreement, the Company agreed to supply UltraDx with HD-X instruments (both fully assembled and disassembled), assays and assay components, and granted a co-exclusive license to manufacture, seek Chinese regulatory approval of (including performance of any necessary research and development activities), and commercialize, HD-X instruments assembled in China and related assays in the Chinese in vitro diagnostic (“IVD”) market. Refer to Note 17 - Related Party Transactions for a discussion of the related party relationships between Quanterix and these entities. The Company determined that the instruments, components, and licenses formed a single, combined performance obligation. The consideration due to the Company included (1) cash proceeds of $1.9 million, which was received and recognized as revenue in the third quarter of 2022 when the instruments, components, and licenses were delivered to and paid for by UltraDx, and (2) contingent, non-cash consideration in the form of ordinary shares of UltraDx with a deemed fair value of $1.0 million. The issuance of the shares was contingent on UltraDx completing a preferred share financing under the terms and conditions in the UltraDx Agreement. Given the uncertainty of the completion of the preferred share financing, the Company concluded that the non-cash consideration related to the ordinary shares was variable consideration that was fully constrained at contract inception. In the second quarter of 2023, UltraDx completed the qualified preferred share financing and issued to the Company one million ordinary shares. Refer to Note 7 - Fair Value of Financial Instruments for the Company’s disclosures related to determining the fair value of the shares received. Also refer to Note 20 - Variable Interest Entities for additional information on the Company’s investment in UltraDx as a result of the share issuance. During the year ended December 31, 2023, the Company recognized $1.8 million of revenue from UltraDx, which includes the one-time revenue from the receipt of the UltraDx shares in the second quarter of 2023. During the year ended December 31, 2022, the Company recognized $1.9 million of revenue from UltraDx. Eli Lilly and Company Service Revenue Agreements On February 25, 2022, the Company entered into a Master Collaboration Agreement with Eli Lilly and Company (“Lilly”) establishing a framework for future projects focused on the development of Simoa immunoassays (the “Lilly Collaboration Agreement”). The Company also entered into a statement of work under the Lilly Collaboration Agreement to perform assay research and development services within the field of Alzheimer’s disease. In connection with the Lilly Collaboration Agreement, the Company received a non-refundable up-front payment of $5.0 million during the first quarter of 2022, which was recognized over a one-year period. In addition, under the statement of work, the Company receives $1.5 million per calendar quarter, which began in the first quarter of 2022. The statement of work automatically renews on a quarterly basis until Lilly provides a termination notice in accordance with the terms of the Lilly Collaboration Agreement. As of December 31, 2023, the Lilly Collaboration Agreement and the statement of work were still in effect. Concurrent with the execution of the Lilly Collaboration Agreement, the Company entered into a Technology License Agreement (the “Lilly License”) under which Lilly granted the Company a non-exclusive license to Lilly’s proprietary p-Tau 217 antibody technology for use in research use only products and services and future IVD applications within the field of Alzheimer’s disease. In consideration of the Lilly License, the Company paid an upfront fee, is required to make milestone payments based on the achievement of predetermined regulatory and commercial events, and will pay royalties on net sales of licensed products. The Company recognized revenue from the Lilly Collaboration Agreement of $6.0 million during the year ended December 31, 2023, and $10.9 million during the year ended December 31, 2022. Contract Assets There were no contract assets as of December 31, 2023. Deferred Revenue During the years ended December 31, 2023 and 2022, the Company recognized $7.7 million and $5.5 million of revenue, respectively, related to its deferred revenue balance at January 1 of each such period. Remaining Performance Obligations As of December 31, 2023, the aggregate amount of transaction prices allocated to performance obligations that were not yet satisfied, or were partially satisfied, was $10.7 million. Of the performance obligations not yet satisfied or partially satisfied, $9.5 million is expected to be recognized as revenue in the next 12 months, with the remainder expected to be recognized thereafter. The $10.7 million primarily consists of amounts billed for undelivered services related to initial and extended service-type warranties and research services. Costs to Obtain a Contract Changes in costs to obtain a contract were as follows (in thousands): 2023 2022 2021 Balance at December 31 of prior year $ 377 $ 440 $ 248 Capitalization of costs to obtain a contract 528 1,387 905 Recognition of costs to obtain a contract (617) (1,450) (713) Balance at December 31 $ 288 $ 377 $ 440 The Company evaluates potential impairment of these amounts at each balance sheet date, and no related impairments were recorded during the years ended December 31, 2023, 2022, and 2021, respectively. Grant Revenue All of the Company's grant revenue is generated within North America. Grant revenue for the years ended December 31, 2023, 2022, and 2021 was $1.2 million, $0.6 million, and $5.2 million, respectively. NIH Grant On September 21, 2022, the Company and the National Institutes of Health (the “NIH”), an agency of the U.S. Department of Health and Human Services, entered into a contract (the “NIH Grant”) with a total award value of $1.7 million. The NIH granted the Company funding in support of the development of certain point-of-care diagnostic technologies through collaborative efforts. Grant funding is to be used solely for activities related to the point-of-care diagnostic device development project and the contract period runs through August 2025. Receipt of the award value occurs throughout the term of the contract period and after the Company submits for reimbursement of activities related to the grant. As of December 31, 2023, the Company had received $0.6 million of the award value. During the year ended December 31, 2023, grant revenue recognized and research and development expenses incurred were $0.7 million a nd $0.6 million , respectively . During the year ended December 31, 2022, grant revenue recognized and research and development expenses incurred were not material. ADDF Grant On March 24, 2022, the Company and the Alzheimer’s Drug Discovery Foundation (the “ADDF”) entered into a contract (the “ADDF Grant”) with a total funding value of $2.3 million. The ADDF is a charitable venture philanthropy entity that granted the Company funding in support of certain activities for the development of an IVD test for early detection of Alzheimer's disease. The ADDF Grant restricts the Company’s use of the granted funds solely for activities related to the Company’s Alzheimer’s diagnostic test development project and the contract period runs through June 2024. Receipt of the contract funding was subject to achievement of pre-defined milestones, and as of December 31, 2023, the Company had received the total funding value of $2.3 million. During the year ended December 31, 2023, grant revenue recognized and research and development expenses incurred was $0.5 million a nd $0.5 million , respectively . During the year ended December 31, 2022, grant revenue recognized and research and development expenses incurred was $0.6 million and $0.6 million, respectively. As of December 31, 2023, the Company had $1.1 million of deferred revenue related to the ADDF Grant. RADx Grant On September 29, 2020, the Company entered into a contract with the NIH under its Rapid Acceleration of Diagnostics (“RADx”) program (the “RADx Grant”), with a total award value of $18.2 million. The RADx Grant was to accelerate the continued development, scale-up, and deployment of the novel SARS-CoV-2 antigen detection test using the Company’s Simoa technology. Grant funding was used to expand assay kit manufacturing capacity and commercial deployment readiness, and the contract ran through the final milestone on May 31, 2022. Receipt of the award value occurred throughout the term of the contract period and after the Company submitted for reimbursement of activities related to the grant. During the first half of 2022, the Company received $0.5 million which represented the final and total funding value of the $18.2 million award. During the years ended December 31, 2023 and 2022, the Company recognized no grant revenue and research and development expenses incurred were not material. During the year ended December 31, 2021 , the Company recognized $5.2 million in grant revenue and inc urred $3.4 million in research an d development expenses. As of December 31, 2023, the Company had no future obligations under the RADx Grant.

Allowance for Credit Losses

Allowance for Credit Losses	12 Months Ended
	Dec. 31, 2023
Allowance for Credit Loss [Abstract]
Allowance for Credit Losses	Note 5. Allowance for Credit Losses The change in the allowance for credit losses on accounts receivable is summarized as follows (in thousands): 2023 2022 2021 Balance at December 31 of prior year $ 118 $ 419 $ 370 Provision for expected credit losses 729 752 213 Write-offs and recoveries collected (393) (1,053) (164) Balance at December 31 $ 454 $ 118 $ 419

Marketable Securities

Marketable Securities	12 Months Ended
	Dec. 31, 2023
Investments, Debt and Equity Securities [Abstract]
Marketable Securities	Note 6. Marketable Securities The amortized cost, gross unrealized gains, gross unrealized losses, and fair value of the Company’s marketable securities by major security type were as follows (in thousands): As of December 31, 2023 Amortized cost Unrealized Gains Unrealized Losses Fair Value Commercial paper $ 53,482 $ 23 $ (12) $ 53,493 U.S. Treasuries 4,896 1 — 4,897 U.S. Government agency bonds 28,366 39 (7) 28,398 Corporate bonds 66,726 289 (8) 67,007 Total marketable securities $ 153,470 $ 352 $ (27) $ 153,795 Marketable securities are reported in the following Consolidated Balance Sheets captions: Cash and cash equivalents $ 6,893 Marketable securities 146,902 Total marketable securities $ 153,795 The Company did not have any marketable securities as of December 31, 2022. The following table shows the fair value and gross unrealized losses of the Company’s available-for-sale securities with unrealized losses that are not deemed to be other-than-temporary, aggregated by major security type and length of time that the individual securities have been in a continuous unrealized loss position (in thousands): Less Than 12 Months As of December 31, 2023 Fair Value Unrealized Losses Commercial paper $ 32,137 $ (12) U.S. Government agency bonds 15,861 (7) Corporate bonds 8,367 (8) Total $ 56,365 $ (27) The Company did not have any individual securities in a continuous loss position for greater than 12 months, and there were no individual securities that were in a significant unrealized loss position as of December 31, 2023. For marketable securities in an unrealized loss position, the Company does not intend to sell them before recovery of their amortized cost bases, it is not more likely than not that the Company will be required to sell them before recovery of their amortized cost bases, and the unrealized losses are not credit related. Accordingly, the Company has not recorded any impairment losses or a credit loss allowance. The Company did not sell any marketable securities or record any realized gains or losses for the year ended December 31, 2023. At December 31, 2023, the Company had $1.0 million of accrued interest receivable on its marketable securities. The following table summarizes the contractual maturities of the Company’s marketable securities (in thousands): As of December 31, 2023 Amortized cost Fair Value Due within one year $ 95,188 $ 95,232 Due in one to two years 58,282 58,563 Total $ 153,470 $ 153,795

Fair value of Financial Instrum

Fair value of Financial Instruments (As Restated)	12 Months Ended
	Dec. 31, 2023
Fair Value Disclosures [Abstract]
Fair value of Financial Instruments (As Restated)	Note 7. Fair Value of Financial Instruments (As Restated) Recurring Fair Value Measurements The following tables present the Company’s fair value hierarchy for its financial assets that are measured at fair value on a recurring basis (in thousands): As of December 31, 2023 Total Level 1 Level 2 Level 3 Financial assets: Cash equivalents: (1) Money market funds $ 155,367 $ 155,367 $ — $ — Commercial paper 1,996 — 1,996 — U.S. Treasuries 4,897 — 4,897 — Total cash equivalents 162,260 155,367 6,893 — Marketable securities: (2) Commercial paper 51,498 — 51,498 — U.S. Treasuries — — — — U.S. Government agency bonds 28,398 — 28,398 — Corporate bonds 67,006 — 67,006 — Total marketable securities 146,902 — 146,902 — Total financial assets $ 309,162 $ 155,367 $ 153,795 $ — As of December 31, 2022 Total Level 1 Level 2 Level 3 Financial assets: Money market funds (1) $ 306,097 $ 306,097 $ — $ — Total financial assets $ 306,097 $ 306,097 $ — $ — (1) Included in cash and cash equivalents on the Consolidated Balance Sheets. (2) Marketable securities are initially valued at their purchase price and subsequently fair valued at the end of each reporting period utilizing third party pricing services or other observable data. The pricing services utilize industry standard valuation methods, including both income and market-based approaches and observable market inputs to determine the fair value. These observable market inputs include reportable trades, benchmark yields, credit spreads, broker/dealer quotes, bids, offers, current spot rates, and other industry and economic events. Nonrecurring Fair Value Measurements The Company has a non-marketable equity investment in an entity that is evaluated under the VIE guidance (refer to Note 20 - Variable Interest Entities for further discussion). Pursuant to ASC 321 – Investments – Equity Securities , the Company uses the measurement alternative for equity investments without readily determinable fair values and recognizes its non-marketable equity investment at cost, less any impairment, and adjusted for any observable price changes in orderly transactions. On June 26, 2023, the Company received ordinary shares in UltraDx (refer to Note 4 - Revenue and Related Matters ) which were valued at $0.8 million upon receipt, primarily using the third-party purchase price of similar interests issued during UltraDx’s financing event that closed in the second quarter of 2023. As UltraDx is a recently formed, privately held entity, there was minimal market activity or other financial information available to determine the fair value of UltraDx’s shares and therefore this investment is considered a Level 3 financial asset. Changes in the inputs and assumptions used to determine the fair value would have resulted in a higher or lower fair value measurement. As of December 31, 2023 the carrying value of the Company’s Level 3 financial asset was $0.8 million and is included in other non-current assets on the Consolidated Balance Sheets. Other Fair Value Disclosures During the years ended December 31, 2023 and 2022, the Company did not transfer financial assets between levels of the fair value hierarchy. Additionally, there have been no changes to the valuation techniques for Level 2 or Level 3 financial assets.

Inventory

Inventory	12 Months Ended
	Dec. 31, 2023
Inventory Disclosure [Abstract]
Inventory	Note 8. Inventory Inventory, net of inventory reserves, consisted of the following (in thousands): As of December 31, 2023 2022 (As Restated) (As Restated) Raw materials $ 5,114 $ 5,509 Work in process 5,439 3,415 Finished goods 15,570 8,040 Total inventory $ 26,123 $ 16,964

Property and Equipment

Property and Equipment	12 Months Ended
	Dec. 31, 2023
Property, Plant and Equipment [Abstract]
Property and Equipment	Note 9. Property and Equipment Property and equipment consisted of the following (in thousands): As of December 31, 2023 2022 (As Restated) Laboratory and manufacturing equipment $ 13,141 $ 11,753 Office furniture and equipment 1,905 1,798 Computers and software 3,927 3,581 Leasehold improvements 13,074 13,688 Total cost $ 32,047 $ 30,820 Less: accumulated depreciation (14,121) (11,009) Property and equipment, net $ 17,926 $ 19,811 The Company incurred depreciation expense of $4.7 million, $3.5 million, and $2.8 million for the years ended December 31, 2023, 2022, and 2021, respectively. Substantially all of the Company’s property and equipment is located in North America. During the year ended December 31, 2023, the Company had $0.8 million of disposals related to equipment no longer being used by the Company. There were no material disposals during the years ended December 31, 2022, and 2021. For the years ended December 31, 2023, and 2022, except for the recorded impairments related to leasehold improvements associated with two leased facilities not being used (refer to Note 15 - Leases for further discussion), the Company's impairments were not material. There were no material impairments during the year ended December 31, 2021. As of December 31, 2023, the Company capitalized $1.1 million of software development costs, all of which are included in the computers and software category above. Amounts capitalized as of December 31, 2022 and 2021 were not material. Depreciation of capitalized software costs for the years ended December 31, 2023, 2022, and 2021 was not material.

Accrued Expenses and Other Curr

Accrued Expenses and Other Current Liabilities	12 Months Ended
	Dec. 31, 2023
Accrued Expenses and Other Current Liabilities
Accrued Expenses and Other Current Liabilities	Note 10. Accrued Expenses and Other Current Liabilities Accrued expenses and other current liabilities consisted of the following (in thousands): As of December 31, 2023 2022 (As Restated) (As Restated) Accrued professional services $ 1,596 $ 1,409 Accrued royalties 1,689 815 Accrued tax liabilities 822 186 Other accrued expenses 1,948 2,737 Total accrued expenses and other current liabilities $ 6,055 $ 5,147

Stock-Based Compensation

Stock-Based Compensation	12 Months Ended
	Dec. 31, 2023
Share-Based Payment Arrangement [Abstract]
Stock-Based Compensation	Note 11. Stock-Based Compensation Stock-Based Compensation Plans In December 2017, the Company adopted the 2017 Employee, Director and Consultant Equity Incentive Plan (the “2017 Plan”), under which it may grant incentive stock options, non-qualified stock options, RSUs, and other stock-based awards. As of December 31, 2017, the 2017 Plan allowed for the issuance of (1) up to 1.0 million shares of common stock and (2) up to 2.5 million shares of common stock represented by awards granted under the 2007 Stock Option and Grant Plan (which was terminated upon completion of the Company’s initial public offering) that were forfeited, expired, or cancelled without delivery of shares or which result in the forfeiture of shares of common stock back to the Company on or after the date the 2017 Plan became effective. The 2017 Plan contains an “evergreen” provision, which allows for an annual increase in the number of shares of common stock available for issuance under the 2017 Plan on the first day of each fiscal year during the period beginning in 2019 and ending in 2027. The annual increase is equal to the lowest of (1) 4% of the number of shares of common stock outstanding as of such date and (2) an amount determined by the Company’s Board of Directors or Compensation Committee. As of December 31, 2023, 4.0 million shares were outstanding and there were 2.0 million shares available for grant under the 2017 Plan. In December 2017, the Company adopted the 2017 Employee Stock Purchase Plan (the “2017 ESPP”). As of December 31, 2019, the 2017 ESPP allowed for the issuance of up to 0.6 million shares of common stock. The 2017 ESPP contains an “evergreen” provision, which allows for an increase in the number of shares under the plan on the first day of each fiscal year beginning with 2018 and ending in 2027. The increase is equal to the lowest of: (1) 1% of the number of shares of common stock outstanding on the last day of the immediately preceding fiscal year and (2) an amount determined by the Company’s Board of Directors or Compensation Committee. As of December 31, 2023, 1.7 million shares were available for grant under the 2017 ESPP. The 2017 ESPP provides for six-month offering periods commencing and ending on March 1 through August 31, and September 1 through February 28. During the years ended December 31, 2023, 2022, and 2021, employees purchased 121 thousand, 57 thousand, and 29 thousand shares, respectively, of the Company’s common stock pursuant to the 2017 ESPP. Stock Options Under the 2017 Plan, stock options may not be granted with exercise prices of less than fair market value on the date of the grant. Options generally vest ratably over a four-year period with 25% vesting on the first anniversary and the remaining 75% vesting ratably on a monthly basis over the remaining three years. These options expire ten years after the grant date. Stock option activity for the year ended December 31, 2023 is presented below (in thousands, except per share and contractual life amounts): Number of options Weighted-average Weighted-average remaining contractual Aggregate Outstanding at December 31, 2022 2,188 $ 20.69 8.1 $ 4,273 Granted 1,187 16.23 Exercised (161) 11.77 Forfeited/expired (440) 18.65 Outstanding at December 31, 2023 2,774 $ 19.62 7.9 $ 26,941 Exercisable at December 31, 2023 1,117 $ 22.81 6.4 $ 9,176 Vested and expected to vest at December 31, 2023 2,774 $ 19.62 7.9 $ 26,941 The total intrinsic value of stock options exercised was $1.9 million in 2023, $3.4 million in 2022, and $24.3 million in 2021. Restricted Stock Units RSUs represent the right to receive shares of common stock upon meeting specified vesting requirements. Shares are delivered to the grantee upon vesting, less shares for the payment of withholding taxes. RSU activity for the year ended December 31, 2023 is presented below (in thousands, except per share amounts): Number of shares Weighted-average grant Unvested as of December 31, 2022 1,188 $ 21.18 Granted 868 15.90 Vested (453) 23.00 Forfeited (275) 17.50 Unvested as of December 31, 2023 1,328 $ 17.87 Expected to convert at December 31, 2023 1,328 $ 17.87 The weighted average grant-date fair value per share of awards granted was $15.90 in 2023, $18.32 in 2022, and $58.20 in 2021.The total fair value of shares that vested was $10.4 million in 2023, $9.8 million in 2022 and $7.3 million in 2021. Stock-Based Compensation Expense Stock-based compensation expense was recorded in the following categories on the Consolidated Statements of Operations (in thousands): Year Ended December 31, 2023 2022 2021 (As Restated) (As Restated) Cost of product revenue $ 839 $ 596 $ 471 Cost of service and other revenue 1,124 819 403 Research and development 1,713 1,629 1,807 Selling, general and administrative 13,147 12,339 13,294 Total stock-based compensation $ 16,823 $ 15,383 $ 15,975 As of December 31, 2023, there was $35.4 million of total unrecognized stock-based compensation expense related to unvested RSUs and stock options, which is expected to be recognized over the remaining weighted-average vesting period of 2.7 years. The fair value of the Company’s stock options granted and purchase rights to the ESPP were estimated using the Black-Scholes valuation model with the following assumptions: Year Ended December 31, 2023 2022 2021 Stock Options: Risk-free interest rate 3.5% - 4.7% 1.4% - 4.1% 0.4% - 1.3% Expected dividend yield None None None Expected term (in years) 5.0 - 5.2 5.0 - 5.8 6.0 Expected volatility 71.1% - 83.1% 55.0% - 70.8% 49.2% - 55.6% Weighted-average grant date fair value per share $ 10.63 $ 9.88 $ 29.96 Employee Stock Purchase Plan: Risk-free interest rate 5.2% - 5.5% 0.7% - 3.9% 0.1% Expected dividend yield None None None Expected term (in years) 0.5 0.5 0.5 Expected volatility 72.8% - 82.5% 51.9% - 117.3% 46.3% - 59.8% Weighted-average grant date fair value per share $ 3.19 $ 3.53 $ 8.06

Net Loss Per Share

Net Loss Per Share	12 Months Ended
	Dec. 31, 2023
Earnings Per Share [Abstract]
Net Loss Per Share	Note 12. Net Loss Per Share The following table presents the computation of basic and diluted net loss per share (in thousands, except per share data): Year Ended December 31, 2023 2022 2021 (As Restated) (As Restated) (As Restated) Numerator: Net loss $ (28,354) $ (99,574) $ (55,548) Denominator: Weighted average common shares outstanding, basic and diluted 37,594 36,991 35,997 Net loss per share, basic and diluted $ (0.75) $ (2.69) $ (1.54) As the Company was in a net loss position, the following common share equivalents (calculated on a weighted average basis) were excluded from the calculation of diluted net loss per share (in thousands): Year Ended December 31, 2023 2022 2021 Stock options 2,783 446 2,305 Common stock and RSUs 1,512 702 531 Estimated ESPP purchases 23 52 8 Total dilutive shares 4,318 1,200 2,844

Income Taxes (As Restated)

Income Taxes (As Restated)	12 Months Ended
	Dec. 31, 2023
Income Tax Disclosure [Abstract]
Income Taxes (As Restated)	Note 13. Income Taxes (As Restated) The following table presents the components of loss before income taxes (in thousands): Year Ended December 31, 2023 2022 2021 United States $ (30,355) $ (92,360) $ (54,414) Foreign 2,720 (7,050) (1,170) Total loss before income taxes $ (27,635) $ (99,410) $ (55,584) The following table summarizes income tax (expense) benefit (in thousands): Year Ended December 31, 2023 2022 2021 Current: United States Federal $ — $ — $ — State (161) (87) (33) Foreign (850) (372) (340) Total current income tax provision (1,011) (459) (373) Deferred United States Federal — 10 5 State — 13 (6) Foreign 292 272 410 Total deferred income tax benefit 292 295 409 Total income tax (expense) benefit $ (719) $ (164) $ 36 A reconciliation of the federal statutory income tax rate to the effective tax rate is as follows: Year Ended December 31, 2023 2022 2021 Federal statutory income tax rate 21.0 % 21.0 % 21.0 % Non-deductible executive compensation (2.6) % — % — % State taxes, net of federal benefit (0.3 %) 2.8 % 6.6 % Tax credits 4.9 % 1.3 % 2.1 % Stock-based compensation (1.6) % (2.4) % 7.7 % Permanent items (0.6) % (0.3) % (1.9) % Deferred tax rate change — % — % 0.2 % Change in valuation allowance (24.3) % (20.8) % (35.2) % Impairment of goodwill — % (1.6) % — % Other 0.9 % (0.2 %) (0.4) % Effective income tax rate (2.6) % (0.2) % 0.1 % The effective income tax rate differs from the U.S. federal statutory rate of 21.0% primarily as a result of the valuation allowance maintained against the Company’s net deferred tax assets. Deferred tax assets and liabilities reflect the net tax effects of temporary differences between the carrying amount of assets and liabilities for financial reporting and the amounts used for income tax purposes. Significant components of the Company’s deferred tax assets and liabilities were as follows (in thousands): As of December 31, 2023 2022 Deferred tax assets: Net operating loss carryforwards $ 77,731 $ 77,689 Tax credits 8,645 7,303 Deferred revenue 2,499 2,385 Amortization 727 847 Stock-based compensation 3,186 2,773 Inventory 1,391 1,507 Capitalized R&D Costs 9,353 5,743 Lease liability 9,948 10,856 Other deferred tax assets 2,357 1,536 Total deferred tax assets 115,837 110,639 Less: valuation allowances (110,082) (103,372) Net deferred tax assets 5,755 7,267 Deferred tax liabilities: Right-of-Use Assets (4,372) (5,219) Depreciation (1,307) (1,756) Amortization acquired intangibles (1,253) (1,570) Other deferred tax liabilities — (189) Net deferred tax (liability) asset (1,177) (1,467) The Company records deferred tax liabilities in other non-current liabilities on the Consolidated Balance Sheets. The Company’s change in its valuation allowance account related to deferred tax assets was as follows (in thousands): 2023 2022 Balance at December 31 of prior year $ 103,372 $ 82,654 Change in valuation allowance 6,710 20,718 Balance at December 31 $ 110,082 $ 103,372 The valuation allowance increased during the year ended December 31, 2023 primarily as a result of the U.S. operating losses incurred. In determining the need for a valuation allowance, the Company considers the cumulative book income and loss positions of each of its entities as well as its worldwide cumulative loss position. The Company has assessed, on a jurisdictional basis, the available means of recovering deferred tax assets, including the ability to carryback net operating losses (“NOLs”), the existence of reversing taxable temporary differences, the availability of tax planning strategies, and forecasted future taxable income. At December 31, 2023, the Company maintained a full valuation allowance against its worldwide net deferred tax assets, as it concluded that it was more likely than not that the deferred assets will not be utilized. As of December 31, 2023, the Company had U.S. federal net operating losses (“NOLs”) of approximately $308.8 million. U.S. federal NOLs generated through December 31, 2017 of approximately $108.5 million expire at various dates through 2037, and U.S. federal NOLs generated after December 31, 2017 of approximately $200.3 million do not expire. As of December 31, 2023, the Company had U.S. federal tax credit carryforwards of approximately $6.7 million that expire at various dates through 2043. As of December 31, 2023, the Company had $214.3 million of state NOLs, approximately $203.8 million of which expire at various dates through 2043, and approximately $10.5 million of which do not expire. As of December 31, 2023, the Company had U.S. state tax credit carryforwards of approximately $2.5 million that expire at various dates through 2038. Under Sections 382 and 383 of the U.S. Internal Revenue Code, if a corporation undergoes an ownership change, the corporation’s ability to use its pre-ownership change NOLs and other pre-ownership change tax attributes, such as research tax credits, to offset its post-change income and taxes may be limited. In general, an ownership change occurs if there is a cumulative change in an entity’s ownership by 5% stockholders that exceeds 50 percentage points over a rolling three-year period. Similar rules may apply under U.S. state tax laws. Under the Tax Cuts and Jobs Act of 2017 (“TCJA”), the use of federal NOLs arising in taxable years beginning after December 31, 2017 is limited to 80% of current year taxable income and NOLs arising in taxable years ending after December 31, 2017 may not be carried back (though any such NOLs may be carried forward indefinitely). The Company may have experienced an ownership change in the past and may experience ownership changes in the future as a result of future transactions in its share capital, some of which may be outside of the control of the Company. As a result, if the Company earns net taxable income, its ability to use its pre-ownership change NOLs, or other pre-ownership change tax attributes, to offset U.S. federal and state taxable income and taxes may be subject to significant limitations. For the years ended December 31, 2023, 2022, and 2021, the Company had no tax reserves accrued for uncertain tax positions. The Company is subject to taxation in the United States as well as the Netherlands, Sweden, and China. At December 31, 2023, the Company is generally no longer subject to examination by taxing authorities in the United States for years prior to 2020. However, NOLs and tax credits in the United States may be subject to adjustments by taxing authorities in future years in which they are utilized. The Company’s foreign subsidiaries remain open to examination by taxing authorities from 2018 onward. As of December 31, 2023, the Company’s foreign subsidiaries had immaterial undistributed earnings and the tax payable on the earnings that are indefinitely reinvested would be immaterial.

Goodwill and intangible assets

Goodwill and intangible assets	12 Months Ended
	Dec. 31, 2023
Goodwill and Intangible Assets Disclosure [Abstract]
Goodwill and intangible assets	Note 14. Goodwill and Intangible Assets During the third quarter of 2022, the Company identified certain indicators of impairment, including the significant decline in the Company’s stock price, actions taken under the Restructuring Plan (Refer to Note 18 - Restructuring) , and a reduction of forecasted sales and profitability. As a result, the Company performed a goodwill impairment test and determined the carrying amount of the Company’s sole reporting unit exceeded its estimated fair value. The Company concluded that its entire goodwill balance was impaired and recognized an $8.2 million impairment charge during the third quarter of 2022. As of December 31, 2023 and December 31, 2022, the Company had no remaining goodwill balance. Acquired intangible assets consisted of the following (in thousands, except useful life and weighted average life amounts): As of December 31, 2023 Estimated Gross Carrying Accumulated Cumulative Net Carrying Weighted Average Know-how 8.5 $ 13,000 $ (6,326) $ (1,050) $ 5,624 4.0 Developed technology 7 1,650 (1,581) — 69 1.1 Customer relationships 8.5 - 10 1,360 (1,067) (9) 284 4.1 Non-compete agreements 5.5 340 (256) (27) 57 1.0 Trade names 3 50 (50) — — — Total $ 16,400 $ (9,280) $ (1,086) $ 6,034 As of December 31, 2022 Estimated Gross Carrying Accumulated Cumulative Net Carrying Weighted Average Know-how 8.5 $ 13,000 $ (4,763) $ (1,433) $ 6,804 5.0 Developed technology 7 1,650 (1,458) — 192 2.1 Customer relationships 8.5 - 10 1,360 (938) (12) 410 5.1 Non-compete agreements 5.5 340 (193) (37) 110 2.0 Trade names 3 50 (50) — — — Total $ 16,400 $ (7,402) $ (1,482) $ 7,516 The Company recorded amortization expense of $1.6 million, $1.8 million, and $2.0 million for the years ended December 31, 2023, 2022, and 2021, respectively. Amortization of know-how is recorded in cost of goods sold; amortization of developed technology is recorded in research and development expenses; and amortization of customer relationships, non-compete agreements and trade names are recorded in selling, general and administrative expenses on the Consolidated Statements of Operations. Future estimated amortization expense is as follows (amounts in thousands): As of December 31, 2023 2024 $ 1,631 2025 1,493 2026 1,465 2027 1,443 2028 2 Thereafter — Total amortization expense $ 6,034

Leases

Leases	12 Months Ended
	Dec. 31, 2023
Lessee Disclosure [Abstract]
Leases	Note 15. Leases As part of the Restructuring Plan in the third quarter of 2022 (refer to Note 18 - Restructuring ), the Company decided not to use two leased facilities in its operations and intends to sublease the vacant space to recover a portion of the total lease costs. The Company’s decision to not use the leased facilities triggered an impairment assessment and the related ROU assets and leasehold improvements became their own asset group. The impairment analysis evaluated the present value of net cash flows under the original lease and the estimated cash flows under estimated subleases to identify any impairment amount. The impairment assessment considered all industry and economic factors such as rental rates, interest rates, and recent real estate activities to estimate the net cash flows analysis and impairment amount. During 2023, the Company continued to reassess the remaining ROU assets and leasehold improvements associated with the leased facilities that remained vacant. The above assessments resulted in the Company recording an impairment charge of $1.3 million and $16.3 million during the years ended December 31, 2023 and 2022, respectively, which was recorded in impairment and restructuring on the Consolidated Statements of Operations. For the years ended December 31, 2023 and 2022, impairment charges included $1.1 million and $12.0 million impairment of ROU assets, respectively. For the related leasehold improvements, during the year ended December 31, 2023, impairment charges were not material and for the year ended December 31, 2022 impairment charges were $4.3 million. After recording impairments for the year ended December 31, 2023 and 2022, the carrying value of ROU assets and related leasehold improvements for facilities not being used were $10.1 million and $12.8 million, respectively. There were no ROU asset or leasehold improvement impairments recorded in 2021. The components of the lease costs and supplemental cash flow information relating to the Company's leases were as follows (in thousands): Year Ended December 31, 2023 2022 2021 Operating lease cost $ 5,209 $ 5,488 $ 2,726 Short-term and variable lease cost 3,996 3,417 2,699 Total lease cost $ 9,205 $ 8,905 $ 5,425 Year Ended December 31, 2023 2022 Cash paid for amounts included in the measurement of operating lease liabilities $ 8,935 $ 6,539 Operating ROU assets obtained in exchange for lease obligations $ — $ 22,494 Weighted average remaining lease term - operating leases (years) 6.8 7.8 Weighted average discount rate - operating leases 7.86 % 7.83 % The undiscounted future lease payments for non-cancelable operating leases were as follows (in thousands): Maturity of lease liabilities As of December 31, 2023 2024 $ 7,094 2025 7,254 2026 7,408 2027 7,641 2028 7,880 Thereafter 15,741 Total lease payments $ 53,018 Less: imputed interest (11,554) Total operating lease liabilities $ 41,464 Operating lease balances presented on the Consolidated Balance Sheets were as follows (in thousands): As of December 31, 2023 Operating lease ROU assets $ 18,251 Operating lease liabilities $ 4,241 Operating lease liabilities, net of current portion 37,223 Total operating lease liabilities $ 41,464

Commitments and contingencies

Commitments and contingencies	12 Months Ended
	Dec. 31, 2023
Commitments and Contingencies Disclosure [Abstract]
Commitments and contingencies	Note 16. Commitments and Contingencies Purchase Commitments STRATEC During 2022, the Company and STRATEC Consumables GmbH (“STRATEC”) entered into an amendment to the supply agreement with STRATEC (as amended, the “STRATEC Supply Agreement”), related to the supply of discs used in Simoa bead-based instruments. As part of the STRATEC Supply Agreement, the Company agreed to purchase a total of 515 thousand discs to be shipped at various points starting in 2022 and continuing through 2024 at an agreed purchase price per disc. The total purchase commitment under the STRATEC Supply Agreement is $3.7 million, of which $2.1 million has been paid, and $1.0 million is due within one year from December 31, 2023. In 2023 and 2022, STRATEC shipped 218 thousand and 75 thousand discs, respectively, to the Company. The Company recorded cost of product revenue related to these shipments of $1.6 million and $0.5 million for the years ended December 31, 2023 and 2022, respectively. During 2024, STRATEC is required to ship 222 thousand discs to the Company. Other Purchase Commitments The Company’s other non-cancellable purchase commitments primarily consist of purchases of raw materials for manufacturing operations under annual and multi-year agreements, some of which have minimum quantity requirements. The Company’s total purchase commitments under these agreements as of December 31, 2023 was $3.1 million, most of which the Company expects to incur in the year ending December 31, 2024. License Agreements Harvard University In August 2022, the Company and Harvard University (“Harvard”) entered into an exclusive license agreement (the “Harvard License Agreement”) for certain intellectual property owned by Harvard. Pursuant to the Harvard License Agreement, the Company paid an upfront fee of $0.6 million, which was recorded in research and development expenses on the Consolidated Statements of Operations. Under this license, the Company is required to pay Harvard low single-digit royalties on net sales of products and services using the licensed technology, as well as a portion of its applicable sublicense revenues. The Company incurred no royalty expense under the Harvard License Agreement for the years ended December 31, 2023 and 2022. Refer to Note 17 - Related Party Transactions for a discussion of a related party relationship with Harvard. Tufts University In June 2007, the Company and Tufts University (“Tufts”) entered into a license agreement (the “Tufts License Agreement”) for certain intellectual property owned by Tufts. The Tufts License Agreement, which was subsequently amended, is exclusive and sub-licensable, and will continue in effect on a country by country basis as long as there is a valid claim of a licensed patent in a country. The Company is contractually obligated to pay license and maintenance fees that are creditable against royalties, in addition to low single-digit royalties on direct sales and services, and a royalty on sublicense income. The Company incurred royalty expenses related to the Tufts License Agreement of $1.7 million, $1.4 million, and $1.6 million, during the years ended December 31, 2023, 2022, and 2021, respectively, which was recorded in cost of product revenue on the Consolidated Statements of Operations. Refer to Note 17 - Related Party Transactions for a discussion of a related party relationship with Tufts. Legal Contingencies The Company is subject to claims in the ordinary course of business; however, the Company is not currently a party to any pending or threatened litigation, the outcome of which would be expected to have a material adverse effect on its financial condition or results of operations. The Company accrues for contingent liabilities when losses are probable and estimable. If an estimate of a probable loss is a range and no amount within the range is more likely than any other amount in the range, the Company accrues the minimum amount of the range.

Related Party Transactions

Related Party Transactions	12 Months Ended
	Dec. 31, 2023
Related Party Transactions [Abstract]
Related Party Transactions	Note 17. Related Party Transactions In June 2007, the Company entered into the Tufts License Agreement for certain intellectual property owned by Tufts (refer to Note 16 - Commitments and Contingencies ). A member of the Company’s Board of Directors was previously affiliated with Tufts. This Board member continues to receive compensation from Tufts on a formulaic basis based on royalties and license payments the Company makes to Tufts. At December 31, 2023 and 2022, open payable balances to Tufts were not material. In August 2022, the Company entered into the Harvard License Agreement for certain intellectual property owned by Harvard (refer to Note 16 - Commitments and Contingencies ). Harvard is obligated to pay a portion of the payments received from the Company under the Harvard License Agreement to a member of the Company’s Board of Directors. A member of the Company’s Board of Directors is also affiliated with Harvard and Mass General Brigham. Revenue recorded from sales of products and services to Harvard and its affiliates and to Mass General Brigham and its affiliates totaled $1.3 million and $0.7 million for the years ended December 31, 2023 and 2022, respectively, and was not material for the year ended December 31, 2021. Cost of product revenue and operating expenses with Harvard and its affiliates and Mass General Brigham and its affiliates was $0.3 million for the year ended December 31, 2023, and was not material for the years ended December 31, 2022 and 2021. At December 31, 2023 and 2022, open payables to and receivable balances from Harvard and Mass General Brigham were not material. As discussed in Note 4 - Revenue and Related Matters

Restructuring

Restructuring	12 Months Ended
	Dec. 31, 2023
Restructuring and Related Activities [Abstract]
Restructuring	Note 18. Restructuring Following a strategic review and assessment of the Company’s operations and cost structure, on August 8, 2022, the Company announced a plan of restructuring and strategic re-alignment plan (the “Restructuring Plan”). As part of the Restructuring Plan, the Company began an assay redevelopment program with the ultimate objective of improving its ability to manufacture and deliver high-quality assays at scale. The Restructuring Plan aligned the Company’s investments to best serve the needs of its customers, focused the Company’s innovation efforts on key platforms, and provided a foundation for the Company’s entry into translational pharma and clinical markets. In accordance with the Restructuring Plan, the Company implemented a workforce reduction, which was substantially completed by the end of the third quarter of 2022. The Restructuring Plan included the elimination of 119 positions and other cost-saving measures. During the year ended December 31, 2022, the Company incurred approximately $3.8 million of expenses related to the Restructuring Plan, which represents the total amount expected to be incurred and the amount incurred to date. These costs were recorded in impairment and restructuring on the Consolidated Statements of Operations and were substantially for cash payments of severance and employee benefits, $3.5 million of which was paid by December 31, 2022. As a result of the Restructuring Plan, the Company performed an impairment assessment of its goodwill, long-lived assets, including ROU assets and related leasehold improvements, and intangibles. The assessments resulted in the Company recording an impairment charge of $25.6 million during the year ended December 31, 2022, which was recorded in impairment and restructuring on the Consolidated Statements of Operations. The impairment charge included (1) $8.2 million of goodwill (refer to Note 14 - Goodwill and Intangible Assets ), (2) $16.3 million associated with the ROU assets and related leasehold improvements at two leased facilities not being used (refer to Note 15 - Leases ), and (3) $1.1 million for software costs related to projects that were rationalized as part of the Restructuring Plan or the related exit and disposal costs. During the year ended December 31, 2023, the Company recorded an additional impairment of $1.3 million associated with the ROU asset and related leasehold improvements at the leased facilities not being used (refer to Note 15 − Leases ). The following table presents the restructuring reserve and provision activity for the year ended December 31, 2023 (in thousands): Severance and Employee Benefit Costs Balance at December 31, 2022 $ 328 Accrual adjustments (33) Cash payments (16) Foreign currency translation 8 Balance at December 31, 2023 $ 287 The Company did not have any restructuring activities related to the Restructuring Plan during the year ended December 31, 2023.

Employee benefit plans

Employee benefit plans	12 Months Ended
	Dec. 31, 2023
Retirement Benefits [Abstract]
Employee benefit plans	Note 19. Employee Benefit Plans The Company sponsors a 401(k) savings plan for employees and may make discretionary contributions. During the years ended December 31, 2023, 2022, and 2021, the Company made contributions of $0.8 million, $1.2 million, and $1.1 million, respectively.

Variable Interest Entities (As

Variable Interest Entities (As Restated)	12 Months Ended
	Dec. 31, 2023
Variable Interest Entities
Variable Interest Entities (As Restated)	Note 20. Variable Interest Entities (As Restated) The Company enters into relationships with, or has investments in, other entities that may be VIEs. The Company assesses the criteria in ASC 810 – Consolidation to determine if any of these entities meet the definition of a VIE and require consolidation into its financial statements. The Company’s analysis determines whether it has a controlling financial interest and also identifies the primary beneficiary of a VIE as the enterprise that has both (1) the power to direct activities of a VIE that most significantly impact the entity’s economic performance and (2) the obligation to absorb losses of, or the right to receive benefits from, the entity that could potentially be significant to that entity. As discussed in Note 4 - Revenue and Related Matters , during the second quarter of 2023 the Company received one million ordinary shares of UltraDx under the UltraDx Agreement. Primarily due to having less than a 5% ownership interest in UltraDx, the Company concluded that it does not have the power to direct activities impacting UltraDx’s economic performance and therefore the Company is not the primary beneficiary of the VIE. Based on the Company’s assessments, it does not have any controlling financial interests in any VIEs, and therefore did not consolidate any VIEs into its Consolidated Financial Statements during the years ended December 31, 2023 and 2022. As of December 31, 2023, the carrying value of the Company’s investment in a VIEs was $0.8 million , which was recorded in other non-current assets on the Consolidated Balance Sheets. Refer to Note 7 - Fair Value of Financial Instruments for the Company’s related valuation disclosures. Maximum exposure to losses related to this VIE is limited to its carrying value and the Company does not have any future funding commitments to this VIE.

Quarterly Financial Data (Unaud

Quarterly Financial Data (Unaudited and Restated)	12 Months Ended
	Dec. 31, 2023
Quarterly Financial Information Disclosure [Abstract]
Quarterly Financial Data (Unaudited and Restated)	Note 21. Quarterly Financial Data (Unaudited and Restated) The following tables present restated unaudited Consolidated Financial Statements for the quarterly and year-to-date (as applicable) periods ended March 31, 2023, June 30, 2023, September 30, 2023, March 31, 2022, June 30, 2022, and September 30, 2022. The amounts labeled "As Previously Reported" in the following tables were derived from the previously filed Quarterly Reports on Form 10-Q. The Consolidated Statements of Operations for the quarterly periods ended December 31, 2023 and 2022 are also included and derived from the Company's earnings releases. The amounts labeled “Adjustments” represent the impact of correcting the Combined Misstatements identified by the Company. Refer to Note 1 - Restatement of Financial Statements for further discussion of the Restatement. QUANTERIX CORPORATION RESTATED CONSOLIDATED BALANCE SHEETS (amounts in thousands, except per share data) As of March 31, 2023 As Previously Reported Adjustments As Restated ASSETS Current assets: Cash and cash equivalents $ 329,354 $ — $ 329,354 Accounts receivable, net of allowance for expected credit losses 22,546 — 22,546 Inventory 17,070 (441) 16,629 Prepaid expenses and other current assets 7,002 (57) 6,945 Total current assets 375,972 (498) 375,474 Restricted cash 2,920 (322) 2,598 Property and equipment, net 19,056 (366) 18,690 Intangible assets, net 7,129 — 7,129 Right-of-use assets 20,891 — 20,891 Other non-current assets 1,345 (407) 938 Total assets $ 427,313 $ (1,593) $ 425,720 LIABILITIES AND STOCKHOLDERS' EQUITY Current liabilities: Accounts payable $ 2,585 $ — $ 2,585 Accrued compensation and benefits 4,880 531 5,411 Other accrued expenses 4,624 14 4,638 Deferred revenue 10,682 — 10,682 Short-term lease liabilities 3,875 — 3,875 Other current liabilities 291 — 291 Total current liabilities 26,937 545 27,482 Deferred revenue, net of current portion 1,419 — 1,419 Long-term lease liabilities 40,409 — 40,409 Other non-current liabilities 1,216 — 1,216 Total liabilities 69,981 545 70,526 Commitments and contingencies (Note 16) Stockholders’ equity: Common stock, $0.001 par value per share: Authorized shares: 120,000 shares; Issued and outstanding: 37,424 shares at March 31, 2023. 37 — 37 Additional paid-in capital 768,141 (18) 768,123 Accumulated other comprehensive (loss) income (2,581) 85 (2,496) Accumulated deficit (408,265) (2,205) (410,470) Total stockholders’ equity 357,332 (2,138) 355,194 Total liabilities and stockholders’ equity $ 427,313 $ (1,593) $ 425,720 As of June 30, 2023 As Previously Reported Adjustments As Restated ASSETS Current assets: Cash and cash equivalents $ 329,525 $ — $ 329,525 Accounts receivable, net of allowance for expected credit losses 24,423 (54) 24,369 Inventory 18,156 177 18,333 Prepaid expenses and other current assets 6,954 (57) 6,897 Total current assets 379,058 66 379,124 Restricted cash 2,686 (85) 2,601 Property and equipment, net 18,328 (280) 18,048 Intangible assets, net 6,476 — 6,476 Operating lease right-of-use assets 20,380 — 20,380 Other non-current assets 2,282 (645) 1,637 Total assets $ 429,210 $ (944) $ 428,266 LIABILITIES AND STOCKHOLDERS' EQUITY Current liabilities: Accounts payable $ 3,238 $ — $ 3,238 Accrued compensation and benefits 7,525 650 8,175 Accrued expenses and other current liabilities 6,777 14 6,791 Deferred revenue 10,421 — 10,421 Operating lease liabilities 3,986 — 3,986 Total current liabilities 31,947 664 32,611 Deferred revenue, net of current portion 1,304 — 1,304 Operating lease liabilities, net of current portion 39,378 — 39,378 Other non-current liabilities 1,225 — 1,225 Total liabilities 73,854 664 74,518 Commitments and contingencies (Note 16) Stockholders’ equity: Common stock, $0.001 par value per share: Authorized shares: 120,000,000 shares; Issued and outstanding: 37,566 shares at June 30, 2023. 37 — 37 Additional paid-in capital 772,473 (1) 772,472 Accumulated other comprehensive loss (2,825) 85 (2,740) Accumulated deficit (414,329) (1,692) (416,021) Total stockholders’ equity 355,356 (1,608) 353,748 Total liabilities and stockholders’ equity $ 429,210 $ (944) $ 428,266 As of September 30, 2023 As Previously Reported Adjustments As Restated ASSETS Current assets: Cash and cash equivalents $ 201,261 $ — $ 201,261 Marketable securities 126,449 — 126,449 Accounts receivable, net of allowance for expected credit losses 24,083 — 24,083 Inventory 19,945 912 20,857 Prepaid expenses and other current assets 9,273 (57) 9,216 Total current assets 381,011 855 381,866 Restricted cash 2,647 (45) 2,602 Property and equipment, net 17,517 (230) 17,287 Intangible assets, net 6,003 — 6,003 Operating lease right-of-use assets 19,860 — 19,860 Other non-current assets 2,004 (401) 1,603 Total assets $ 429,042 $ 179 $ 429,221 LIABILITIES AND STOCKHOLDERS' EQUITY Current liabilities: Accounts payable $ 4,786 $ — $ 4,786 Accrued compensation and benefits 9,775 730 10,505 Accrued expenses and other current liabilities 6,672 14 6,686 Deferred revenue 9,827 — 9,827 Operating lease liabilities 4,093 — 4,093 Total current liabilities 35,153 744 35,897 Deferred revenue, net of current portion 1,126 — 1,126 Operating lease liabilities, net of current portion 38,306 — 38,306 Other non-current liabilities 1,105 — 1,105 Total liabilities 75,690 744 76,434 Commitments and contingencies (Note 16) Stockholders’ equity: Common stock, $0.001 par value per share: Authorized shares: 120,000,000 shares; Issued and outstanding: 37,839 shares at September 30, 2023. 38 — 38 Additional paid-in capital 778,615 — 778,615 Accumulated other comprehensive loss (3,214) 85 (3,129) Accumulated deficit (422,087) (650) (422,737) Total stockholders’ equity 353,352 (565) 352,787 Total liabilities and stockholders’ equity $ 429,042 $ 179 $ 429,221 Three Months Ended March 31, 2023 As Previously Reported Adjustments As Restated Revenues: Product revenue $ 19,287 $ 73 $ 19,360 Service and other revenue 8,579 (73) 8,506 Collaboration revenue 368 — 368 Grant revenue 222 — 222 Total revenue 28,456 — 28,456 Costs of goods sold: Cost of product revenue 7,033 846 7,879 Cost of service and other revenue 4,497 87 4,584 Total costs of goods sold and services 11,530 933 12,463 Gross profit 16,926 (933) 15,993 Operating expenses: Research and development 4,720 267 4,987 Selling, general and administrative 20,883 (66) 20,817 Other lease costs 776 24 800 Impairment and restructuring (33) — (33) Total operating expenses 26,346 225 26,571 Loss from operations (9,420) (1,158) (10,578) Interest income 3,449 — 3,449 Other income (expense), net 8 (13) (5) Loss before income taxes (5,963) (1,171) (7,134) Income tax expense (140) — (140) Net loss $ (6,103) $ (1,171) $ (7,274) Net loss per common share, basic and diluted $ (0.16) $ (0.03) $ (0.19) Weighted-average common shares outstanding, basic and diluted 37,327 — 37,327 Three Months Ended June 30, 2023 Six Months Ended June 30, 2023 As Previously Reported Adjustments As Restated As Previously Reported Adjustments As Restated Revenues: Product revenue $ 19,692 $ (143) $ 19,549 $ 38,979 $ (70) $ 38,909 Service revenue 10,552 (82) 10,470 19,131 (155) 18,976 Collaboration and license revenue 629 — 629 997 — 997 Grant revenue 156 — 156 378 — 378 Total revenues 31,029 (225) 30,804 59,485 (225) 59,260 Costs of goods sold and services: Cost of product revenue 7,236 (535) 6,701 14,269 311 14,580 Cost of service and other revenue 4,655 32 4,687 9,152 119 9,271 Total costs of goods sold and services 11,891 (503) 11,388 23,421 430 23,851 Gross profit 19,138 278 19,416 36,064 (655) 35,409 Operating expenses: Research and development 5,946 182 6,128 10,666 449 11,115 Selling, general, and administrative 21,591 (283) 21,308 42,474 (349) 42,125 Other lease costs 1,162 (194) 968 1,938 (170) 1,768 Impairment and restructuring — — — (33) — (33) Total operating expenses 28,699 (295) 28,404 55,045 (70) 54,975 Loss from operations (9,561) 573 (8,988) (18,981) (585) (19,566) Interest income 3,886 — 3,886 7,335 — 7,335 Other expense (154) (60) (214) (146) (73) (219) Loss before income taxes (5,829) 513 (5,316) (11,792) (658) (12,450) Income tax expense (235) — (235) (375) — (375) Net loss $ (6,064) $ 513 $ (5,551) $ (12,167) $ (658) $ (12,825) Net loss per common share, basic and diluted $ (0.16) $ 0.01 $ (0.15) $ (0.33) $ (0.01) $ (0.34) Weighted-average common shares outstanding, basic and diluted 37,494 — 37,494 37,411 — 37,411 Three Months Ended September 30, 2023 Nine Months Ended September 30, 2023 As Previously Reported Adjustments As Restated As Previously Reported Adjustments As Restated Revenues: Product revenue $ 19,660 $ 280 $ 19,940 $ 58,639 $ 210 $ 58,849 Service revenue 10,938 (55) 10,883 30,069 (210) 29,859 Collaboration and license revenue 237 — 237 1,234 — 1,234 Grant revenue 499 — 499 877 — 877 Total revenues 31,334 225 31,559 90,819 — 90,819 Costs of goods sold and services: Cost of product revenue 8,342 (1,133) 7,209 22,611 (822) 21,789 Cost of service and other revenue 5,209 (268) 4,941 14,361 (149) 14,212 Total costs of goods sold and services 13,551 (1,401) 12,150 36,972 (971) 36,001 Gross profit 17,783 1,626 19,409 53,847 971 54,818 Operating expenses: Research and development 7,200 539 7,739 17,866 988 18,854 Selling, general, and administrative 23,595 (45) 23,550 66,069 (394) 65,675 Other lease costs 758 170 928 2,696 — 2,696 Impairment and restructuring — — — (33) — (33) Total operating expenses 31,553 664 32,217 86,598 594 87,192 Loss from operations (13,770) 962 (12,808) (32,751) 377 (32,374) Interest income 4,185 — 4,185 11,520 — 11,520 Other income 2,030 80 2,110 1,884 7 1,891 Loss before income taxes (7,555) 1,042 (6,513) (19,347) 384 (18,963) Income tax expense (203) — (203) (578) — (578) Net loss $ (7,758) $ 1,042 $ (6,716) $ (19,925) $ 384 $ (19,541) Net loss per common share, basic and diluted $ (0.21) $ 0.03 $ (0.18) $ (0.53) $ 0.01 $ (0.52) Weighted-average common shares outstanding, basic and diluted 37,657 — 37,657 37,494 — 37,494 Three Months Ended December 31, 2023 As Previously Reported in the Company's Earnings Release Adjustments As Restated Revenues: Product revenue $ 20,821 $ — $ 20,821 Service and other revenue 10,230 — 10,230 Collaboration revenue 146 — 146 Grant revenue 352 — 352 Total revenue 31,549 — 31,549 Costs of goods sold: Cost of product revenue 10,025 (2,711) 7,314 Cost of service and other revenue 4,725 104 4,829 Total costs of goods sold and services 14,750 (2,607) 12,143 Gross profit 16,799 2,607 19,406 Operating expenses: Research and development 6,991 219 7,210 Selling, general and administrative 24,172 (736) 23,436 Other lease costs 1,016 — 1,016 Impairment and restructuring 1,570 (209) 1,361 Total operating expenses 33,749 (726) 33,023 Loss from operations (16,950) 3,333 (13,617) Interest income 4,319 — 4,319 Other income 363 263 626 Loss before income taxes (12,268) 3,596 (8,672) Income tax expense (141) — (141) Net loss $ (12,409) $ 3,596 $ (8,813) Net loss per common share, basic and diluted $ (0.33) $ 0.10 $ (0.23) Weighted-average common shares outstanding, basic and diluted 37,890 — 37,890 QUANTERIX CORPORATION RESTATED CONSOLIDATED STATEMENTS OF COMPREHENSIVE LOSS (amounts in thousands) Three Months Ended March 31, 2023 As Previously Reported Adjustments As Restated Net loss $ (6,103) $ (1,171) $ (7,274) Other comprehensive income, net of tax: Foreign currency translation adjustment 42 — 42 Total other comprehensive income 42 — 42 Comprehensive loss $ (6,061) $ (1,171) $ (7,232) Three Months Ended June 30, 2023 Six Months Ended June 30, 2023 As Previously Reported Adjustments As Restated As Previously Reported Adjustments As Restated Net loss $ (6,064) $ 513 $ (5,551) $ (12,167) $ (658) $ (12,825) Other comprehensive loss, net of tax: Foreign currency translation adjustment (244) — (244) (202) — (202) Total other comprehensive loss (244) — (244) (202) — (202) Comprehensive loss $ (6,308) $ 513 $ (5,795) $ (12,369) $ (658) $ (13,027) Three Months Ended September 30, 2023 Nine Months Ended September 30, 2023 As Previously Reported Adjustments As Restated As Previously Reported Adjustments As Restated Net loss $ (7,758) $ 1,042 $ (6,716) $ (19,925) $ 384 $ (19,541) Other comprehensive loss, net of tax: Unrealized losses on marketable securities (241) — (241) (241) — (241) Foreign currency translation adjustment (148) — (148) (350) — (350) Total other comprehensive loss (389) — (389) (591) — (591) Comprehensive loss $ (8,147) $ 1,042 $ (7,105) $ (20,516) $ 384 $ (20,132) QUANTERIX CORPORATION RESTATED CONSOLIDATED STATEMENTS OF STOCKHOLDERS’ EQUITY (amounts in thousands) Three Months Ended, March 31, 2023 Common Stock As Previously Reported Shares Amount Additional paid-in capital Accumulated other comprehensive income (loss) Accumulated Total stockholders' equity Balance at December 31, 2022 37,280 $ 37 $ 763,688 $ (2,623) $ (402,162) $ 358,940 Issuance of common stock under stock plans, including tax effects 144 — 551 — — 551 Stock-based compensation expense — — 3,902 — — 3,902 Foreign currency translation adjustment — — — 42 — 42 Net loss — — — — (6,103) (6,103) Balance at March 31, 2023 37,424 $ 37 $ 768,141 $ (2,581) $ (408,265) $ 357,332 Adjustments Balance at December 31, 2022 — $ — $ (59) $ 85 $ (1,034) $ (1,008) Issuance of common stock under stock plans, including tax effects — — — — — — Stock-based compensation expense — — 41 — — 41 Foreign currency translation adjustment — — — — — — Net loss — — — — (1,171) (1,171) Balance at March 31, 2023 — $ — $ (18) $ 85 $ (2,205) $ (2,138) As Restated Balance at December 31, 2022 37,280 $ 37 $ 763,629 $ (2,538) $ (403,196) $ 357,932 Issuance of common stock under stock plans, including tax effects 144 — 551 — — 551 Stock-based compensation expense — — 3,943 — — 3,943 Foreign currency translation adjustment — — — 42 — 42 Net loss — — — — (7,274) (7,274) Balance at March 31, 2023 37,424 $ 37 $ 768,123 $ (2,496) $ (410,470) $ 355,194 Three Months Ended June 30, 2023 Common Stock As Previously Reported Shares Amount Additional paid-in capital Accumulated other comprehensive income (loss) Accumulated Total stockholders' equity Balance at March 31, 2023 37,424 $ 37 $ 768,141 $ (2,581) $ (408,265) $ 357,332 Issuance of common stock under stock plans, including tax effects 142 — 139 — — 139 Stock-based compensation expense — — 4,193 — — 4,193 Foreign currency translation adjustment — — — (244) — (244) Net loss — — — — (6,064) (6,064) Balance at June 30, 2023 37,566 $ 37 $ 772,473 $ (2,825) $ (414,329) $ 355,356 Adjustments Balance at March 31, 2023 — $ — $ (18) $ 85 $ (2,205) $ (2,138) Issuance of common stock under stock plans, including tax effects — — — — — — Stock-based compensation expense — — 17 — — 17 Foreign currency translation adjustment — — — — — — Net loss — — — — 513 513 Balance at June 30, 2023 — $ — $ (1) $ 85 $ (1,692) $ (1,608) As Restated Balance at March 31, 2023 37,424 $ 37 $ 768,123 $ (2,496) $ (410,470) $ 355,194 Issuance of common stock under stock plans, including tax effects 142 — 139 — — 139 Stock-based compensation expense — — 4,210 — — 4,210 Foreign currency translation adjustment — — — (244) — (244) Net loss — — — — (5,551) (5,551) Balance at June 30, 2023 37,566 $ 37 $ 772,472 $ (2,740) $ (416,021) $ 353,748 Six Months Ended June 30, 2023 Common Stock As Previously Reported Shares Amount Additional paid-in capital Accumulated other comprehensive income (loss) Accumulated Total stockholders' equity Balance at December 31, 2022 37,280 $ 37 $ 763,688 $ (2,623) $ (402,162) $ 358,940 Issuance of common stock under stock plans, including tax effects 286 — 690 — — 690 Stock-based compensation expense — — 8,095 — — 8,095 Foreign currency translation adjustment — — — (202) — (202) Net loss — — — — (12,167) (12,167) Balance at June 30, 2023 37,566 $ 37 $ 772,473 $ (2,825) $ (414,329) $ 355,356 Adjustments Balance at December 31, 2022 — $ — $ (59) $ 85 $ (1,034) $ (1,008) Issuance of common stock under stock plans, including tax effects — — — — — — Stock-based compensation expense — — 58 — — 58 Foreign currency translation adjustment — — — — — — Net loss — — — — (658) (658) Balance at June 30, 2023 — $ — $ (1) $ 85 $ (1,692) $ (1,608) As Restated Balance at December 31, 2022 37,280 $ 37 $ 763,629 $ (2,538) $ (403,196) $ 357,932 Issuance of common stock under stock plans, including tax effects 286 — 690 — — 690 Stock-based compensation expense — — 8,153 — — 8,153 Foreign currency translation adjustment — — — (202) — (202) Net loss — — — — (12,825) (12,825) Balance at June 30, 2023 37,566 $ 37 $ 772,472 $ (2,740) $ (416,021) $ 353,748 Three Months Ended, September 30, 2023 Common Stock As Previously Reported Shares Amount Additional paid-in capital Accumulated other comprehensive income (loss) Accumulated Total stockholders' equity Balance at June 30, 2023 37,566 $ 37 $ 772,473 $ (2,825) $ (414,329) $ 355,356 Issuance of common stock under stock plans, including tax effects 273 1 1,799 — — 1,800 Stock-based compensation expense — — 4,343 — — 4,343 Unrealized loss on marketable securities, net of tax — — — (241) — (241) Foreign currency translation adjustment — — — (148) — (148) Net loss — — — — (7,758) (7,758) Balance at September 30, 2023 37,839 $ 38 $ 778,615 $ (3,214) $ (422,087) $ 353,352 Adjustments Balance at June 30, 2023 — $ — $ (1) $ 85 $ (1,692) $ (1,608) Issuance of common stock under stock plans, including tax effects — — — — — — Stock-based compensation expense — — 1 — — 1 Unrealized loss on marketable securities, net of tax — — — — — — Foreign currency translation adjustment — — — — — — Net loss — — — — 1,042 1,042 Balance at September 30, 2023 — $ — $ — $ 85 $ (650) $ (565) As Restated Balance at June 30, 2023 37,566 $ 37 $ 772,472 $ (2,740) $ (416,021) $ 353,748 Issuance of common stock under stock plans, including tax effects 273 1 1,799 — — 1,800 Stock-based compensation expense — — 4,344 — — 4,344 Unrealized loss on marketable securities, net of tax — — — (241) — (241) Foreign currency translation adjustment — — — (148) — (148) Net loss — — — — (6,716) (6,716) Balance at September 30, 2023 37,839 $ 38 $ 778,615 $ (3,129) $ (422,737) $ 352,787 Nine Months Ended, September 30, 2023 Common Stock As Previously Reported Shares Amount Additional paid-in capital Accumulated other comprehensive income (loss) Accumulated Total stockholders' equity Balance at December 31, 2022 37,280 $ 37 $ 763,688 $ (2,623) $ (402,162) $ 358,940 Issuance of common stock under stock plans, including tax effects 559 1 2,489 — — 2,490 Stock-based compensation expense — — 12,438 — — 12,438 Unrealized loss on marketable securities, net of tax — — — (241) — (241) Foreign currency translation adjustment — — — (350) — (350) Net loss — — — — (19,925) (19,925) Balance at September 30, 2023 37,839 $ 38 $ 778,615 $ (3,214) $ (422,087) $ 353,352 Adjustments Balance at December 31, 2022 — $ — $ (59) $ 85 $ (1,034) $ (1,008) Issuance of common stock under stock plans, including tax effects — — — — — — Stock-based compensation expense — — 59 — — 59 Unrealized loss on marketable securities, net of tax — — — — — — Foreign currency translation adjustment — — — — — — Net loss — — — — 384 384 Balance at September 30, 2023 — $ — $ — $ 85 $ (650) $ (565) As Restated Balance at December 31, 2022 37,280 $ 37 $ 763,629 $ (2,538) $ (403,196) $ 357,932 Issuance of common stock under stock plans, including tax effects 559 1 2,489 — — 2,490 Stock-based compensation expense — — 12,497 — — 12,497 Unrealized loss on marketable securities, net of tax — — — (241) — (241) Foreign currency translation adjustment — — — (350) — (350) Net loss — — — — (19,541) (19,541) Balance at September 30, 2023 37,839 $ 38 $ 778,615 $ (3,129) $ (422,737) $ 352,787 QUANTERIX CORPORATION RESTATED CONSOLIDATED STATEMENTS OF CASH FLOWS (amounts in thousands) Three Months Ended March 31, 2023 As Previously Reported Adjustments As Restated Operating activities: Net loss $ (6,103) $ (1,171) $ (7,274) Adjustments to reconcile net loss to net cash used in operating activities: Depreciation and amortization expense 1,439 15 1,454 Credit losses on accounts receivable 178 — 178 Non-cash operating lease expense 334 — 334 Stock-based compensation expense 3,902 41 3,943 Deferred taxes 270 — 270 Changes in operating assets and liabilities: Accounts receivable (3,741) — (3,741) Inventory (89) 619 530 Prepaid expenses and other assets (422) — (422) Other non-current assets (33) — (33) Accounts payable (1,271) — (1,271) Accrued compensation and benefits, other accrued expenses and other current liabilities (5,983) 174 (5,809) Deferred revenue 2,041 — 2,041 Operating lease liabilities 179 — 179 Other non-current liabilities (203) — (203) Net cash used in operating activities (9,502) (322) (9,824) Investing activities Purchases of property and equipment (136) — (136) Net cash used in investing activities (136) — (136) Financing activities Proceeds from stock options exercised 13 — 13 Proceeds from ESPP purchase 551 — 551 Payments for employee taxes on units withheld (13) — (13) Net cash provided by financing activities 551 — 551 Net decrease in cash, cash equivalents and restricted cash (9,087) (322) (9,409) Effect of foreign currency exchange rate on cash 24 — 24 Cash, cash equivalents and restricted cash at beginning of period 341,337 — 341,337 Cash, cash equivalents and restricted cash at end of period $ 332,274 $ (322) $ 331,952 Supplemental disclosure of cash flow information: Cash paid for income taxes $ 246 $ — $ 246 Noncash transactions: Purchases of property and equipment included in accounts payable and accrued expenses $ 147 $ — $ 147 Reconciliation of cash, cash equivalents and restricted cash: Cash and cash equivalents $ 329,354 $ — $ 329,354 Restricted cash $ 2,920 $ (322) $ 2,598 Total cash, cash equivalents, and restricted cash $ 332,274 $ (322) $ 331,952 Six Months Ended June 30, 2023 As Previously Reported Adjustments As Restated Cash flows from operating activities: Net loss $ (12,167) $ (658) $ (12,825) Adjustments to reconcile net loss to net cash used in operating activities: Depreciation and amortization expense 2,845 38 2,883 Credit losses on accounts receivable 324 — 324 Operating lease right-of-use asset amortization 1,002 — 1,002 Stock-based compensation expense 8,095 58 8,153 Other operating activity 548 54 602 Loss on disposal of fixed assets Impairment — 54 54 Changes in assets and liabilities: Accounts receivable (5,750) — (5,750) Inventory (1,181) 1 (1,180) Prepaid expenses and other current assets (527) — (527) Other non-current assets (965) 258 (707) Accounts payable (631) — (631) Accrued compensation and benefits, accrued expenses, and other current liabilities (1,326) 273 (1,053) Deferred revenue 1,666 — 1,666 Operating lease liabilities (730) — (730) Other non-current liabilities (72) — (72) Net cash used in operating activities (8,869) 78 (8,791) Cash flows from investing activities: Purchases of property and equipment (784) (163) (947) Net cash used in investing activities (784) (163) (947) Cash flows from financing activities: Proceeds from common stock issued under stock plans 777 — 777 Payments for employee taxes withheld on stock-based compensation awards (87) — (87) Net cash provided by financing activities 690 — 690 Net decrease in cash, cash equivalents, and restricted cash (8,963) (85) (9,048) Effect of exchange rate changes on cash, cash equivalents, and restricted cash (163) — (163) Cash, cash equivalents, and restricted cash at beginning of period 341,337 — 341,337 Cash, cash equivalents, and restricted cash at end of period $ 332,211 $ (85) $ 332,126 Supplemental disclosure of cash flow information: Cash paid for taxes $ 502 $ — $ 502 Shares received as consideration under product sales agreement (Note 4, 7) $ 1,000 $ — $ 1,000 Nine Months Ended September 30, 2023 As Previously Reported Adjustments As Restated Cash flows from operating activities: Net loss $ (19,925) $ 384 $ (19,541) Adjustments to reconcile net loss to net cash used in operating activities: Depreciation and amortization expense 4,788 55 4,843 Credit losses on accounts receivable 311 — 311 Foreign currency losses 359 — 359 Unrealized losses on marketable securities (241) — (241) Amortization of (discount) premium on marketable securities (1,249) — (1,249) Operating lease right-of-use asset amortization 1,518 — 1,518 Stock-based compensation expense 12,438 59 12,497 Impairment — 130 130 Deferred income taxes 242 — 242 Loss on disposal of fixed assets 46 — 46 Accounts receivable (5,615) — (5,615) Inventory (2,966) (734) (3,700) Prepaid expenses and other current assets (2,829) — (2,829) Other non-current assets (716) 14 (702) Accounts payable 948 — 948 Accrued compensation and benefits, accrued expenses, and other current liabilities 876 353 1,229 Deferred revenue 894 — 894 Operating lease liabilities (1,690) — (1,690) Other non-current liabilities (107) — (107) Net cash used in operating activities (12,918) 261 (12,657) Cash flows from investing activities: Purchases of marketable securities (125,200) — (125,200) Proceeds from marketable debt securities Purchases of property and equipment (1,572) (306) (1,878) Proceeds from RADx grant on assets purchased Net cash used in investing activities (126,772) (306) (127,078) Cash flows from financing activities: Proceeds from common stock issued under stock plans 2,632 — 2,632 Payments for employee taxes withheld on stock-based compensation awards (142) — (142) Net cash provided by financing activities 2,490 — 2,490 Net decrease in cash, cash equivalents, and restricted cash (137,200) (45) (137,245) Effect of exchange rate changes on cash, cash equivalents, and restricted cash (229) — (229) Cash, cash equivalents, and restricted cash at beginning of period 341,337 — 341,337 Cash, cash equivalents, and restricted cash at end of period $ 203,908 $ (45) $ 203,863 Supplemental disclosure of cash flow information: Cash paid for taxes $ 719 $ — $ 719 Shares received as consideration under product sales agreement (Note 4, 7) $ 775 $ — $ 775 QUANTERIX CORPORATION RESTATED CONSOLIDATED BALANCE SHEETS (amounts in thousands, except per share data) As of March 31, 2022 As Previously Reported Adjustments As Restated ASSETS Current assets: Cash and cash equivalents $ 374,317 $ 226 $ 374,543 Accounts receivable, net of allowance for expected credit losses 22,616 — 22,616 Inventory 22,669 1,757 24,426 Prepaid expenses and other current assets 14,104 (226) 13,878 Total current assets 433,706 1,757 435,463 Restricted cash 2,577 — 2,577 Property and equipment, net 19,683 (44) 19,639 Intangible assets, net 9,692 — 9,692 Goodwill 9,323 — 9,323 Right-of-use assets 29,298 617 29,915 Other non-current assets 378 (300) 78 Total assets $ 504,657 $ 2,030 $ 506,687 LIABILITIES AND STOCKHOLDERS' EQUITY Current liabilities: Accounts payable $ 4,262 $ (75) $ 4,187 Accrued compensation and benefits 8,139 57 8,196 Other accrued expenses 8,024 — 8,024 Deferred revenue 9,194 — 9,194 Short term lease liabilities 1,886 — 1,886 Other current liabilities 268 — 268 Total current liabilities 31,773 (18) 31,755 Deferred revenue, net of current portion 1,222 — 1,222 Long term lease liabilities 43,563 — 43,563 Other non-current liabilities 1,691 — 1,691 Total liabilities 78,249 (18) 78,231 Commitments and contingencies (Note 16) Stockholders’ equity: Common stock, $0.001 par value: Authorized shares: 120,000 shares; Issued and outstanding: 36,899 shares at March 31, 2022. 37 — 37 Additional paid-in capital 750,742 (25) 750,717 Accumulated other comprehensive (loss) income (756) — (756) Accumulated deficit (323,615) 2,073 (321,542) Total stockholders’ equity 426,408 2,048 428,456 Total liabilities and stockholders’ equity $ 504,657 $ 2,030 $ 506,687 As of June 30, 2022 As Previously Reported Adjustments As Restated ASSETS Current assets: Cash and cash equivalents $ 361,293 $ 258 $ 361,551 Accounts receivable, net of allowance for expected credit losses 19,683 — 19,683 Inventory 21,985 700 22,685 Prepaid expenses and other current assets 10,237 (258) 9,979 Total current assets 413,198 700 413,898 Restricted cash 2,594 — 2,594 Property and equipment, net 22,295 (319) 21,976 Intangible assets, net 8,527 — 8,527 Goodwill 8,675 — 8,675 Right-of-use assets 32,935 — 32,935 Other non-current assets 377 (300) 77 Total assets $ 488,601 $ 81 $ 488,682 LIABILITIES AND STOCKHOLDERS' EQUITY Current liabilities: Accounts payable $ 5,983 $ (75) $ 5,908 Accrued compensation and benefits 10,638 139 10,777 Other accrued expenses 4,799 — 4,799 Deferred revenue 12,157 — 12,157 Short term lease liabilities 2,696 — 2,696 Other current liabilities 396 — 396 Total current liabilities 36,669 64 36,733 Deferred revenue, net of current portion 1,531 — 1,531 Long term lease liabilities 43,135 — 43,135 Other non-current liabilities 1,810 — 1,810 Total liabilities 83,145 64 83,209 Commitments and contingencies (Note 16) Stockholders’ equity: Common stock, $0.001 par value: Authorized shares: 120,000 shares; Issued and outstanding: 36,975 shares at June 30, 2022. 37 — 37 Additional paid-in capital 756,139 (52) 756,087 Accumulated other comprehensive (loss) income (2,203) — (2,203) Accumulated deficit (348,517) 69 (348,448) Total stockholders’ equity 405,456 17 405,473 Total liabilities and stockholders’ equity $ 488,601 $ 81 $ 488,682 As of September 30, 2022 As Previously Reported Adjustments As Restated ASSETS Current assets: Cash and cash equivalents $ 343,743 $ — $ 343,743 Accounts receivable, net of allowance for expected credit losses 18,330 — 18,330 Inventory 18,236 (192) 18,044 Prepaid expenses and other current assets 6,475 (400) 6,075 Total current assets 386,784 (592) 386,192 Restricted cash 2,596 — 2,596 Property and equipment, net 21,441 (246) 21,195 Intangible assets, net 7,511 — 7,511 Right-of-use assets 27,165 — 27,165 Other non-current assets 1,200 (290) 910 Total assets $ 446,697 $ (1,128) $ 445,569 LIABILITIES AND STOCKHOLDERS' EQUITY Current liabilities: Accounts payable $ 2,106 $ — $ 2,106 Accrued compensation and benefits 10,503 257 10,760 Other accrued expenses 5,951 90 6,041 Deferred revenue 8,976 — 8,976 Short-term lease liabilities 767 — 767 Other current liabilities 268 — 268 Total current liabilities 28,571 347 28,918 Deferred revenue, net of current portion 1,591 — 1,591 Long-term lease liabilities 42,196 — 42,196 Other non-current liabilities 1,570 — 1,570 Total liabilities 73,928 347 74,275 Commitments and contingencies (Note 16) Stockholders’ equity: Common stock, $0.001 par value: Authorized shares: 120,000 shares; Issued and outstanding: 37,094 shares at September 30, 2022. 37 — 37 Additional paid-in capital 759,312 (58) 759,254 Accumulated other comprehensive (loss) income (2,999) — (2,999) Accumulated deficit (383,581) (1,417) (384,998) Total stockholders’ equity 372,769 (1,475) 371,294 Total liabilities and stockholders’ equity $ 446,697 $ (1,128) $ 445,569 QUANTERIX CORPORATION RESTATED CONSOLIDATED STATEMENTS OF OPERATIONS (amounts in thousands, except per share data) Three Months Ended March 31, 2022 As Previously Reported Adjustments As Restated Revenues: Product revenue $ 20,656 $ — $ 20,656 Service and other revenue 8,810 — 8,810 Collaboration revenue 86 — 86 Total revenue 29,552 — 29,552 Costs of goods sold: Cost of product revenue 10,746 711 11,457 Cost of service and other revenue 4,247 (249) 3,998 Total costs of goods sold and services 14,993 462 15,455 Gross profit 14,559 (462) 14,097 Operating expenses: Research and development 7,034 (154) 6,880 Selling, general and administrative 25,712 (541) 25,171 Total operating expenses 32,746 (695) 32,051 Loss from operations (18,187) 233 (17,954) Interest income 52 — 52 Other expense (217) — (217) Loss before income taxes (18,352) 233 (18,119) Income tax benefit 199 — 199 Net loss $ (18,153) $ 233 $ (17,920) Net loss per common share, basic and diluted $ (0.49) $ — $ (0.49) Weighted-average common shares outstanding, basic and diluted 36,851 — 36,851 Three Months Ended June 30, 2022 Six Months Ended June 30, 2022 As Previously Reported Adjustments As Restated As Previously Reported Adjustments As Restated Revenues: Product revenue $ 14,785 $ — $ 14,785 $ 35,441 $ — $ 35,441 Service and other revenue 8,548 — 8,548 17,358 — 17,358 Collaboration revenue 92 — 92 178 — 178 Grant revenue 75 — 75 75 — 75 Total revenue 23,500 — 23,500 53,052 — 53,052 Costs of goods sold: Cost of product revenue 9,921 625 10,546 20,667 1,336 22,003 Cost of service and other revenue 4,868 (231) 4,637 9,115 (480) 8,635 Total costs of goods sold and services 14,789 394 15,183 29,782 856 30,638 Gross profit 8,711 (394) 8,317 23,270 (856) 22,414 Operating expenses: Research and de

Significant Accounting Polici_2

Significant Accounting Policies (Policies)	12 Months Ended
	Dec. 31, 2023
Accounting Policies [Abstract]
Basis of Presentation	Basis of Presentation The accompanying Consolidated Financial Statements and notes herein have been prepared in accordance with generally accepted accounting principles in the United States of America (“U.S. GAAP”) and pursuant to the rules and regulations of the U.S. Securities and Exchange Commission (“SEC”) regarding annual financial reporting on Form 10-K.
Use of Estimates	Use of Estimates The preparation of the Consolidated Financial Statements and Notes to Consolidated Financial Statements in conformity with U.S. GAAP requires management to make estimates and assumptions that affect the reported amounts of assets and liabilities at the dates of the Consolidated Financial Statements, and the reported amounts of revenues and expenses during each fiscal year. Such estimates include, but are not limited to, revenue recognition, valuation of inventory, leases, valuation and impairment of long-lived assets, recoverability of deferred tax assets, and stock-based compensation expense. The Company bases its estimates on historical experience, known trends, worldwide economic conditions, both general and specific to the life sciences industry, and other relevant factors it believes to be reasonable under the circumstances. On an ongoing basis, management evaluates its estimates and changes in estimates are recorded in the period in which they become known. Actual results could differ from those estimates.
Principles of Consolidation	Principles of Consolidation The Consolidated Financial Statements include the accounts of Quanterix and its wholly-owned subsidiaries. All intercompany transactions have been eliminated in consolidation. In accordance with Accounting Standards Codification (“ASC”) 810 – Consolidation , the Company assesses the terms of its non-marketable equity investments in entities to determine if any meet the definition of a variable interest entity (“VIE”) and require consolidation into its Consolidated Financial Statements. Refer to Note 20 - Variable Interest Entities for further discussion.
Foreign Currency (As Restated)	Foreign Currency (As Restated) The functional currency of the Company’s subsidiaries is their respective local currencies. These subsidiary financial statements are translated into U.S. dollars using the period-end exchange rates for assets and liabilities, average exchange rates during the corresponding period for revenue and expenses, and historical rates for equity. The effects of foreign currency translation adjustments are recorded in accumulated other comprehensive income (loss), a component of stockholders’ equity on the Consolidated Balance Sheets. Foreign currency transaction gains (losses) are included in other income (expense), net on the Consolidated Statements of Operations. Foreign exchange losses were $0.2 million, $1.0 million and $0.4 million during the years ended December 31, 2023, 2022 and 2021, respectively.
Segment Information	Segment Information Operating segments are defined as components of an enterprise about which separate discrete information is available for evaluation by the chief operating decision-maker (“CODM”), in deciding how to allocate resources and assess performance. The Company’s CODM is the chief executive officer, who reviews the Company's operations and manages its business as a single operating segment as of December 31, 2023.
Revenue from Contracts with Customers	Revenue from Contracts with Customers The Company generates the majority of its revenues from contracts with customers and accounts for them pursuant to the Financial Accounting Standards Board (“FASB”) Accounting Standards Codification Topic 606 - Revenue from Contracts with Customers (“ASC 606”). Refer to the section titled “Grant Revenue” below for discussion the Company’s accounting policy for revenue generated from grant awards. For contracts with customers, the Company recognizes revenue when a customer obtains control of promised products or services, for an amount that reflects the consideration expected to be received in exchange for those products or services. The Company follows the five step revenue model prescribed by ASC 606 to determine revenue recognition: (i) identify the contract(s) with a customer; (ii) identify the performance obligations in the contract; (iii) determine the transaction price, including variable consideration, if any; (iv) allocate the transaction price to the performance obligations in the contract; and (v) recognize revenue when (or as) the entity satisfies a performance obligation. Revenues are presented net of any sales, value added, or similar taxes collected from customers and remitted to the government. The Company accounts for a contract when it has approval and commitment from both parties, the fees, payment terms and rights of the parties regarding the products or services to be transferred are identified, the contract has commercial substance, and it is probable that substantially all of the consideration for the products and services expected to be transferred is collectible. The Company applies judgment in determining the customer’s ability and intention to pay for services expected to be transferred, which is based on factors including the customer’s payment history, management’s ability to mitigate exposure to credit risk (for example, requiring payment in advance of the transfer of products or services, or the ability to stop transferring promised products or services in the event a customer fails to pay consideration when due), and experience selling to similarly situated customers. The Company’s contracts may include either a single promise (referred to as a performance obligation) to transfer a product or service, or a combination of multiple performance obligations to transfer products or services. The Company evaluates the existence of multiple performance obligations within its contracts by using judgment to determine if (1) the customer can benefit from each contractual promise on its own or together with readily available resources and (2) the transfer of each contractual promise is separately identifiable from other promises in a contract. When both criteria are met, each promise is accounted for as a separate performance obligation. Additionally, the Company has elected the practical expedient under ASC 606 to account for shipping and handling as an activity to fulfill a promise to transfer a product, and therefore does not evaluate whether shipping and handling activities are promised services to its customers. Contracts that include rights to additional products or services that are exercisable at a customer’s discretion are generally considered options. The Company assesses if these options provide a material right to the customer and if so, the material right is considered a performance obligation. The identification of material rights requires judgment to determine the value of the option to purchase additional products and services in relation to options that may be provided to, and prices paid by, customers in the normal course of business. Material rights are recognized when exercised by a customer or upon expiration of the right. The Company determines the transaction price of its contracts based on the amount of consideration it expects to be entitled to, which is generally equal to the contract amount. In some cases, contracts contain variable consideration which primarily relates to (1) sales- and usage-based royalties related to the license of intellectual property in collaboration and license contracts and (2) contracts with minimum purchase commitments. For sales and usage-based royalties, ASC 606 provides an exception to estimating variable consideration. Under this exception, the Company recognizes revenues from sales- or usage-based royalty revenue at the later of when the sales or usage occurs or the satisfaction (or partial satisfaction) of the performance obligation to which the royalty has been allocated. All other variable amounts are constrained to the minimum guaranteed contract amount so that a reversal of cumulative revenue does not occur in future periods. Once there is no longer uncertainty over a variable amount, any incremental fees the Company is entitled to are allocated to the related performance obligation(s). For contracts that contain multiple performance obligations, the Company allocates the transaction price among the performance obligations on a relative basis according to their standalone selling prices (“SSP”). Determining the SSP for performance obligations requires judgment and is based on factors including prices charged to customers in observable transactions, internal pricing objectives and list prices, and pricing of similar products. The Company uses a range of amounts to estimate SSP and has more than one range for certain products and services based on the geographic location of customers and sales channel. Product Revenue The Company’s product revenues are composed of instruments, assay kits, replacement parts, and other consumables such as reagents and antibodies. Products are sold directly to customers and are also sold through distributors in EMEA and Asia Pacific regions. Direct instrument sales include installation and an initial year of implied service-type warranties. The Company has determined that the instrument and installation are a combined performance obligation as the customer cannot benefit from the instrument without the installation and no other vendors can provide the installation of the Company’s specialized instruments. The implied service-type warranty is considered a separate performance obligation since a customer could benefit from it independently with readily available resources and is capable of being sold on its own. Sales of instruments to distributors include a license to import and resell the instruments. The Company has determined these distributor licenses are part of a combined performance obligation with the instrument as the distributor only benefits from the combination of the instrument and ability to resell it. Instrument sales may also be bundled with assays and other consumables, training, and/or an extended service warranty, each of which is considered a separate performance obligation. Product revenues for direct instrument sales to customers are recognized upon completion of the instrument’s installation. For instrument sales to distributors, revenue is recognized based on the agreed upon shipping terms (either upon shipment or delivery) as that is when title passes to the customer. Service Revenue Service revenues are composed of contract research services, initial year of implied service-type warranties, extended services warranty contracts, repair services, and other services such as training. Contract research services are provided through the Company’s Accelerator Laboratory and generally consist of fixed fee contracts. Revenues from contract research services are recognized at the point in time when the Company completes and delivers its research results on each individually completed study, or over time if the contractual provisions allow for the collection of transaction consideration for costs incurred plus a reasonable margin through the period of performance of the services. For contract research services recognized over time, the Company uses the output method based on the number of completed results. Revenues from other services are recognized at the point in time when the training or other services are delivered as the customer simultaneously receives and benefits from the services. Revenues from service-type warranties are recognized ratably over the contract service period. Collaboration and License Revenue Collaboration and license revenues are composed of revenue associated with licensing to third parties the Company’s technology, intellectual property, and know-how associated with our instruments and for related services. License arrangements consist of sales or usage-based fees and/or future royalties. Revenues are recognized at the point in time the license is delivered as the underlying license is considered functional intellectual property. Royalty revenues that are sales- or usage- based are recognized at the later of when the sales or usage occurs and the satisfaction (or partial satisfaction) of the performance obligation to which the royalty has been allocated.
Contract Assets and Liabilities	Contract Assets and Liabilities Accounts Receivable and Allowance for Credit Losses The Company is exposed to credit losses primarily through accounts receivable from sales of its products and services. Accounts receivable includes amounts billed and currently due from customers. Since the only condition for payment of the Company’s invoices is the passage of time, the Company records a receivable on the date the invoice is issued. Also included in accounts receivable are unbilled amounts resulting from revenue exceeding the amount billed to the customer, where the right to payment is unconditional. If the right to payment for services performed was conditional on something other than the passage of time, the unbilled amount would be recorded as a separate contract asset. The Company’s expected loss allowance methodology is developed using historical collection experience, current and future economic and market conditions, and a review of the current status of customers’ accounts receivable. Specific allowance amounts are established to record the appropriate provision for customers that have a higher probability of default. The Company monitors changes to the receivables balance on a timely basis, and balances are written off as they are determined to be uncollectable after collection efforts have been exhausted. Generally, the Company’s contracts are non-cancellable. For contracts that are cancellable by the customer, the Company does not record a receivable when it issues an invoice. The Company records accounts receivable on these contracts only up to the amount of revenue recognized but not yet collected. The Company’s payment terms vary by the type and location of the customer and the products or services offered. Payment from customers is generally required 30 to 45 days from date of shipment or satisfaction of the performance obligation. The Company does not provide financing arrangements to its customers. The Company refers to contract liabilities as deferred revenue on the Consolidated Balance Sheets. For deferred revenue, the Company applies the practical expedient under ASC 606 that allows for the exclusion of (1) contracts with original expected length of one year or less and (2) contracts for which revenue is recognized at the amount to which the Company has the right to invoice for services performed from the amount of remaining performance obligations disclosed.
Costs to Obtain Contracts	Costs to Obtain Contracts The Company capitalizes commissions paid to its sales representatives and related fringe benefit costs that are incremental to obtaining customer contracts. Capitalized commissions are recorded in prepaid expenses and other current assets and other non-current assets on the Consolidated Balance Sheets. These commissions are amortized over the life of the contract and are recorded in cost of goods sold and selling, general and administrative expense on the Consolidated Statements of Operations. The Company has elected the practical expedient allowing commissions with an amortization period of one year or less to be expensed as incurred.
Warranties	Warranties The Company provides an initial year of warranty and maintenance service related to its instruments sold directly to customers and sells extended warranty contracts for additional periods. The Company defers revenue associated with these warranty services and recognizes them ratably over the service period.
Grant Revenue	Grant Revenue Accounting for grants does not fall under ASC 606, as the grantor will not benefit directly from the Company’s expansion or product development, and no products or services are transferred to the grantor. As there is no authoritative guidance under U.S. GAAP on accounting for grants to for profit business entities from government entities, the Company accounts for grants by analogy to International Accounting Standards Topic 20, Accounting for Government Grants and Disclosure of Government Assistance (“IAS 20”) and ASC Topic 958, Not for-Profit Entities (“ASC 958”). The decision to account for a grant under IAS 20 or ASC 958 is based on whether the grantor is a government entity. The Company records grants related to assets as a deduction in calculating the carrying value of the asset, and to record grants related to income separately on the Consolidated Statements of Operations on a gross basis as grant revenue. The related expenses are recorded on a gross basis within operating expenses. These methods are elections an entity can make under both IAS 20 and ASC 958. The Company recognizes grant revenue as the Company performs services under the arrangement when the funding is committed and as each grant’s activities are performed. The timing of revenue recognition and receipt of funding varies by grant and can be independent from performance of the related activities, such as an upfront payment of the award value, or subsequent to the Company’s requests for reimbursement for already performed activities (subject to the approval of the granting organization).
Cost of Goods Sold and Services	Cost of Goods Sold and Services Cost of Product Revenue Cost of product revenue consists of manufacturing and assembly costs for instruments, related reagents, other consumables, contract manufacturer costs, personnel costs, royalties, overhead, and other direct costs related to product sales. Raw material part costs include inbound freight, shipping and handling costs associated with purchased goods, contract manufacturer costs, personnel costs, royalties, overhead and other direct costs related to product sales. Additionally included in cost of product revenue are royalty fees due to third parties from revenue generated by collaboration or license deals. Cost of Service and Other Revenue Cost of services and other revenue consists of direct costs associated with operating the Company’s Accelerator Laboratory on behalf of its customers, including raw materials, personnel costs, royalties, allocated overhead costs that include facility and other related costs, and other direct costs. Additional costs include costs related to warranties, and other costs of servicing equipment at customer sites.
Research and Development Expenses	Research and Development Expenses Research and development expense consists primarily of personnel costs, research supplies, third-party development costs for new products, materials for prototypes, quality assurance, and allocated overhead costs that include facility and other related costs. The Company accounts for nonrefundable advance payments for products and services that will be used in future research and development activities as expense when the service has been performed or when the products have been received. For arrangements in which the Company receives funding from third parties for research and development activities (excluding the government sponsored arrangements), the Company assesses whether the arrangement is within the scope of ASC 730 – Research and Development . When the Company is entitled to receive reimbursements, does not have an obligation to repay, does not transfer products or services, and is the primary beneficiary of the activities, the Company records the reimbursable amounts as a reduction to research and development expense. Amounts reimbursed in excess of the cost incurred by the Company are recorded within other income (expense), net on the Consolidated Statements of Operations. Reimbursable amounts recorded as a reduction to research and development expenses were not material during the years ended December 31, 2023, 2022, and 2021. Amounts reimbursed in excess of costs incurred by the Company related to activities funded by third parties were $0.4 million during the year ended December 31, 2023. No reimbursable amounts were received during the years ended December 31, 2022, and 2021.
Selling, General and Administrative Expenses	Selling, General and Administrative Expenses
Net Loss Per Share	Net Loss Per Share Basic net loss per common share attributable to common stockholders is calculated by dividing the loss attributable to common stockholders by the weighted-average number of common shares outstanding. Diluted net loss per common share attributable to common stockholders is calculated under the treasury stock method by dividing the loss attributable to common stockholders by the diluted weighted-average number of common shares outstanding. Diluted weighted-average shares outstanding reflect the dilutive effect, if any, of potential common shares issued, such as unvested common stock, unvested restricted stock units (“RSUs”), common stock options, and shares estimated to be purchased under the Company’s employee stock purchase plan (“ESPP”). During periods when the Company is in a net loss position, these potential common shares are excluded from the diluted net loss per common share attributable to common stockholders because their effect would be anti-dilutive. Accordingly, basic and diluted net loss per common share attributable to common stockholders were the same for all periods presented.
Cash and Cash Equivalents	Cash and Cash Equivalents Cash and cash equivalents consist of cash deposits and short-term, highly liquid marketable securities that are readily convertible into cash, with original maturities of three months or less. Cash and cash equivalents consist of the following (in thousands): As of December 31, 2023 2022 Cash $ 12,162 $ 32,643 Money market funds 155,367 306,097 Marketable securities 6,893 — Total cash and cash equivalents $ 174,422 $ 338,740
Restricted Cash	Restricted Cash The following table summarizes the period ending cash and cash equivalents as presented on the Consolidated Balance Sheets and the total cash, cash equivalents, and restricted cash as presented on the Consolidated Statements of Cash Flows (in thousands): As of December 31, 2023 2022 Cash and cash equivalents $ 174,422 $ 338,740 Restricted cash 2,604 2,597 Cash, cash equivalents, and restricted cash $ 177,026 $ 341,337 Restricted cash consists of collateral for a letter of credit issued as security for two of the Company’s leased facilities and to secure the Company’s corporate credit card program. The short-term or long-term classification is determined in accordance with the expiration of the underlying letter of credit and security.
Marketable Securities	Marketable Securities The Company’s current portfolio of marketable securities is entirely debt securities and may at any time include commercial paper, U.S. Treasuries, corporate notes and bonds, U.S. Government agency bonds, certificates of deposit, and similar types of debt securities. Marketable debt securities with original maturities of three months or less at the time of purchase are recorded in cash equivalents on the Consolidated Balance Sheets as they are considered highly liquid and readily convertible into cash. All other marketable securities, including those with maturities beyond one year, are recorded as current assets on the Consolidated Balance Sheets based on their highly liquid nature and because such securities are available for use in current operations. The Company classifies its marketable securities as either held to maturity, available-for-sale, or trading at the time of purchase and re-evaluates such classification at each balance sheet date. All of the Company’s marketable securities are currently classified as available-for-sale as it may use them in current operations. Available-for-sale securities are recorded at fair value (refer to Note 7 - Fair Value of Financial Instruments ). Unrealized gains and losses (other than impairment or credit related losses) are recorded in accumulated other comprehensive income (loss), a component of stockholders’ equity on the Consolidated Balance Sheets. Realized gains and losses are determined using the specific identification method and are recorded in other income (expense), net on the Consolidated Statements of Operations. Quarterly, or more frequently if circumstances warrant, the Company monitors its marketable securities for impairment. In the event a security’s fair value is less than its amortized cost basis, the Company evaluates whether an impairment exists and if the impairment is a result of credit loss or other factors. For a security in an unrealized loss position, if the Company intends to sell the security in an unrealized loss position, or it is more likely than not that the Company will be required to sell the security before recovery of its amortized cost basis, an impairment loss equal to the difference between the security’s fair value and amortized cost basis is recorded in other income (expense), net. Additionally, the Company determines if a credit loss exists by considering information about the collectability of the security, current market conditions, and the issuer’s financial condition. If a decline in fair value is a result of a credit loss, an allowance for credit losses is recorded in other income (expense), net on the Consolidated Statement of Operations, limited to the portion attributed to the credit loss. The Company has also elected the practical expedient to separately present accrued interest receivable from its marketable securities balance. Such accrued interest is recorded in prepaid expenses and other current assets on the Consolidated Balance Sheets and is not included in the assessment and measurement of impairment of its marketable securities.
Inventory	Inventory Inventory consists of instruments, assays, and the materials required to manufacture instruments and assays. Inventory is stated at the lower of cost or net realizable value on a first-in, first-out (“FIFO”) basis and includes the cost of materials, labor, and manufacturing overhead. The Company analyzes its inventory levels on each reporting date for slow-moving, excess, and obsolete inventory, and inventory expected to expire prior to being used. These analyses require judgment and are based on factors including, but not limited to, recent historical activity, anticipated or forecasted demand for the Company’s products (developed through its planning and sales and marketing inputs, and market conditions). If the Company identifies adverse conditions exist, the carrying value of the inventory is reduced to its estimated net realizable value by providing estimated reserves for excess or obsolete inventory. The Company adjusts the reserves for excess or obsolete inventory and records additional inventory write downs based on unfavorable changes in estimated customer demand or actual market conditions that may differ from its projections.
Property and Equipment	Property and Equipment Property and equipment, including leasehold improvements, are stated at cost, net of accumulated depreciation. These assets are depreciated over their estimated useful lives using the straight-line method. Expenditures for maintenance and repairs are charged to expense as incurred, whereas significant expenditures that extend the useful lives of existing assets are capitalized as additions to property and equipment. Depreciation is calculated based upon the following estimated useful lives: Estimated Useful Life Laboratory and manufacturing equipment 5 years Office furniture and equipment 7 years Computers and software 3 years Leasehold improvements Shorter of asset's life or remaining lease term
Leases	Leases The Company enters into operating leases for office, laboratory, and manufacturing spaces, as well as office equipment, and determines whether an arrangement is a lease at inception of the arrangement. The Company accounts for a lease when it has the right to control the leased asset for a period of time, while obtaining substantially all of the assets’ economic benefits. Leases are recorded on the Consolidated Balance Sheets as operating lease right-of-use (“ROU”) assets and current or non-current operating lease liabilities, as applicable. All of the Company’s leases are classified as operating leases. Additionally, the Company does not separate lease and non-lease components. Operating lease ROU assets and lease liabilities are recognized at the lease commencement date based on the estimated present value of the future minimum lease payments over the lease term and any initial direct costs incurred. Initial direct costs are incremental costs of a lease that would not have been incurred had the lease not been executed. The discount rate used to determine the present value of the lease payments is the Company’s incremental borrowing rate on a collateralized basis for a similar term and amount, as generally an implicit rate in the lease is not readily determinable. To estimate its incremental borrowing rate, a credit rating applicable to the Company is estimated using a synthetic credit rating analysis since the Company does not currently have an agency-based credit rating. The Company’s lease agreements can contain lease and non-lease components. The Company accounts for the lease and fixed payments for non-lease components as a single lease component under ASC 842– Leases , which increases the amount of the ROU assets and lease liabilities. Most of the Company’s lease agreements also contain variable payments, primarily maintenance, utility, and other-related costs, which are expensed as incurred and not included in the measurement of the ROU assets and lease liabilities. Some of the Company’s leases contain options to extend or terminate the lease. When determining the lease term, these options are included in the measurement and recognition of the Company’s ROU assets and lease liabilities when it is reasonably certain that the Company will exercise the option(s). The Company considers various economic factors when making this determination, including, but not limited to, the significance of leasehold improvements incurred in the leased space, the difficulty in replacing the asset, underlying contractual obligations, and specific characteristics unique to a particular lease. Subsequent to entering into a lease arrangement, the Company reassesses the certainty of exercising options to extend or terminate a lease. When it becomes reasonably certain that the Company will exercise an option that was not included in the lease term, the Company accounts for the change in circumstances as a lease modification, which results in the remeasurement of the ROU asset and lease liability as of the modification date. Leases with a term of 12 months or less upon commencement are not recorded on the Consolidated Balance Sheets and are recorded to expense on a straight-line basis over the lease term.
Impairment of Goodwill	Impairment of Goodwill The Company assesses goodwill for impairment at the reporting unit level annually and whenever events occur or circumstances change that would more likely than not reduce the fair value of a reporting unit below its carrying amount. Such events or circumstances could include the occurrence of operating losses, a significant decline in earnings, or significant changes in or restructuring of the business. The impairment test is first performed at the reporting unit level using a qualitative assessment to determine whether it is more likely than not that the fair value of the reporting unit is less than the carrying value. If the reporting unit does not pass the qualitative assessment, the reporting unit’s carrying value is compared to its fair value, using estimates including forecasts of discounted future cash flows and peer market multiples. An impairment charge is recorded equal to the excess of the reporting unit’s carrying value over its fair value.
Impairment of Long-Lived Assets	Impairment of Long-Lived Assets The Company’s long-lived assets consist of operating lease ROU assets, property and equipment, and intangible assets. The Company reviews the carrying amount of its long-lived assets for impairment whenever events or circumstances indicate that the estimated useful lives may warrant revision, or that the carrying amount of the assets may not be fully recoverable. To assess whether a long-lived asset or group of assets has been impaired, the estimated undiscounted and discounted future cash flows for the estimated remaining useful life or estimated lease term of the asset (or the primary asset in the asset group) are compared to their carrying values. Significant judgment is required to estimate future cash flows, including, but not limited to, the expected use of the asset (group), historical customer retention rates, technology roadmaps, customer awareness, trademark and trade name history, contractual provisions that could limit or extend an asset's useful life, market data, discount rates, potential sublease opportunities including rent and rent escalation rates, time to sublease, and free rent periods. To the extent that the future cash flows are less than the carrying value, the asset(s) are impaired and written down to its estimated fair value.
Software Development Costs	Software Development Costs The Company develops and modifies software related to the operation of some of its instruments and internal use software supporting the Company’s operations. Certain costs incurred during the application development stage including external direct costs of services used in the development or internal personnel costs for employees directly associated with the development, are capitalized. The Company begins depreciating these costs over the life of the related asset upon completion of a working model or when it is ready for its intended use. Capitalized software development costs related to software owned or developed by the Company are recorded in property and equipment on the Consolidated Balance Sheets. Capitalized software development costs related to software hosted by a vendor (i.e. cloud computing) are recorded in prepaid expenses and other current assets, and other non-current assets on the Consolidated Balance Sheets.
Fair Value of Financial Instruments	Fair Value of Financial Instruments The carrying amount reflected in the Consolidated Balance Sheets for cash, restricted cash, accounts receivable, prepaid expenses and other current assets, accounts payable, accrued compensation and benefits, and accrued expenses and other current liabilities approximate their fair values due to their short-term nature. Additionally, the Company has certain financial assets that are required to be measured at fair value on a recurring basis including cash equivalents and marketable securities. Pursuant to the accounting standards for fair value measurements, the fair values of these financial assets are classified as Level 1, 2, or 3 within the fair value hierarchy as follows: • Level 1: Observable inputs based on unadjusted quoted prices in active markets for identical assets. • Level 2: Inputs, other than Level 1 inputs, that are observable either directly or indirectly, such as quoted prices for similar assets, quoted prices in markets that are not active, other inputs that are observable or can be corroborated by observable market data for substantially the full term of the assets. • Level 3: Unobservable inputs for which there is little or no market data and such inputs are significant to the fair value of the assets. These inputs reflect the Company’s assumptions about the inputs that market participants would use in pricing the asset. To the extent that the valuation is based on models or inputs that are less observable or unobservable in the market, the determination of fair value requires more judgment.
Income Taxes	Income Taxes The Company recognizes deferred tax assets and liabilities for the expected future tax consequences of events that have been recognized in the Consolidated Financial Statements or tax returns. Deferred tax assets and liabilities are determined based on differences between the carrying amount and the tax basis of the assets and liabilities using the enacted tax rates in effect in the years in which the differences are expected to reverse. A valuation allowance against deferred tax assets is recorded if, based on the weight of the available evidence, it is more likely than not that some or all of the deferred tax assets will not be realized. The Company accounts for uncertain tax positions using a more likely than not threshold for recognizing uncertain tax positions, in accordance with ASC 740 – Income Taxes . The evaluation of uncertain tax positions is based on factors that include, but are not limited to, changes in tax law, the measurement of tax positions taken or expected to be taken in tax returns, the effective settlement of matters subject to audit, new audit activity, and changes in facts or circumstances related to a tax position. The Company evaluates uncertain tax positions on an ongoing basis and adjusts any liability recorded to reflect subsequent changes in the relevant facts surrounding the uncertain positions. Amounts recorded for uncertain tax positions, including interest and penalties, are recorded in income tax (expense) benefit on the Consolidated Statement of Operations.
Credit, Product, and Supplier Concentrations and Off-Balance-Sheet Risk	Credit, Product, and Supplier Concentrations and Off-Balance-Sheet Risk Financial instruments that potentially expose the Company to concentrations of credit risk primarily consist of cash, cash equivalents, marketable securities, and accounts receivable. The Company limits its risk exposure by having its cash, cash equivalents, and marketable securities held at large commercial banks. Customers outside the United States represented 38% and 41% of the Company’s gross trade accounts receivable balance as of December 31, 2023 and 2022, respectively. For the year ended December 31, 2023, one customer of approximately $14.0 million revenue accounted for greater than 10% of the Company’s total revenue. For the year ended December 31, 2022, one customer of approximately $13.7 million revenue accounted for greater than 10% of the Company’s total revenue. For the year ended December 31, 2021, no customer individually accounted for greater than 10% of the Company’s total revenue. The Company is also subject to supply chain risks related to outsourced manufacturing of its instruments. Although there are a limited number of manufacturers for its instruments, the Company believes that other suppliers could provide similar manufacturing on comparable terms. A change in suppliers, however, could cause a delay in manufacturing and a possible loss of sales, which would adversely affect operating results. In addition to outsourced manufacturing of its instruments, the Company also purchases antibodies through a number of different suppliers. Although a disruption in service from any one of its antibody suppliers is possible, the Company believes that it would be able to find an adequate supply from alternative suppliers.
Stock-Based Compensation	Stock-Based Compensation The Company measures and recognizes stock-based compensation expense by calculating the estimated fair value of stock options, RSUs, or purchase rights issued under the Company’s ESPP on the grant date. The Company generally issues new common shares upon exercise of options and vesting of RSUs. Awards granted by the Company are routine in nature including new hire, annual, and promotion grants. The fair value of stock options and purchase rights under the ESPP is estimated using the Black-Scholes option-pricing model. The Black-Scholes model requires the Company to make assumptions about the expected or contractual term of the option or purchase right, the expected volatility, risk-free interest rates, and expected dividend yield. The Company estimates the expected term of options granted to employees utilizing historical exercise data. The expected term is applied to the stock option grant group as a whole, as the Company does not expect substantially different exercise or post-vesting termination behavior among its employee population. During 2023, the expected volatility was based on the Company’s historical volatility. Prior to 2023, the expected volatility was estimated based on both the Company’s volatility and the average volatility for comparable publicly traded companies over a period equal to the expected term of the related stock-based awards. The risk-free interest rate is based on the U.S. Treasury yield curve in effect at the time of grant, commensurate with the expected term. The expected dividend yield is zero as the Company has never paid dividends and has no current plans to pay any dividends on common stock. The fair value of RSUs is determined using the closing market price of the Company’s common stock on the grant date. The Company recognizes stock-based compensation expense on a straight-line basis over the award’s requisite service period, which is the vesting period for stock options and RSUs, and the offering period for purchase rights under the ESPP. The Company recognizes forfeitures as they occur.
Advertising	Advertising
Recent Accounting Pronouncements	Recent Accounting Pronouncements Recent Accounting Standards To Be Adopted In June 2022, the FASB issued ASC Update No. 2022-03, Fair Value Measurement (Topic 820): Fair Value Measurement of Equity Securities Subject to Contractual Sale Restrictions . This update clarifies the guidance in Topic 820 related to measuring the fair value of an equity security subject to contractual restrictions that prohibit the sale of an equity security, as well as introduces new disclosure requirements for these types of equity securities. The new standard will be effective for the Company on January 1, 2024. The adoption of this standard is not expected to have a material impact on the Company’s financial position or results of operations. In November 2023, the FASB issued ASC Update No. 2023-07, Segment Reporting (Topic 280): Improvement to Reportable Segment Disclosures. This update enhances reportable segment disclosure requirements by requiring public entities to provide disclosures of significant segment expenses and other segment items, as well as disclosures about a reportable segment’s profit or loss and assets that are currently required annually in interim periods. The new standard will be effective for the Company’s annual financial statements for the period beginning on January 1, 2024. The adoption of this standard is not expected to have a material impact on the Company’s disclosures. In December 2023, the FASB issued ASC Update No. 2023-09, Income Taxes (Topic 740): Improvements to Income Tax Disclosures. This update enhances income tax disclosure requirements by requiring public entities to provide additional information in its tax rate reconciliation and additional disclosures about income taxes paid. The update is effective for annual periods beginning after December 15, 2024. Early adoption is permitted for annual financial statements that have not yet been issued or made available for issuance and the amendments in this update should be applied prospectively, but entities have the option to apply it retrospectively. The Company is currently evaluating the impact that the standard will have on its Consolidated Financial Statements.

Restatement of Financial Stat_3

Restatement of Financial Statements (Tables)	12 Months Ended
	Dec. 31, 2023
Accounting Changes and Error Corrections [Abstract]
Schedule of Error Corrections and Prior Period Adjustments	QUANTERIX CORPORATION RESTATED CONSOLIDATED BALANCE SHEETS (amounts in thousands, except per share data) As of December 31, 2023 As Previously Reported Adjustments As Restated ASSETS Current assets: Cash and cash equivalents $ 174,422 $ — $ 174,422 Marketable securities 146,902 — 146,902 Accounts receivable, net of allowance for expected credit losses 25,414 — 25,414 Inventory 22,365 3,758 26,123 Prepaid expenses and other current assets 9,291 (57) 9,234 Total current assets 378,394 3,701 382,095 Restricted cash 2,604 — 2,604 Property and equipment, net 17,926 — 17,926 Intangible assets, net 6,034 — 6,034 Operating lease right-of-use assets 18,251 — 18,251 Other non-current assets 1,802 (145) 1,657 Total assets $ 425,011 $ 3,556 $ 428,567 LIABILITIES AND STOCKHOLDERS' EQUITY Current liabilities: Accounts payable $ 5,048 $ — $ 5,048 Accrued compensation and benefits 13,659 511 14,170 Accrued expenses and other current liabilities 6,041 14 6,055 Deferred revenue 9,468 — 9,468 Operating lease liabilities 4,241 — 4,241 Total current liabilities 38,457 525 38,982 Deferred revenue, net of current portion 1,227 — 1,227 Operating lease liabilities, net of current portion 37,223 — 37,223 Other non-current liabilities 1,177 — 1,177 Total liabilities 78,084 525 78,609 Commitments and contingencies (Note 16) Stockholders’ equity: Common stock, $0.001 par value per share: Authorized: 120,000 shares; Issued and outstanding: 38,014 shares at December 31, 2023 38 — 38 Additional paid-in capital 783,142 — 783,142 Accumulated other comprehensive loss (1,757) 85 (1,672) Accumulated deficit (434,496) 2,946 (431,550) Total stockholders’ equity 346,927 3,031 349,958 Total liabilities and stockholders’ equity $ 425,011 $ 3,556 $ 428,567 As of December 31, 2022 As Previously Reported Adjustments As Restated ASSETS Current assets: Cash and cash equivalents $ 338,740 $ — $ 338,740 Accounts receivable, net of allowance for expected credit losses 19,017 — 19,017 Inventory 16,786 178 16,964 Prepaid expenses and other current assets 6,860 (57) 6,803 Total current assets 381,403 121 381,524 Restricted cash 2,597 — 2,597 Property and equipment, net 20,162 (351) 19,811 Intangible assets, net 7,516 — 7,516 Operating lease right-of-use assets 21,223 — 21,223 Other non-current assets 1,298 (387) 911 Total assets $ 434,199 $ (617) $ 433,582 LIABILITIES AND STOCKHOLDERS' EQUITY Current liabilities: Accounts payable $ 3,836 $ — $ 3,836 Accrued compensation and benefits 10,658 377 11,035 Accrued expenses and other current liabilities 5,133 14 5,147 Deferred revenue 8,644 — 8,644 Operating lease liabilities 2,687 — 2,687 Total current liabilities 30,958 391 31,349 Deferred revenue, net of current portion 1,415 — 1,415 Operating lease liabilities, net of current portion 41,417 — 41,417 Other non-current liabilities 1,469 — 1,469 Total liabilities 75,259 391 75,650 Commitments and contingencies (Note 16) Stockholders’ equity: Common stock, $0.001 par value per share: Authorized: 120,000 shares; Issued and outstanding: 37,280 shares at December 31, 2022 37 — 37 Additional paid-in capital 763,688 (59) 763,629 Accumulated other comprehensive loss (2,623) 85 (2,538) Accumulated deficit (402,162) (1,034) (403,196) Total stockholders’ equity 358,940 (1,008) 357,932 Total liabilities and stockholders’ equity $ 434,199 $ (617) $ 433,582 QUANTERIX CORPORATION RESTATED CONSOLIDATED STATEMENTS OF OPERATIONS (amounts in thousands, except per share data) Year Ended December 31, 2023 As Previously Reported Adjustments As Restated Revenues: Product revenue $ 79,460 $ 210 $ 79,670 Service and other revenue 40,299 (210) 40,089 Collaboration and license revenue 1,380 — 1,380 Grant revenue 1,229 — 1,229 Total revenues 122,368 — 122,368 Costs of goods sold and services: Cost of product revenue 32,636 (3,533) 29,103 Cost of service and other revenue 19,086 (45) 19,041 Total costs of goods sold and services 51,722 (3,578) 48,144 Gross profit 70,646 3,578 74,224 Operating expenses: Research and development 24,857 1,207 26,064 Selling, general and administrative 90,241 (1,130) 89,111 Other lease costs 3,712 — 3,712 Impairment and restructuring 1,537 (209) 1,328 Total operating expenses 120,347 (132) 120,215 Loss from operations (49,701) 3,710 (45,991) Interest income 15,839 — 15,839 Other income 2,247 270 2,517 Loss before income taxes (31,615) 3,980 (27,635) Income tax expense (719) — (719) Net loss $ (32,334) $ 3,980 $ (28,354) Net loss per common share, basic and diluted $ (0.86) $ 0.11 $ (0.75) Weighted-average common shares outstanding, basic and diluted 37,594 — 37,594 Year Ended December 31, 2022 As Previously Reported Adjustments As Restated Revenues: Product revenue $ 69,808 $ — $ 69,808 Service and other revenue 34,495 — 34,495 Collaboration and license revenue 649 — 649 Grant revenue 570 — 570 Total revenues 105,522 — 105,522 Costs of goods sold and services: Cost of product revenue 40,809 2,032 42,841 Cost of service and other revenue 17,907 (589) 17,318 Total costs of goods sold and services 58,716 1,443 60,159 Gross profit 46,806 (1,443) 45,363 Operating expenses: Research and development 25,890 919 26,809 Selling, general and administrative 91,995 (144) 91,851 Other lease costs 1,278 133 1,411 Impairment and restructuring 29,347 209 29,556 Total operating expenses 148,510 1,117 149,627 Loss from operations (101,704) (2,560) (104,264) Interest income 5,131 — 5,131 Other expense (62) (215) (277) Loss before income taxes (96,635) (2,775) (99,410) Income tax expense (65) (99) (164) Net loss $ (96,700) $ (2,874) $ (99,574) Net loss per common share, basic and diluted $ (2.61) $ (0.08) $ (2.69) Weighted-average common shares outstanding, basic and diluted 36,991 — 36,991 Year Ended December 31, 2021 As Previously Reported Adjustments As Restated Revenues: Product revenue $ 81,062 $ — $ 81,062 Service and other revenue 23,629 — 23,629 Collaboration and license revenue 648 — 648 Grant revenue 5,217 — 5,217 Total revenues 110,556 — 110,556 Costs of goods sold and services: Cost of product revenue 34,149 (2,035) 32,114 Cost of service and other revenue 14,679 (1,009) 13,670 Total costs of goods sold and services 48,828 (3,044) 45,784 Gross profit 61,728 3,044 64,772 Operating expenses: Research and development 27,978 405 28,383 Selling, general and administrative 92,336 574 92,910 Total operating expenses 120,314 979 121,293 Loss from operations (58,586) 2,065 (56,521) Interest expense (403) — (403) Other income 1,265 75 1,340 Loss before income taxes (57,724) 2,140 (55,584) Income tax benefit 36 — 36 Net loss $ (57,688) $ 2,140 $ (55,548) Net loss per common share, basic and diluted $ (1.60) $ 0.06 $ (1.54) Weighted-average common shares outstanding, basic and diluted 35,997 — 35,997 QUANTERIX CORPORATION RESTATED CONSOLIDATED STATEMENTS OF COMPREHENSIVE LOSS (amounts in thousands) Year Ended December 31, 2023 As Previously Reported Adjustments As Restated Net loss $ (32,334) $ 3,980 $ (28,354) Other comprehensive income (loss), net of tax: Unrealized gains on marketable securities 325 — 325 Foreign currency translation adjustment 541 — 541 Total other comprehensive income 866 — 866 Comprehensive loss $ (31,468) $ 3,980 $ (27,488) Year Ended December 31, 2022 As Previously Reported Adjustments As Restated Net loss $ (96,700) $ (2,874) $ (99,574) Other comprehensive income (loss), net of tax: Foreign currency translation adjustment (3,064) 85 (2,979) Total other comprehensive income (loss) (3,064) 85 (2,979) Comprehensive loss $ (99,764) $ (2,789) $ (102,553) Year Ended December 31, 2021 As Previously Reported Adjustments As Restated Net loss $ (57,688) $ 2,140 $ (55,548) Other comprehensive income (loss), net of tax: Foreign currency translation adjustment (1,993) — (1,993) Total other comprehensive loss (1,993) — (1,993) Comprehensive loss $ (59,681) $ 2,140 $ (57,541) QUANTERIX CORPORATION RESTATED CONSOLIDATED STATEMENTS OF STOCKHOLDERS’ EQUITY (amounts in thousands) Year Ended December 31, 2023 Common Stock As Previously Reported Shares Amount Additional paid-in capital Accumulated other comprehensive income (loss) Accumulated Total stockholders' equity Balance at December 31, 2022 37,280 $ 37 $ 763,688 $ (2,623) $ (402,162) $ 358,940 Issuance of common stock under stock plans, including tax effects 734 1 2,690 — — 2,691 Stock-based compensation expense — — 16,764 — — 16,764 Unrealized gains on marketable securities, net of tax — — — 325 — 325 Foreign currency translation adjustment — — — 541 — 541 Net loss — — — — (32,334) (32,334) Balance at December 31, 2023 38,014 $ 38 $ 783,142 $ (1,757) $ (434,496) $ 346,927 Adjustments Balance at December 31, 2022 — $ — $ (59) $ 85 $ (1,034) $ (1,008) Issuance of common stock under stock plans, including tax effects — — — — — — Stock-based compensation expense — — 59 — — 59 Unrealized gains on marketable securities, net of tax — — — — — — Foreign currency translation adjustment — — — — — — Net loss — — — — 3,980 3,980 Balance at December 31, 2023 — $ — $ — $ 85 $ 2,946 $ 3,031 As Restated Balance at December 31, 2022 37,280 $ 37 $ 763,629 $ (2,538) $ (403,196) $ 357,932 Issuance of common stock under stock plans, including tax effects 734 1 2,690 — — 2,691 Stock-based compensation expense — — 16,823 — — 16,823 Unrealized gains on marketable securities, net of tax — — — 325 — 325 Foreign currency translation adjustment — — — 541 — 541 Net loss — — — — (28,354) (28,354) Balance at December 31, 2023 38,014 $ 38 $ 783,142 $ (1,672) $ (431,550) $ 349,958 Year Ended December 31, 2022 Common Stock As Previously Reported Shares Amount Additional paid-in capital Accumulated other comprehensive income (loss) Accumulated Total stockholders' equity Balance at December 31, 2021 36,768 $ 37 $ 745,936 $ 441 $ (305,462) $ 440,952 Issuance of common stock under stock plans, including tax effects 512 — 2,310 — — 2,310 Stock-based compensation expense — — 15,442 — — 15,442 Foreign currency translation adjustment — — — (3,064) — (3,064) Net loss — — — — (96,700) (96,700) Balance at December 31, 2022 37,280 $ 37 $ 763,688 $ (2,623) $ (402,162) $ 358,940 Adjustments Balance at December 31, 2021 — $ — $ — $ — $ 1,840 $ 1,840 Issuance of common stock under stock plans, including tax effects — — — — — — Stock-based compensation expense — — (59) — — (59) Foreign currency translation adjustment — — — 85 — 85 Net loss — — — — (2,874) (2,874) Balance at December 31, 2022 — $ — $ (59) $ 85 $ (1,034) $ (1,008) As Restated Balance at December 31, 2021 36,768 $ 37 $ 745,936 $ 441 $ (303,622) $ 442,792 Issuance of common stock under stock plans, including tax effects 512 — 2,310 — — 2,310 Stock-based compensation expense — — 15,383 — — 15,383 Foreign currency translation adjustment — — — (2,979) — (2,979) Net loss — — — — (99,574) (99,574) Balance at December 31, 2022 37,280 $ 37 $ 763,629 $ (2,538) $ (403,196) $ 357,932 Year Ended December 31, 2021 Common Stock As Previously Reported Shares Amount Additional paid-in capital Accumulated other comprehensive income (loss) Accumulated Total stockholders' equity Balance at December 31, 2020 31,797 $ 32 $ 451,433 $ 2,434 $ (247,774) $ 206,125 Issuance of common stock under stock plans, including tax effects 864 1 8,814 — — 8,815 Sale of common stock in underwritten public offering, net 4,107 4 269,714 — — 269,718 Stock-based compensation expense — — 15,975 — — 15,975 Foreign currency translation adjustment — — — (1,993) — (1,993) Net loss — — — — (57,688) (57,688) Balance at December 31, 2021 36,768 $ 37 $ 745,936 $ 441 $ (305,462) $ 440,952 Adjustments Balance at December 31, 2020 — $ — $ — $ — $ (300) $ (300) Issuance of common stock under stock plans, including tax effects — — — — — — Sale of common stock in underwritten public offering, net — — — — — — Stock-based compensation expense — — — — — — Foreign currency translation adjustment — — — — — — Net loss — — — — 2,140 2,140 Balance at December 31, 2021 — $ — $ — $ — $ 1,840 $ 1,840 As Restated Balance at December 31, 2020 31,797 $ 32 $ 451,433 $ 2,434 $ (248,074) $ 205,825 Issuance of common stock under stock plans, including tax effects 864 1 8,814 — — 8,815 Sale of common stock in underwritten public offering, net 4,107 4 269,714 — — 269,718 Stock-based compensation expense — — 15,975 — — 15,975 Foreign currency translation adjustment — — — (1,993) — (1,993) Net loss — — — — (55,548) (55,548) Balance at December 31, 2021 36,768 $ 37 $ 745,936 $ 441 $ (303,622) $ 442,792 QUANTERIX CORPORATION RESTATED CONSOLIDATED STATEMENTS OF CASH FLOWS (amounts in thousands) Year Ended December 31, 2023 As Previously Reported Adjustments As Restated Cash flows from operating activities: Net loss $ (32,334) $ 3,980 $ (28,354) Adjustments to reconcile net loss to net cash used in operating activities: Depreciation and amortization expense 6,364 (89) 6,275 Credit losses on accounts receivable 336 — 336 Accretion of marketable securities (1,964) — (1,964) Operating lease right-of-use asset amortization 2,015 — 2,015 Stock-based compensation expense 16,764 59 16,823 Impairment 1,570 (209) 1,361 Other operating activity 150 (300) (150) Changes in assets and liabilities: Accounts receivable (6,695) — (6,695) Inventory (5,364) (3,580) (8,944) Prepaid expenses and other current assets (2,371) — (2,371) Other non-current assets (775) 58 (717) Accounts payable 1,189 — 1,189 Accrued compensation and benefits, accrued expenses, and other current liabilities 4,276 134 4,410 Deferred revenue 635 — 635 Operating lease liabilities (2,645) — (2,645) Other non-current liabilities (53) — (53) Net cash used in operating activities (18,902) 53 (18,849) Cash flows from investing activities: Purchases of marketable debt securities (175,613) — (175,613) Proceeds from marketable debt securities 31,000 — 31,000 Purchases of property and equipment (3,788) (53) (3,841) Net cash used in investing activities (148,401) (53) (148,454) Cash flows from financing activities: Proceeds from common stock issued under stock plans 2,889 — 2,889 Payments for employee taxes withheld on stock-based compensation awards (198) — (198) Net cash provided by financing activities 2,691 — 2,691 Net increase (decrease) in cash, cash equivalents, and restricted cash (164,612) — (164,612) Effect of exchange rate changes on cash, cash equivalents, and restricted cash 301 — 301 Cash, cash equivalents, and restricted cash at beginning of period 341,337 — 341,337 Cash, cash equivalents, and restricted cash at end of period $ 177,026 $ — $ 177,026 Supplemental disclosure of cash flow information: Cash paid for taxes $ 808 $ — $ 808 Purchases of property and equipment in accounts payable and accruals $ 419 $ — $ 419 Shares received as consideration under product sales agreement (Note 4, 7) $ 775 $ — $ 775 Year Ended December 31, 2022 As Previously Reported Adjustments As Restated Cash flows from operating activities: Net loss $ (96,700) $ (2,874) $ (99,574) Adjustments to reconcile net loss to net cash used in operating activities: Depreciation and amortization expense 5,349 60 5,409 Credit losses on accounts receivable (301) — (301) Operating lease right-of-use asset amortization 715 — 715 Stock-based compensation expense 15,442 (59) 15,383 Impairment 25,592 209 25,801 Other operating activity (439) 99 (340) Changes in assets and liabilities: Accounts receivable 5,156 — 5,156 Inventory 5,386 1,857 7,243 Prepaid expenses and other current assets (568) (170) (738) Other non-current assets (909) 87 (822) Accounts payable (5,362) 75 (5,287) Accrued compensation and benefits, accrued expenses, and other current liabilities (3,976) 377 (3,599) Deferred revenue 2,599 — 2,599 Operating lease liabilities (266) — (266) Other non-current liabilities 10 — 10 Net cash used in operating activities (48,272) (339) (48,611) Cash flows from investing activities: Purchases of property and equipment (11,726) 112 (11,614) Proceeds from RADx grant on assets purchased 520 — 520 Net cash used in investing activities (11,206) 112 (11,094) Cash flows from financing activities: Proceeds from common stock issued under stock plans 2,311 — 2,311 Net cash provided by financing activities 2,311 — 2,311 Net increase (decrease) in cash, cash equivalents, and restricted cash (57,167) (227) (57,394) Effect of exchange rate changes on cash, cash equivalents, and restricted cash (538) — (538) Cash, cash equivalents, and restricted cash at beginning of period 399,042 227 399,269 Cash, cash equivalents, and restricted cash at end of period $ 341,337 $ — $ 341,337 Supplemental disclosure of cash flow information: Cash paid for taxes $ 684 $ — $ 684 Purchases of property and equipment in accounts payable and accruals $ 152 $ — $ 152 Operating lease right-of-use assets obtained in exchange for lease liabilities $ 22,494 $ — $ 22,494 Year Ended December 31, 2021 As Previously Reported Adjustments As Restated Cash flows from operating activities: Net loss $ (57,688) $ 2,140 $ (55,548) Adjustments to reconcile net loss to net cash used in operating activities: Depreciation and amortization expense 4,851 29 4,880 Credit losses on accounts receivable 213 — 213 Operating lease right-of-use asset amortization 499 — 499 Stock-based compensation expense 15,975 — 15,975 Other operating activity 692 — 692 Changes in assets and liabilities: Accounts receivable (6,853) — (6,853) Inventory (8,090) (2,035) (10,125) Prepaid expenses and other current assets (393) 227 (166) Other non-current assets (2) — (2) Accounts payable 2,414 (75) 2,339 Accrued compensation and benefits, accrued expenses, and other current liabilities 606 — 606 Deferred revenue 1,462 — 1,462 Operating lease liabilities (1,230) — (1,230) Other non-current liabilities (363) — (363) Net cash used in operating activities (47,907) 286 (47,621) Cash flows from investing activities: Purchases of marketable debt securities — — — Proceeds from marketable debt securities — — — Purchases of property and equipment (13,616) (59) (13,675) Proceeds from RADx grant on assets purchased 7,278 — 7,278 Net cash used in investing activities (6,338) (59) (6,397) Cash flows from financing activities: Proceeds from common stock issued under stock plans 8,815 — 8,815 Sale of common stock in underwritten public offering, net 269,718 — 269,718 Payments on notes payable (7,738) — (7,738) Net cash provided by financing activities 270,795 — 270,795 Net increase (decrease) in cash, cash equivalents, and restricted cash 216,550 227 216,777 Effect of exchange rate changes on cash, cash equivalents, and restricted cash (92) — (92) Cash, cash equivalents, and restricted cash at beginning of period 182,584 — 182,584 Cash, cash equivalents, and restricted cash at end of period $ 399,042 $ 227 $ 399,269 Supplemental disclosure of cash flow information: Cash paid for interest $ 389 $ — $ 389 Purchases of property and equipment in accounts payable and accruals $ 229 $ — $ 229

Significant Accounting Polici_3

Significant Accounting Policies (Tables)	12 Months Ended
	Dec. 31, 2023
Accounting Policies [Abstract]
Schedule of cash and cash equivalents	As of December 31, 2023 2022 Cash $ 12,162 $ 32,643 Money market funds 155,367 306,097 Marketable securities 6,893 — Total cash and cash equivalents $ 174,422 $ 338,740
Schedule of restricted cash and cash equivalents	The following table summarizes the period ending cash and cash equivalents as presented on the Consolidated Balance Sheets and the total cash, cash equivalents, and restricted cash as presented on the Consolidated Statements of Cash Flows (in thousands): As of December 31, 2023 2022 Cash and cash equivalents $ 174,422 $ 338,740 Restricted cash 2,604 2,597 Cash, cash equivalents, and restricted cash $ 177,026 $ 341,337
Schedule of Property, Plant and Equipment, Useful Life	Estimated Useful Life Laboratory and manufacturing equipment 5 years Office furniture and equipment 7 years Computers and software 3 years Leasehold improvements Shorter of asset's life or remaining lease term

Revenue and Related Matters (Ta

Revenue and Related Matters (Tables)	12 Months Ended
	Dec. 31, 2023
Revenue from Contract with Customer [Abstract]
Schedule of disaggregated revenue	Year Ended December 31, 2023 North America EMEA Asia Pacific Total (As Restated) (As Restated) (As Restated) (As Restated) Product revenue: Instruments $ 6,374 $ 4,384 $ 4,947 $ 15,705 Consumable and other products 35,122 21,216 7,627 63,965 Total $ 41,496 $ 25,600 $ 12,574 $ 79,670 Service revenue: Service-type warranties $ 6,265 $ 3,001 $ 613 $ 9,879 Research services 24,706 2,000 1,122 27,828 Other services 1,436 951 (5) 2,382 Total $ 32,407 $ 5,952 $ 1,730 $ 40,089 Collaboration and license revenue: Total $ 1,380 $ — $ — $ 1,380 Year Ended December 31, 2022 North America EMEA Asia Pacific Total Product revenue: Instruments $ 9,254 $ 8,362 $ 7,388 $ 25,004 Consumable and other products 25,894 14,514 4,396 44,804 Total $ 35,148 $ 22,876 $ 11,784 $ 69,808 Service and other revenues: Service-type warranties $ 5,581 $ 2,779 $ 480 $ 8,840 Research services 22,493 1,013 147 23,653 Other services 1,144 722 136 2,002 Total $ 29,218 $ 4,514 $ 763 $ 34,495 Collaboration and license revenue: Total $ 274 $ 323 $ 52 $ 649 Year Ended December 31, 2021 North America EMEA Asia Pacific Total Product revenue: Instruments $ 12,138 $ 8,178 $ 5,657 $ 25,973 Consumable and other products 34,997 16,122 3,970 55,089 Total $ 47,135 $ 24,300 $ 9,627 $ 81,062 Service and other revenue: Service-type warranties $ 4,334 $ 2,039 $ 255 $ 6,628 Research services 12,101 2,600 124 14,825 Other services 1,372 695 109 2,176 Total $ 17,807 $ 5,334 $ 488 $ 23,629 Collaboration and license revenue: Total $ 360 $ 288 $ — $ 648
Schedule of costs to obtain a contract	Changes in costs to obtain a contract were as follows (in thousands): 2023 2022 2021 Balance at December 31 of prior year $ 377 $ 440 $ 248 Capitalization of costs to obtain a contract 528 1,387 905 Recognition of costs to obtain a contract (617) (1,450) (713) Balance at December 31 $ 288 $ 377 $ 440

Allowance for Credit Losses (Ta

Allowance for Credit Losses (Tables)	12 Months Ended
	Dec. 31, 2023
Allowance for Credit Loss [Abstract]
Schedule of the allowance for credit losses	The change in the allowance for credit losses on accounts receivable is summarized as follows (in thousands): 2023 2022 2021 Balance at December 31 of prior year $ 118 $ 419 $ 370 Provision for expected credit losses 729 752 213 Write-offs and recoveries collected (393) (1,053) (164) Balance at December 31 $ 454 $ 118 $ 419

Marketable Securities (Tables)

Marketable Securities (Tables)	12 Months Ended
	Dec. 31, 2023
Investments, Debt and Equity Securities [Abstract]
Debt Securities, Available-for-Sale	The amortized cost, gross unrealized gains, gross unrealized losses, and fair value of the Company’s marketable securities by major security type were as follows (in thousands): As of December 31, 2023 Amortized cost Unrealized Gains Unrealized Losses Fair Value Commercial paper $ 53,482 $ 23 $ (12) $ 53,493 U.S. Treasuries 4,896 1 — 4,897 U.S. Government agency bonds 28,366 39 (7) 28,398 Corporate bonds 66,726 289 (8) 67,007 Total marketable securities $ 153,470 $ 352 $ (27) $ 153,795 Marketable securities are reported in the following Consolidated Balance Sheets captions: Cash and cash equivalents $ 6,893 Marketable securities 146,902 Total marketable securities $ 153,795
Unrealized Gain (Loss) on Investments	The following table shows the fair value and gross unrealized losses of the Company’s available-for-sale securities with unrealized losses that are not deemed to be other-than-temporary, aggregated by major security type and length of time that the individual securities have been in a continuous unrealized loss position (in thousands): Less Than 12 Months As of December 31, 2023 Fair Value Unrealized Losses Commercial paper $ 32,137 $ (12) U.S. Government agency bonds 15,861 (7) Corporate bonds 8,367 (8) Total $ 56,365 $ (27)
Investments Classified by Contractual Maturity Date	The following table summarizes the contractual maturities of the Company’s marketable securities (in thousands): As of December 31, 2023 Amortized cost Fair Value Due within one year $ 95,188 $ 95,232 Due in one to two years 58,282 58,563 Total $ 153,470 $ 153,795

Fair value of Financial Instr_2

Fair value of Financial Instruments (As Restated) (Tables)	12 Months Ended
	Dec. 31, 2023
Fair Value Disclosures [Abstract]
Schedule of fair value measurements	The following tables present the Company’s fair value hierarchy for its financial assets that are measured at fair value on a recurring basis (in thousands): As of December 31, 2023 Total Level 1 Level 2 Level 3 Financial assets: Cash equivalents: (1) Money market funds $ 155,367 $ 155,367 $ — $ — Commercial paper 1,996 — 1,996 — U.S. Treasuries 4,897 — 4,897 — Total cash equivalents 162,260 155,367 6,893 — Marketable securities: (2) Commercial paper 51,498 — 51,498 — U.S. Treasuries — — — — U.S. Government agency bonds 28,398 — 28,398 — Corporate bonds 67,006 — 67,006 — Total marketable securities 146,902 — 146,902 — Total financial assets $ 309,162 $ 155,367 $ 153,795 $ — As of December 31, 2022 Total Level 1 Level 2 Level 3 Financial assets: Money market funds (1) $ 306,097 $ 306,097 $ — $ — Total financial assets $ 306,097 $ 306,097 $ — $ — (1) Included in cash and cash equivalents on the Consolidated Balance Sheets. (2) Marketable securities are initially valued at their purchase price and subsequently fair valued at the end of each reporting period utilizing third party pricing services or other observable data. The pricing services utilize industry standard valuation methods, including both income and market-based approaches and observable market inputs to determine the fair value. These observable market inputs include reportable trades, benchmark yields, credit spreads, broker/dealer quotes, bids, offers, current spot rates, and other industry and economic events.

Inventory (Tables)

Inventory (Tables)	12 Months Ended
	Dec. 31, 2023
Inventory Disclosure [Abstract]
Summary of inventory	Inventory, net of inventory reserves, consisted of the following (in thousands): As of December 31, 2023 2022 (As Restated) (As Restated) Raw materials $ 5,114 $ 5,509 Work in process 5,439 3,415 Finished goods 15,570 8,040 Total inventory $ 26,123 $ 16,964

Property and Equipment (Tables)

Property and Equipment (Tables)	12 Months Ended
	Dec. 31, 2023
Property, Plant and Equipment [Abstract]
Schedule of Property and equipment	Property and equipment consisted of the following (in thousands): As of December 31, 2023 2022 (As Restated) Laboratory and manufacturing equipment $ 13,141 $ 11,753 Office furniture and equipment 1,905 1,798 Computers and software 3,927 3,581 Leasehold improvements 13,074 13,688 Total cost $ 32,047 $ 30,820 Less: accumulated depreciation (14,121) (11,009) Property and equipment, net $ 17,926 $ 19,811

Accrued Expenses and Other Cu_2

Accrued Expenses and Other Current Liabilities (Tables)	12 Months Ended
	Dec. 31, 2023
Accrued Expenses and Other Current Liabilities
Summary of Accrued expenses and other current liabilities	Accrued expenses and other current liabilities consisted of the following (in thousands): As of December 31, 2023 2022 (As Restated) (As Restated) Accrued professional services $ 1,596 $ 1,409 Accrued royalties 1,689 815 Accrued tax liabilities 822 186 Other accrued expenses 1,948 2,737 Total accrued expenses and other current liabilities $ 6,055 $ 5,147

Stock-Based Compensation (Table

Stock-Based Compensation (Tables)	12 Months Ended
	Dec. 31, 2023
Share-Based Payment Arrangement [Abstract]
Summary of stock option activity	Stock option activity for the year ended December 31, 2023 is presented below (in thousands, except per share and contractual life amounts): Number of options Weighted-average Weighted-average remaining contractual Aggregate Outstanding at December 31, 2022 2,188 $ 20.69 8.1 $ 4,273 Granted 1,187 16.23 Exercised (161) 11.77 Forfeited/expired (440) 18.65 Outstanding at December 31, 2023 2,774 $ 19.62 7.9 $ 26,941 Exercisable at December 31, 2023 1,117 $ 22.81 6.4 $ 9,176 Vested and expected to vest at December 31, 2023 2,774 $ 19.62 7.9 $ 26,941
Summary of restricted stock units activity	RSU activity for the year ended December 31, 2023 is presented below (in thousands, except per share amounts): Number of shares Weighted-average grant Unvested as of December 31, 2022 1,188 $ 21.18 Granted 868 15.90 Vested (453) 23.00 Forfeited (275) 17.50 Unvested as of December 31, 2023 1,328 $ 17.87 Expected to convert at December 31, 2023 1,328 $ 17.87
Summary of share-based compensation expense for all stock awards	Stock-based compensation expense was recorded in the following categories on the Consolidated Statements of Operations (in thousands): Year Ended December 31, 2023 2022 2021 (As Restated) (As Restated) Cost of product revenue $ 839 $ 596 $ 471 Cost of service and other revenue 1,124 819 403 Research and development 1,713 1,629 1,807 Selling, general and administrative 13,147 12,339 13,294 Total stock-based compensation $ 16,823 $ 15,383 $ 15,975
Summary of fair value of stock options granted to employees and non-employees is estimated on the grant date using the Black-Scholes option-pricing model	The fair value of the Company’s stock options granted and purchase rights to the ESPP were estimated using the Black-Scholes valuation model with the following assumptions: Year Ended December 31, 2023 2022 2021 Stock Options: Risk-free interest rate 3.5% - 4.7% 1.4% - 4.1% 0.4% - 1.3% Expected dividend yield None None None Expected term (in years) 5.0 - 5.2 5.0 - 5.8 6.0 Expected volatility 71.1% - 83.1% 55.0% - 70.8% 49.2% - 55.6% Weighted-average grant date fair value per share $ 10.63 $ 9.88 $ 29.96 Employee Stock Purchase Plan: Risk-free interest rate 5.2% - 5.5% 0.7% - 3.9% 0.1% Expected dividend yield None None None Expected term (in years) 0.5 0.5 0.5 Expected volatility 72.8% - 82.5% 51.9% - 117.3% 46.3% - 59.8% Weighted-average grant date fair value per share $ 3.19 $ 3.53 $ 8.06

Net Loss Per Share (Tables)

Net Loss Per Share (Tables)	12 Months Ended
	Dec. 31, 2023
Earnings Per Share [Abstract]
Schedule of basic and diluted shares	The following table presents the computation of basic and diluted net loss per share (in thousands, except per share data): Year Ended December 31, 2023 2022 2021 (As Restated) (As Restated) (As Restated) Numerator: Net loss $ (28,354) $ (99,574) $ (55,548) Denominator: Weighted average common shares outstanding, basic and diluted 37,594 36,991 35,997 Net loss per share, basic and diluted $ (0.75) $ (2.69) $ (1.54)
Schedule of common share equivalents have been excluded from the calculation of diluted net loss per share	As the Company was in a net loss position, the following common share equivalents (calculated on a weighted average basis) were excluded from the calculation of diluted net loss per share (in thousands): Year Ended December 31, 2023 2022 2021 Stock options 2,783 446 2,305 Common stock and RSUs 1,512 702 531 Estimated ESPP purchases 23 52 8 Total dilutive shares 4,318 1,200 2,844

Income Taxes (As Restated) (Tab

Income Taxes (As Restated) (Tables)	12 Months Ended
	Dec. 31, 2023
Income Tax Disclosure [Abstract]
Schedule of components of loss before income taxes	The following table presents the components of loss before income taxes (in thousands): Year Ended December 31, 2023 2022 2021 United States $ (30,355) $ (92,360) $ (54,414) Foreign 2,720 (7,050) (1,170) Total loss before income taxes $ (27,635) $ (99,410) $ (55,584)
Schedule of provision for (benefit from) income taxes	The following table summarizes income tax (expense) benefit (in thousands): Year Ended December 31, 2023 2022 2021 Current: United States Federal $ — $ — $ — State (161) (87) (33) Foreign (850) (372) (340) Total current income tax provision (1,011) (459) (373) Deferred United States Federal — 10 5 State — 13 (6) Foreign 292 272 410 Total deferred income tax benefit 292 295 409 Total income tax (expense) benefit $ (719) $ (164) $ 36
Schedule of reconciliation of the federal statutory income tax rate to the effective tax rate	Year Ended December 31, 2023 2022 2021 Federal statutory income tax rate 21.0 % 21.0 % 21.0 % Non-deductible executive compensation (2.6) % — % — % State taxes, net of federal benefit (0.3 %) 2.8 % 6.6 % Tax credits 4.9 % 1.3 % 2.1 % Stock-based compensation (1.6) % (2.4) % 7.7 % Permanent items (0.6) % (0.3) % (1.9) % Deferred tax rate change — % — % 0.2 % Change in valuation allowance (24.3) % (20.8) % (35.2) % Impairment of goodwill — % (1.6) % — % Other 0.9 % (0.2 %) (0.4) % Effective income tax rate (2.6) % (0.2) % 0.1 %
Schedule of deferred tax assets and liabilities	Deferred tax assets and liabilities reflect the net tax effects of temporary differences between the carrying amount of assets and liabilities for financial reporting and the amounts used for income tax purposes. Significant components of the Company’s deferred tax assets and liabilities were as follows (in thousands): As of December 31, 2023 2022 Deferred tax assets: Net operating loss carryforwards $ 77,731 $ 77,689 Tax credits 8,645 7,303 Deferred revenue 2,499 2,385 Amortization 727 847 Stock-based compensation 3,186 2,773 Inventory 1,391 1,507 Capitalized R&D Costs 9,353 5,743 Lease liability 9,948 10,856 Other deferred tax assets 2,357 1,536 Total deferred tax assets 115,837 110,639 Less: valuation allowances (110,082) (103,372) Net deferred tax assets 5,755 7,267 Deferred tax liabilities: Right-of-Use Assets (4,372) (5,219) Depreciation (1,307) (1,756) Amortization acquired intangibles (1,253) (1,570) Other deferred tax liabilities — (189) Net deferred tax (liability) asset (1,177) (1,467)
Summary of Valuation Allowance	The Company’s change in its valuation allowance account related to deferred tax assets was as follows (in thousands): 2023 2022 Balance at December 31 of prior year $ 103,372 $ 82,654 Change in valuation allowance 6,710 20,718 Balance at December 31 $ 110,082 $ 103,372

Goodwill and Intangible Assets

Goodwill and Intangible Assets (Tables)	12 Months Ended
	Dec. 31, 2023
Goodwill and Intangible Assets Disclosure [Abstract]
Schedule of future estimated amortization expense of acquired intangible assets	Future estimated amortization expense is as follows (amounts in thousands): As of December 31, 2023 2024 $ 1,631 2025 1,493 2026 1,465 2027 1,443 2028 2 Thereafter — Total amortization expense $ 6,034
Summary of intangible assets	Acquired intangible assets consisted of the following (in thousands, except useful life and weighted average life amounts): As of December 31, 2023 Estimated Gross Carrying Accumulated Cumulative Net Carrying Weighted Average Know-how 8.5 $ 13,000 $ (6,326) $ (1,050) $ 5,624 4.0 Developed technology 7 1,650 (1,581) — 69 1.1 Customer relationships 8.5 - 10 1,360 (1,067) (9) 284 4.1 Non-compete agreements 5.5 340 (256) (27) 57 1.0 Trade names 3 50 (50) — — — Total $ 16,400 $ (9,280) $ (1,086) $ 6,034 As of December 31, 2022 Estimated Gross Carrying Accumulated Cumulative Net Carrying Weighted Average Know-how 8.5 $ 13,000 $ (4,763) $ (1,433) $ 6,804 5.0 Developed technology 7 1,650 (1,458) — 192 2.1 Customer relationships 8.5 - 10 1,360 (938) (12) 410 5.1 Non-compete agreements 5.5 340 (193) (37) 110 2.0 Trade names 3 50 (50) — — — Total $ 16,400 $ (7,402) $ (1,482) $ 7,516

Leases (Tables)

Leases (Tables)	12 Months Ended
	Dec. 31, 2023
Lessee Disclosure [Abstract]
Schedule of components of the lease costs	The components of the lease costs and supplemental cash flow information relating to the Company's leases were as follows (in thousands): Year Ended December 31, 2023 2022 2021 Operating lease cost $ 5,209 $ 5,488 $ 2,726 Short-term and variable lease cost 3,996 3,417 2,699 Total lease cost $ 9,205 $ 8,905 $ 5,425
Schedule of supplemental cash flow information	Year Ended December 31, 2023 2022 Cash paid for amounts included in the measurement of operating lease liabilities $ 8,935 $ 6,539 Operating ROU assets obtained in exchange for lease obligations $ — $ 22,494 Weighted average remaining lease term - operating leases (years) 6.8 7.8 Weighted average discount rate - operating leases 7.86 % 7.83 %
Schedule of undiscounted future lease payments for non-cancelable operating leases	The undiscounted future lease payments for non-cancelable operating leases were as follows (in thousands): Maturity of lease liabilities As of December 31, 2023 2024 $ 7,094 2025 7,254 2026 7,408 2027 7,641 2028 7,880 Thereafter 15,741 Total lease payments $ 53,018 Less: imputed interest (11,554) Total operating lease liabilities $ 41,464
Schedule of balance sheet information	Operating lease balances presented on the Consolidated Balance Sheets were as follows (in thousands): As of December 31, 2023 Operating lease ROU assets $ 18,251 Operating lease liabilities $ 4,241 Operating lease liabilities, net of current portion 37,223 Total operating lease liabilities $ 41,464

Commitments and contingencies (

Commitments and contingencies (Tables)	12 Months Ended
	Dec. 31, 2023
Commitments and Contingencies Disclosure [Abstract]
Schedule of future minimum lease payments	The undiscounted future lease payments for non-cancelable operating leases were as follows (in thousands): Maturity of lease liabilities As of December 31, 2023 2024 $ 7,094 2025 7,254 2026 7,408 2027 7,641 2028 7,880 Thereafter 15,741 Total lease payments $ 53,018 Less: imputed interest (11,554) Total operating lease liabilities $ 41,464

Restructuring (Tables)

Restructuring (Tables)	12 Months Ended
	Dec. 31, 2023
Restructuring and Related Activities [Abstract]
Schedule of rollforward of the restructuring reserve and provision activity	The following table presents the restructuring reserve and provision activity for the year ended December 31, 2023 (in thousands): Severance and Employee Benefit Costs Balance at December 31, 2022 $ 328 Accrual adjustments (33) Cash payments (16) Foreign currency translation 8 Balance at December 31, 2023 $ 287

Quarterly Financial Data (Una_2

Quarterly Financial Data (Unaudited and Restated) (Tables)	12 Months Ended
	Dec. 31, 2023
Quarterly Financial Information Disclosure [Abstract]
Summary Of Quarterly Financial Information	QUANTERIX CORPORATION RESTATED CONSOLIDATED BALANCE SHEETS (amounts in thousands, except per share data) As of March 31, 2023 As Previously Reported Adjustments As Restated ASSETS Current assets: Cash and cash equivalents $ 329,354 $ — $ 329,354 Accounts receivable, net of allowance for expected credit losses 22,546 — 22,546 Inventory 17,070 (441) 16,629 Prepaid expenses and other current assets 7,002 (57) 6,945 Total current assets 375,972 (498) 375,474 Restricted cash 2,920 (322) 2,598 Property and equipment, net 19,056 (366) 18,690 Intangible assets, net 7,129 — 7,129 Right-of-use assets 20,891 — 20,891 Other non-current assets 1,345 (407) 938 Total assets $ 427,313 $ (1,593) $ 425,720 LIABILITIES AND STOCKHOLDERS' EQUITY Current liabilities: Accounts payable $ 2,585 $ — $ 2,585 Accrued compensation and benefits 4,880 531 5,411 Other accrued expenses 4,624 14 4,638 Deferred revenue 10,682 — 10,682 Short-term lease liabilities 3,875 — 3,875 Other current liabilities 291 — 291 Total current liabilities 26,937 545 27,482 Deferred revenue, net of current portion 1,419 — 1,419 Long-term lease liabilities 40,409 — 40,409 Other non-current liabilities 1,216 — 1,216 Total liabilities 69,981 545 70,526 Commitments and contingencies (Note 16) Stockholders’ equity: Common stock, $0.001 par value per share: Authorized shares: 120,000 shares; Issued and outstanding: 37,424 shares at March 31, 2023. 37 — 37 Additional paid-in capital 768,141 (18) 768,123 Accumulated other comprehensive (loss) income (2,581) 85 (2,496) Accumulated deficit (408,265) (2,205) (410,470) Total stockholders’ equity 357,332 (2,138) 355,194 Total liabilities and stockholders’ equity $ 427,313 $ (1,593) $ 425,720 As of June 30, 2023 As Previously Reported Adjustments As Restated ASSETS Current assets: Cash and cash equivalents $ 329,525 $ — $ 329,525 Accounts receivable, net of allowance for expected credit losses 24,423 (54) 24,369 Inventory 18,156 177 18,333 Prepaid expenses and other current assets 6,954 (57) 6,897 Total current assets 379,058 66 379,124 Restricted cash 2,686 (85) 2,601 Property and equipment, net 18,328 (280) 18,048 Intangible assets, net 6,476 — 6,476 Operating lease right-of-use assets 20,380 — 20,380 Other non-current assets 2,282 (645) 1,637 Total assets $ 429,210 $ (944) $ 428,266 LIABILITIES AND STOCKHOLDERS' EQUITY Current liabilities: Accounts payable $ 3,238 $ — $ 3,238 Accrued compensation and benefits 7,525 650 8,175 Accrued expenses and other current liabilities 6,777 14 6,791 Deferred revenue 10,421 — 10,421 Operating lease liabilities 3,986 — 3,986 Total current liabilities 31,947 664 32,611 Deferred revenue, net of current portion 1,304 — 1,304 Operating lease liabilities, net of current portion 39,378 — 39,378 Other non-current liabilities 1,225 — 1,225 Total liabilities 73,854 664 74,518 Commitments and contingencies (Note 16) Stockholders’ equity: Common stock, $0.001 par value per share: Authorized shares: 120,000,000 shares; Issued and outstanding: 37,566 shares at June 30, 2023. 37 — 37 Additional paid-in capital 772,473 (1) 772,472 Accumulated other comprehensive loss (2,825) 85 (2,740) Accumulated deficit (414,329) (1,692) (416,021) Total stockholders’ equity 355,356 (1,608) 353,748 Total liabilities and stockholders’ equity $ 429,210 $ (944) $ 428,266 As of September 30, 2023 As Previously Reported Adjustments As Restated ASSETS Current assets: Cash and cash equivalents $ 201,261 $ — $ 201,261 Marketable securities 126,449 — 126,449 Accounts receivable, net of allowance for expected credit losses 24,083 — 24,083 Inventory 19,945 912 20,857 Prepaid expenses and other current assets 9,273 (57) 9,216 Total current assets 381,011 855 381,866 Restricted cash 2,647 (45) 2,602 Property and equipment, net 17,517 (230) 17,287 Intangible assets, net 6,003 — 6,003 Operating lease right-of-use assets 19,860 — 19,860 Other non-current assets 2,004 (401) 1,603 Total assets $ 429,042 $ 179 $ 429,221 LIABILITIES AND STOCKHOLDERS' EQUITY Current liabilities: Accounts payable $ 4,786 $ — $ 4,786 Accrued compensation and benefits 9,775 730 10,505 Accrued expenses and other current liabilities 6,672 14 6,686 Deferred revenue 9,827 — 9,827 Operating lease liabilities 4,093 — 4,093 Total current liabilities 35,153 744 35,897 Deferred revenue, net of current portion 1,126 — 1,126 Operating lease liabilities, net of current portion 38,306 — 38,306 Other non-current liabilities 1,105 — 1,105 Total liabilities 75,690 744 76,434 Commitments and contingencies (Note 16) Stockholders’ equity: Common stock, $0.001 par value per share: Authorized shares: 120,000,000 shares; Issued and outstanding: 37,839 shares at September 30, 2023. 38 — 38 Additional paid-in capital 778,615 — 778,615 Accumulated other comprehensive loss (3,214) 85 (3,129) Accumulated deficit (422,087) (650) (422,737) Total stockholders’ equity 353,352 (565) 352,787 Total liabilities and stockholders’ equity $ 429,042 $ 179 $ 429,221 Three Months Ended March 31, 2023 As Previously Reported Adjustments As Restated Revenues: Product revenue $ 19,287 $ 73 $ 19,360 Service and other revenue 8,579 (73) 8,506 Collaboration revenue 368 — 368 Grant revenue 222 — 222 Total revenue 28,456 — 28,456 Costs of goods sold: Cost of product revenue 7,033 846 7,879 Cost of service and other revenue 4,497 87 4,584 Total costs of goods sold and services 11,530 933 12,463 Gross profit 16,926 (933) 15,993 Operating expenses: Research and development 4,720 267 4,987 Selling, general and administrative 20,883 (66) 20,817 Other lease costs 776 24 800 Impairment and restructuring (33) — (33) Total operating expenses 26,346 225 26,571 Loss from operations (9,420) (1,158) (10,578) Interest income 3,449 — 3,449 Other income (expense), net 8 (13) (5) Loss before income taxes (5,963) (1,171) (7,134) Income tax expense (140) — (140) Net loss $ (6,103) $ (1,171) $ (7,274) Net loss per common share, basic and diluted $ (0.16) $ (0.03) $ (0.19) Weighted-average common shares outstanding, basic and diluted 37,327 — 37,327 Three Months Ended June 30, 2023 Six Months Ended June 30, 2023 As Previously Reported Adjustments As Restated As Previously Reported Adjustments As Restated Revenues: Product revenue $ 19,692 $ (143) $ 19,549 $ 38,979 $ (70) $ 38,909 Service revenue 10,552 (82) 10,470 19,131 (155) 18,976 Collaboration and license revenue 629 — 629 997 — 997 Grant revenue 156 — 156 378 — 378 Total revenues 31,029 (225) 30,804 59,485 (225) 59,260 Costs of goods sold and services: Cost of product revenue 7,236 (535) 6,701 14,269 311 14,580 Cost of service and other revenue 4,655 32 4,687 9,152 119 9,271 Total costs of goods sold and services 11,891 (503) 11,388 23,421 430 23,851 Gross profit 19,138 278 19,416 36,064 (655) 35,409 Operating expenses: Research and development 5,946 182 6,128 10,666 449 11,115 Selling, general, and administrative 21,591 (283) 21,308 42,474 (349) 42,125 Other lease costs 1,162 (194) 968 1,938 (170) 1,768 Impairment and restructuring — — — (33) — (33) Total operating expenses 28,699 (295) 28,404 55,045 (70) 54,975 Loss from operations (9,561) 573 (8,988) (18,981) (585) (19,566) Interest income 3,886 — 3,886 7,335 — 7,335 Other expense (154) (60) (214) (146) (73) (219) Loss before income taxes (5,829) 513 (5,316) (11,792) (658) (12,450) Income tax expense (235) — (235) (375) — (375) Net loss $ (6,064) $ 513 $ (5,551) $ (12,167) $ (658) $ (12,825) Net loss per common share, basic and diluted $ (0.16) $ 0.01 $ (0.15) $ (0.33) $ (0.01) $ (0.34) Weighted-average common shares outstanding, basic and diluted 37,494 — 37,494 37,411 — 37,411 Three Months Ended September 30, 2023 Nine Months Ended September 30, 2023 As Previously Reported Adjustments As Restated As Previously Reported Adjustments As Restated Revenues: Product revenue $ 19,660 $ 280 $ 19,940 $ 58,639 $ 210 $ 58,849 Service revenue 10,938 (55) 10,883 30,069 (210) 29,859 Collaboration and license revenue 237 — 237 1,234 — 1,234 Grant revenue 499 — 499 877 — 877 Total revenues 31,334 225 31,559 90,819 — 90,819 Costs of goods sold and services: Cost of product revenue 8,342 (1,133) 7,209 22,611 (822) 21,789 Cost of service and other revenue 5,209 (268) 4,941 14,361 (149) 14,212 Total costs of goods sold and services 13,551 (1,401) 12,150 36,972 (971) 36,001 Gross profit 17,783 1,626 19,409 53,847 971 54,818 Operating expenses: Research and development 7,200 539 7,739 17,866 988 18,854 Selling, general, and administrative 23,595 (45) 23,550 66,069 (394) 65,675 Other lease costs 758 170 928 2,696 — 2,696 Impairment and restructuring — — — (33) — (33) Total operating expenses 31,553 664 32,217 86,598 594 87,192 Loss from operations (13,770) 962 (12,808) (32,751) 377 (32,374) Interest income 4,185 — 4,185 11,520 — 11,520 Other income 2,030 80 2,110 1,884 7 1,891 Loss before income taxes (7,555) 1,042 (6,513) (19,347) 384 (18,963) Income tax expense (203) — (203) (578) — (578) Net loss $ (7,758) $ 1,042 $ (6,716) $ (19,925) $ 384 $ (19,541) Net loss per common share, basic and diluted $ (0.21) $ 0.03 $ (0.18) $ (0.53) $ 0.01 $ (0.52) Weighted-average common shares outstanding, basic and diluted 37,657 — 37,657 37,494 — 37,494 Three Months Ended December 31, 2023 As Previously Reported in the Company's Earnings Release Adjustments As Restated Revenues: Product revenue $ 20,821 $ — $ 20,821 Service and other revenue 10,230 — 10,230 Collaboration revenue 146 — 146 Grant revenue 352 — 352 Total revenue 31,549 — 31,549 Costs of goods sold: Cost of product revenue 10,025 (2,711) 7,314 Cost of service and other revenue 4,725 104 4,829 Total costs of goods sold and services 14,750 (2,607) 12,143 Gross profit 16,799 2,607 19,406 Operating expenses: Research and development 6,991 219 7,210 Selling, general and administrative 24,172 (736) 23,436 Other lease costs 1,016 — 1,016 Impairment and restructuring 1,570 (209) 1,361 Total operating expenses 33,749 (726) 33,023 Loss from operations (16,950) 3,333 (13,617) Interest income 4,319 — 4,319 Other income 363 263 626 Loss before income taxes (12,268) 3,596 (8,672) Income tax expense (141) — (141) Net loss $ (12,409) $ 3,596 $ (8,813) Net loss per common share, basic and diluted $ (0.33) $ 0.10 $ (0.23) Weighted-average common shares outstanding, basic and diluted 37,890 — 37,890 QUANTERIX CORPORATION RESTATED CONSOLIDATED STATEMENTS OF COMPREHENSIVE LOSS (amounts in thousands) Three Months Ended March 31, 2023 As Previously Reported Adjustments As Restated Net loss $ (6,103) $ (1,171) $ (7,274) Other comprehensive income, net of tax: Foreign currency translation adjustment 42 — 42 Total other comprehensive income 42 — 42 Comprehensive loss $ (6,061) $ (1,171) $ (7,232) Three Months Ended June 30, 2023 Six Months Ended June 30, 2023 As Previously Reported Adjustments As Restated As Previously Reported Adjustments As Restated Net loss $ (6,064) $ 513 $ (5,551) $ (12,167) $ (658) $ (12,825) Other comprehensive loss, net of tax: Foreign currency translation adjustment (244) — (244) (202) — (202) Total other comprehensive loss (244) — (244) (202) — (202) Comprehensive loss $ (6,308) $ 513 $ (5,795) $ (12,369) $ (658) $ (13,027) Three Months Ended September 30, 2023 Nine Months Ended September 30, 2023 As Previously Reported Adjustments As Restated As Previously Reported Adjustments As Restated Net loss $ (7,758) $ 1,042 $ (6,716) $ (19,925) $ 384 $ (19,541) Other comprehensive loss, net of tax: Unrealized losses on marketable securities (241) — (241) (241) — (241) Foreign currency translation adjustment (148) — (148) (350) — (350) Total other comprehensive loss (389) — (389) (591) — (591) Comprehensive loss $ (8,147) $ 1,042 $ (7,105) $ (20,516) $ 384 $ (20,132) QUANTERIX CORPORATION RESTATED CONSOLIDATED STATEMENTS OF STOCKHOLDERS’ EQUITY (amounts in thousands) Three Months Ended, March 31, 2023 Common Stock As Previously Reported Shares Amount Additional paid-in capital Accumulated other comprehensive income (loss) Accumulated Total stockholders' equity Balance at December 31, 2022 37,280 $ 37 $ 763,688 $ (2,623) $ (402,162) $ 358,940 Issuance of common stock under stock plans, including tax effects 144 — 551 — — 551 Stock-based compensation expense — — 3,902 — — 3,902 Foreign currency translation adjustment — — — 42 — 42 Net loss — — — — (6,103) (6,103) Balance at March 31, 2023 37,424 $ 37 $ 768,141 $ (2,581) $ (408,265) $ 357,332 Adjustments Balance at December 31, 2022 — $ — $ (59) $ 85 $ (1,034) $ (1,008) Issuance of common stock under stock plans, including tax effects — — — — — — Stock-based compensation expense — — 41 — — 41 Foreign currency translation adjustment — — — — — — Net loss — — — — (1,171) (1,171) Balance at March 31, 2023 — $ — $ (18) $ 85 $ (2,205) $ (2,138) As Restated Balance at December 31, 2022 37,280 $ 37 $ 763,629 $ (2,538) $ (403,196) $ 357,932 Issuance of common stock under stock plans, including tax effects 144 — 551 — — 551 Stock-based compensation expense — — 3,943 — — 3,943 Foreign currency translation adjustment — — — 42 — 42 Net loss — — — — (7,274) (7,274) Balance at March 31, 2023 37,424 $ 37 $ 768,123 $ (2,496) $ (410,470) $ 355,194 Three Months Ended June 30, 2023 Common Stock As Previously Reported Shares Amount Additional paid-in capital Accumulated other comprehensive income (loss) Accumulated Total stockholders' equity Balance at March 31, 2023 37,424 $ 37 $ 768,141 $ (2,581) $ (408,265) $ 357,332 Issuance of common stock under stock plans, including tax effects 142 — 139 — — 139 Stock-based compensation expense — — 4,193 — — 4,193 Foreign currency translation adjustment — — — (244) — (244) Net loss — — — — (6,064) (6,064) Balance at June 30, 2023 37,566 $ 37 $ 772,473 $ (2,825) $ (414,329) $ 355,356 Adjustments Balance at March 31, 2023 — $ — $ (18) $ 85 $ (2,205) $ (2,138) Issuance of common stock under stock plans, including tax effects — — — — — — Stock-based compensation expense — — 17 — — 17 Foreign currency translation adjustment — — — — — — Net loss — — — — 513 513 Balance at June 30, 2023 — $ — $ (1) $ 85 $ (1,692) $ (1,608) As Restated Balance at March 31, 2023 37,424 $ 37 $ 768,123 $ (2,496) $ (410,470) $ 355,194 Issuance of common stock under stock plans, including tax effects 142 — 139 — — 139 Stock-based compensation expense — — 4,210 — — 4,210 Foreign currency translation adjustment — — — (244) — (244) Net loss — — — — (5,551) (5,551) Balance at June 30, 2023 37,566 $ 37 $ 772,472 $ (2,740) $ (416,021) $ 353,748 Six Months Ended June 30, 2023 Common Stock As Previously Reported Shares Amount Additional paid-in capital Accumulated other comprehensive income (loss) Accumulated Total stockholders' equity Balance at December 31, 2022 37,280 $ 37 $ 763,688 $ (2,623) $ (402,162) $ 358,940 Issuance of common stock under stock plans, including tax effects 286 — 690 — — 690 Stock-based compensation expense — — 8,095 — — 8,095 Foreign currency translation adjustment — — — (202) — (202) Net loss — — — — (12,167) (12,167) Balance at June 30, 2023 37,566 $ 37 $ 772,473 $ (2,825) $ (414,329) $ 355,356 Adjustments Balance at December 31, 2022 — $ — $ (59) $ 85 $ (1,034) $ (1,008) Issuance of common stock under stock plans, including tax effects — — — — — — Stock-based compensation expense — — 58 — — 58 Foreign currency translation adjustment — — — — — — Net loss — — — — (658) (658) Balance at June 30, 2023 — $ — $ (1) $ 85 $ (1,692) $ (1,608) As Restated Balance at December 31, 2022 37,280 $ 37 $ 763,629 $ (2,538) $ (403,196) $ 357,932 Issuance of common stock under stock plans, including tax effects 286 — 690 — — 690 Stock-based compensation expense — — 8,153 — — 8,153 Foreign currency translation adjustment — — — (202) — (202) Net loss — — — — (12,825) (12,825) Balance at June 30, 2023 37,566 $ 37 $ 772,472 $ (2,740) $ (416,021) $ 353,748 Three Months Ended, September 30, 2023 Common Stock As Previously Reported Shares Amount Additional paid-in capital Accumulated other comprehensive income (loss) Accumulated Total stockholders' equity Balance at June 30, 2023 37,566 $ 37 $ 772,473 $ (2,825) $ (414,329) $ 355,356 Issuance of common stock under stock plans, including tax effects 273 1 1,799 — — 1,800 Stock-based compensation expense — — 4,343 — — 4,343 Unrealized loss on marketable securities, net of tax — — — (241) — (241) Foreign currency translation adjustment — — — (148) — (148) Net loss — — — — (7,758) (7,758) Balance at September 30, 2023 37,839 $ 38 $ 778,615 $ (3,214) $ (422,087) $ 353,352 Adjustments Balance at June 30, 2023 — $ — $ (1) $ 85 $ (1,692) $ (1,608) Issuance of common stock under stock plans, including tax effects — — — — — — Stock-based compensation expense — — 1 — — 1 Unrealized loss on marketable securities, net of tax — — — — — — Foreign currency translation adjustment — — — — — — Net loss — — — — 1,042 1,042 Balance at September 30, 2023 — $ — $ — $ 85 $ (650) $ (565) As Restated Balance at June 30, 2023 37,566 $ 37 $ 772,472 $ (2,740) $ (416,021) $ 353,748 Issuance of common stock under stock plans, including tax effects 273 1 1,799 — — 1,800 Stock-based compensation expense — — 4,344 — — 4,344 Unrealized loss on marketable securities, net of tax — — — (241) — (241) Foreign currency translation adjustment — — — (148) — (148) Net loss — — — — (6,716) (6,716) Balance at September 30, 2023 37,839 $ 38 $ 778,615 $ (3,129) $ (422,737) $ 352,787 Nine Months Ended, September 30, 2023 Common Stock As Previously Reported Shares Amount Additional paid-in capital Accumulated other comprehensive income (loss) Accumulated Total stockholders' equity Balance at December 31, 2022 37,280 $ 37 $ 763,688 $ (2,623) $ (402,162) $ 358,940 Issuance of common stock under stock plans, including tax effects 559 1 2,489 — — 2,490 Stock-based compensation expense — — 12,438 — — 12,438 Unrealized loss on marketable securities, net of tax — — — (241) — (241) Foreign currency translation adjustment — — — (350) — (350) Net loss — — — — (19,925) (19,925) Balance at September 30, 2023 37,839 $ 38 $ 778,615 $ (3,214) $ (422,087) $ 353,352 Adjustments Balance at December 31, 2022 — $ — $ (59) $ 85 $ (1,034) $ (1,008) Issuance of common stock under stock plans, including tax effects — — — — — — Stock-based compensation expense — — 59 — — 59 Unrealized loss on marketable securities, net of tax — — — — — — Foreign currency translation adjustment — — — — — — Net loss — — — — 384 384 Balance at September 30, 2023 — $ — $ — $ 85 $ (650) $ (565) As Restated Balance at December 31, 2022 37,280 $ 37 $ 763,629 $ (2,538) $ (403,196) $ 357,932 Issuance of common stock under stock plans, including tax effects 559 1 2,489 — — 2,490 Stock-based compensation expense — — 12,497 — — 12,497 Unrealized loss on marketable securities, net of tax — — — (241) — (241) Foreign currency translation adjustment — — — (350) — (350) Net loss — — — — (19,541) (19,541) Balance at September 30, 2023 37,839 $ 38 $ 778,615 $ (3,129) $ (422,737) $ 352,787 QUANTERIX CORPORATION RESTATED CONSOLIDATED STATEMENTS OF CASH FLOWS (amounts in thousands) Three Months Ended March 31, 2023 As Previously Reported Adjustments As Restated Operating activities: Net loss $ (6,103) $ (1,171) $ (7,274) Adjustments to reconcile net loss to net cash used in operating activities: Depreciation and amortization expense 1,439 15 1,454 Credit losses on accounts receivable 178 — 178 Non-cash operating lease expense 334 — 334 Stock-based compensation expense 3,902 41 3,943 Deferred taxes 270 — 270 Changes in operating assets and liabilities: Accounts receivable (3,741) — (3,741) Inventory (89) 619 530 Prepaid expenses and other assets (422) — (422) Other non-current assets (33) — (33) Accounts payable (1,271) — (1,271) Accrued compensation and benefits, other accrued expenses and other current liabilities (5,983) 174 (5,809) Deferred revenue 2,041 — 2,041 Operating lease liabilities 179 — 179 Other non-current liabilities (203) — (203) Net cash used in operating activities (9,502) (322) (9,824) Investing activities Purchases of property and equipment (136) — (136) Net cash used in investing activities (136) — (136) Financing activities Proceeds from stock options exercised 13 — 13 Proceeds from ESPP purchase 551 — 551 Payments for employee taxes on units withheld (13) — (13) Net cash provided by financing activities 551 — 551 Net decrease in cash, cash equivalents and restricted cash (9,087) (322) (9,409) Effect of foreign currency exchange rate on cash 24 — 24 Cash, cash equivalents and restricted cash at beginning of period 341,337 — 341,337 Cash, cash equivalents and restricted cash at end of period $ 332,274 $ (322) $ 331,952 Supplemental disclosure of cash flow information: Cash paid for income taxes $ 246 $ — $ 246 Noncash transactions: Purchases of property and equipment included in accounts payable and accrued expenses $ 147 $ — $ 147 Reconciliation of cash, cash equivalents and restricted cash: Cash and cash equivalents $ 329,354 $ — $ 329,354 Restricted cash $ 2,920 $ (322) $ 2,598 Total cash, cash equivalents, and restricted cash $ 332,274 $ (322) $ 331,952 Six Months Ended June 30, 2023 As Previously Reported Adjustments As Restated Cash flows from operating activities: Net loss $ (12,167) $ (658) $ (12,825) Adjustments to reconcile net loss to net cash used in operating activities: Depreciation and amortization expense 2,845 38 2,883 Credit losses on accounts receivable 324 — 324 Operating lease right-of-use asset amortization 1,002 — 1,002 Stock-based compensation expense 8,095 58 8,153 Other operating activity 548 54 602 Loss on disposal of fixed assets Impairment — 54 54 Changes in assets and liabilities: Accounts receivable (5,750) — (5,750) Inventory (1,181) 1 (1,180) Prepaid expenses and other current assets (527) — (527) Other non-current assets (965) 258 (707) Accounts payable (631) — (631) Accrued compensation and benefits, accrued expenses, and other current liabilities (1,326) 273 (1,053) Deferred revenue 1,666 — 1,666 Operating lease liabilities (730) — (730) Other non-current liabilities (72) — (72) Net cash used in operating activities (8,869) 78 (8,791) Cash flows from investing activities: Purchases of property and equipment (784) (163) (947) Net cash used in investing activities (784) (163) (947) Cash flows from financing activities: Proceeds from common stock issued under stock plans 777 — 777 Payments for employee taxes withheld on stock-based compensation awards (87) — (87) Net cash provided by financing activities 690 — 690 Net decrease in cash, cash equivalents, and restricted cash (8,963) (85) (9,048) Effect of exchange rate changes on cash, cash equivalents, and restricted cash (163) — (163) Cash, cash equivalents, and restricted cash at beginning of period 341,337 — 341,337 Cash, cash equivalents, and restricted cash at end of period $ 332,211 $ (85) $ 332,126 Supplemental disclosure of cash flow information: Cash paid for taxes $ 502 $ — $ 502 Shares received as consideration under product sales agreement (Note 4, 7) $ 1,000 $ — $ 1,000 Nine Months Ended September 30, 2023 As Previously Reported Adjustments As Restated Cash flows from operating activities: Net loss $ (19,925) $ 384 $ (19,541) Adjustments to reconcile net loss to net cash used in operating activities: Depreciation and amortization expense 4,788 55 4,843 Credit losses on accounts receivable 311 — 311 Foreign currency losses 359 — 359 Unrealized losses on marketable securities (241) — (241) Amortization of (discount) premium on marketable securities (1,249) — (1,249) Operating lease right-of-use asset amortization 1,518 — 1,518 Stock-based compensation expense 12,438 59 12,497 Impairment — 130 130 Deferred income taxes 242 — 242 Loss on disposal of fixed assets 46 — 46 Accounts receivable (5,615) — (5,615) Inventory (2,966) (734) (3,700) Prepaid expenses and other current assets (2,829) — (2,829) Other non-current assets (716) 14 (702) Accounts payable 948 — 948 Accrued compensation and benefits, accrued expenses, and other current liabilities 876 353 1,229 Deferred revenue 894 — 894 Operating lease liabilities (1,690) — (1,690) Other non-current liabilities (107) — (107) Net cash used in operating activities (12,918) 261 (12,657) Cash flows from investing activities: Purchases of marketable securities (125,200) — (125,200) Proceeds from marketable debt securities Purchases of property and equipment (1,572) (306) (1,878) Proceeds from RADx grant on assets purchased Net cash used in investing activities (126,772) (306) (127,078) Cash flows from financing activities: Proceeds from common stock issued under stock plans 2,632 — 2,632 Payments for employee taxes withheld on stock-based compensation awards (142) — (142) Net cash provided by financing activities 2,490 — 2,490 Net decrease in cash, cash equivalents, and restricted cash (137,200) (45) (137,245) Effect of exchange rate changes on cash, cash equivalents, and restricted cash (229) — (229) Cash, cash equivalents, and restricted cash at beginning of period 341,337 — 341,337 Cash, cash equivalents, and restricted cash at end of period $ 203,908 $ (45) $ 203,863 Supplemental disclosure of cash flow information: Cash paid for taxes $ 719 $ — $ 719 Shares received as consideration under product sales agreement (Note 4, 7) $ 775 $ — $ 775 QUANTERIX CORPORATION RESTATED CONSOLIDATED BALANCE SHEETS (amounts in thousands, except per share data) As of March 31, 2022 As Previously Reported Adjustments As Restated ASSETS Current assets: Cash and cash equivalents $ 374,317 $ 226 $ 374,543 Accounts receivable, net of allowance for expected credit losses 22,616 — 22,616 Inventory 22,669 1,757 24,426 Prepaid expenses and other current assets 14,104 (226) 13,878 Total current assets 433,706 1,757 435,463 Restricted cash 2,577 — 2,577 Property and equipment, net 19,683 (44) 19,639 Intangible assets, net 9,692 — 9,692 Goodwill 9,323 — 9,323 Right-of-use assets 29,298 617 29,915 Other non-current assets 378 (300) 78 Total assets $ 504,657 $ 2,030 $ 506,687 LIABILITIES AND STOCKHOLDERS' EQUITY Current liabilities: Accounts payable $ 4,262 $ (75) $ 4,187 Accrued compensation and benefits 8,139 57 8,196 Other accrued expenses 8,024 — 8,024 Deferred revenue 9,194 — 9,194 Short term lease liabilities 1,886 — 1,886 Other current liabilities 268 — 268 Total current liabilities 31,773 (18) 31,755 Deferred revenue, net of current portion 1,222 — 1,222 Long term lease liabilities 43,563 — 43,563 Other non-current liabilities 1,691 — 1,691 Total liabilities 78,249 (18) 78,231 Commitments and contingencies (Note 16) Stockholders’ equity: Common stock, $0.001 par value: Authorized shares: 120,000 shares; Issued and outstanding: 36,899 shares at March 31, 2022. 37 — 37 Additional paid-in capital 750,742 (25) 750,717 Accumulated other comprehensive (loss) income (756) — (756) Accumulated deficit (323,615) 2,073 (321,542) Total stockholders’ equity 426,408 2,048 428,456 Total liabilities and stockholders’ equity $ 504,657 $ 2,030 $ 506,687 As of June 30, 2022 As Previously Reported Adjustments As Restated ASSETS Current assets: Cash and cash equivalents $ 361,293 $ 258 $ 361,551 Accounts receivable, net of allowance for expected credit losses 19,683 — 19,683 Inventory 21,985 700 22,685 Prepaid expenses and other current assets 10,237 (258) 9,979 Total current assets 413,198 700 413,898 Restricted cash 2,594 — 2,594 Property and equipment, net 22,295 (319) 21,976 Intangible assets, net 8,527 — 8,527 Goodwill 8,675 — 8,675 Right-of-use assets 32,935 — 32,935 Other non-current assets 377 (300) 77 Total assets $ 488,601 $ 81 $ 488,682 LIABILITIES AND STOCKHOLDERS' EQUITY Current liabilities: Accounts payable $ 5,983 $ (75) $ 5,908 Accrued compensation and benefits 10,638 139 10,777 Other accrued expenses 4,799 — 4,799 Deferred revenue 12,157 — 12,157 Short term lease liabilities 2,696 — 2,696 Other current liabilities 396 — 396 Total current liabilities 36,669 64 36,733 Deferred revenue, net of current portion 1,531 — 1,531 Long term lease liabilities 43,135 — 43,135 Other non-current liabilities 1,810 — 1,810 Total liabilities 83,145 64 83,209 Commitments and contingencies (Note 16) Stockholders’ equity: Common stock, $0.001 par value: Authorized shares: 120,000 shares; Issued and outstanding: 36,975 shares at June 30, 2022. 37 — 37 Additional paid-in capital 756,139 (52) 756,087 Accumulated other comprehensive (loss) income (2,203) — (2,203) Accumulated deficit (348,517) 69 (348,448) Total stockholders’ equity 405,456 17 405,473 Total liabilities and stockholders’ equity $ 488,601 $ 81 $ 488,682 As of September 30, 2022 As Previously Reported Adjustments As Restated ASSETS Current assets: Cash and cash equivalents $ 343,743 $ — $ 343,743 Accounts receivable, net of allowance for expected credit losses 18,330 — 18,330 Inventory 18,236 (192) 18,044 Prepaid expenses and other current assets 6,475 (400) 6,075 Total current assets 386,784 (592) 386,192 Restricted cash 2,596 — 2,596 Property and equipment, net 21,441 (246) 21,195 Intangible assets, net 7,511 — 7,511 Right-of-use assets 27,165 — 27,165 Other non-current assets 1,200 (290) 910 Total assets $ 446,697 $ (1,128) $ 445,569 LIABILITIES AND STOCKHOLDERS' EQUITY Current liabilities: Accounts payable $ 2,106 $ — $ 2,106 Accrued compensation and benefits 10,503 257 10,760 Other accrued expenses 5,951 90 6,041 Deferred revenue 8,976 — 8,976 Short-term lease liabilities 767 — 767 Other current liabilities 268 — 268 Total current liabilities 28,571 347 28,918 Deferred revenue, net of current portion 1,591 — 1,591 Long-term lease liabilities 42,196 — 42,196 Other non-current liabilities 1,570 — 1,570 Total liabilities 73,928 347 74,275 Commitments and contingencies (Note 16) Stockholders’ equity: Common stock, $0.001 par value: Authorized shares: 120,000 shares; Issued and outstanding: 37,094 shares at September 30, 2022. 37 — 37 Additional paid-in capital 759,312 (58) 759,254 Accumulated other comprehensive (loss) income (2,999) — (2,999) Accumulated deficit (383,581) (1,417) (384,998) Total stockholders’ equity 372,769 (1,475) 371,294 Total liabilities and stockholders’ equity $ 446,697 $ (1,128) $ 445,569 QUANTERIX CORPORATION RESTATED CONSOLIDATED STATEMENTS OF OPERATIONS (amounts in thousands, except per share data) Three Months Ended March 31, 2022 As Previously Reported Adjustments As Restated Revenues: Product revenue $ 20,656 $ — $ 20,656 Service and other revenue 8,810 — 8,810 Collaboration revenue 86 — 86 Total revenue 29,552 — 29,552 Costs of goods sold: Cost of product revenue 10,746 711 11,457 Cost of service and other revenue 4,247 (249) 3,998 Total costs of goods sold and services 14,993 462 15,455 Gross profit 14,559 (462) 14,097 Operating expenses: Research and development 7,034 (154) 6,880 Selling, general and administrative 25,712 (541) 25,171 Total operating expenses 32,746 (695) 32,051 Loss from operations (18,187) 233 (17,954) Interest income 52 — 52 Other expense (217) — (217) Loss before income taxes (18,352) 233 (18,119) Income tax benefit 199 — 199 Net loss $ (18,153) $ 233 $ (17,920) Net loss per common share, basic and diluted $ (0.49) $ — $ (0.49) Weighted-average common shares outstanding, basic and diluted 36,851 — 36,851 Three Months Ended June 30, 2022 Six Months Ended June 30, 2022 As Previously Reported Adjustments As Restated As Previously Reported Adjustments As Restated Revenues: Product revenue $ 14,785 $ — $ 14,785 $ 35,441 $ — $ 35,441 Service and other revenue 8,548 — 8,548 17,358 — 17,358 Collaboration revenue 92 — 92 178 — 178 Grant revenue 75 — 75 75 — 75 Total revenue 23,500 — 23,500 53,052 — 53,052 Costs of goods sold: Cost of product revenue 9,921 625 10,546 20,667 1,336 22,003 Cost of service and other revenue 4,868 (231) 4,637 9,115 (480) 8,635 Total costs of goods sold and services 14,789 394 15,183 29,782 856 30,638 Gross profit 8,711 (394) 8,317 23,270 (856) 22,414 Operating expenses: Research and development 6,625 632 7,257 13,659 478 14,137 Selling, general and administrative 27,045 771 27,816 52,757 230 52,987 Impairment and restructuring — 207 207 — 207 207 Total operating expenses 33,670 1,610 35,280 66,416 915 67,331 Loss from operations (24,959) (2,004) (26,963) (43,146) (1,771) (44,917) Interest income 552 — 552 604 — 604 Other expense (358) — (358) (575) — (575) Loss before income taxes (24,765) (2,004) (26,769) (43,117) (1,771) (44,888) Income tax (expense) benefit (137) — (137) 62 — 62 Net loss $ (24,902) $ (2,004) $ (26,906) $ (43,055) $ (1,771) $ (44,826) Net loss per common share, basic and diluted $ (0.67) $ (0.06) $ (0.73) $ (1.17) $ (0.05) $ (1.22) Weighted-average common shares outstanding, basic and diluted 36,922 — 36,922 36,887 — 36,887 Three Months Ended September 30, 2022 Nine Months Ended September 30

Restatement of Financial Stat_4

Restatement of Financial Statements - Balance Sheet (Details) - USD ($) $ in Thousands	Dec. 31, 2023	Sep. 30, 2023	Jun. 30, 2023	Mar. 31, 2023	Dec. 31, 2022	Sep. 30, 2022	Jun. 30, 2022	Mar. 31, 2022	Dec. 31, 2021	Dec. 31, 2020
Current assets:
Cash and cash equivalents	$ 174,422	$ 201,261	$ 329,525	$ 329,354	$ 338,740	$ 343,743	$ 361,551	$ 374,543
Marketable securities	146,902	126,449			0
Accounts receivable, net of allowance for expected credit losses	25,414	24,083	24,369	22,546	19,017	18,330	19,683	22,616
Inventory	26,123	20,857	18,333	16,629	16,964	18,044	22,685	24,426
Prepaid expenses and other current assets	9,234	9,216	6,897	6,945	6,803	6,075	9,979	13,878
Total current assets	382,095	381,866	379,124	375,474	381,524	386,192	413,898	435,463
Restricted cash	2,604	2,602	2,601	2,598	2,597	2,596	2,594	2,577
Property and equipment, net	17,926	17,287	18,048	18,690	19,811	21,195	21,976	19,639
Intangible assets, net	6,034	6,003	6,476	7,129	7,516	7,511	8,527	9,692
Operating lease right-of-use assets	18,251	19,860	20,380	20,891	21,223	27,165	32,935	29,915
Other non-current assets	1,657	1,603	1,637	938	911	910	77	78
Total assets	428,567	429,221	428,266	425,720	433,582	445,569	488,682	506,687
Current liabilities:
Accounts payable	5,048	4,786	3,238	2,585	3,836	2,106	5,908	4,187
Accrued compensation and benefits	14,170	10,505	8,175	5,411	11,035	10,760	10,777	8,196
Accrued expenses and other current liabilities	6,055	6,686	6,791		5,147
Deferred revenue	9,468	9,827	10,421	10,682	8,644	8,976	12,157	9,194
Operating lease liabilities	4,241	4,093	3,986	3,875	2,687	767	2,696	1,886
Total current liabilities	38,982	35,897	32,611	27,482	31,349	28,918	36,733	31,755
Deferred revenue, net of current portion	1,227	1,126	1,304	1,419	1,415	1,591	1,531	1,222
Operating lease liabilities, net of current portion	37,223	38,306	39,378	40,409	41,417	42,196	43,135	43,563
Other non-current liabilities	1,177	1,105	1,225	1,216	1,469	1,570	1,810	1,691
Total liabilities	78,609	76,434	74,518	70,526	75,650	74,275	83,209	78,231
Commitments and contingencies (Note 16)
Stockholders’ equity:
Common stock	38	38	37	37	37	37	37	37
Additional paid-in capital	783,142	778,615	772,472	768,123	763,629	759,254	756,087	750,717
Accumulated other comprehensive loss	(1,672)	(3,129)	(2,740)	(2,496)	(2,538)	(2,999)	(2,203)	(756)
Accumulated deficit	(431,550)	(422,737)	(416,021)	(410,470)	(403,196)	(384,998)	(348,448)	(321,542)
Total stockholders’ equity	349,958	352,787	353,748	355,194	357,932	371,294	405,473	428,456	$ 442,792	$ 205,825
Total liabilities and stockholders’ equity	428,567	429,221	428,266	425,720	433,582	445,569	488,682	506,687
As Previously Reported
Current assets:
Cash and cash equivalents	174,422	201,261	329,525	329,354	338,740	343,743	361,293	374,317
Marketable securities	146,902	126,449
Accounts receivable, net of allowance for expected credit losses	25,414	24,083	24,423	22,546	19,017	18,330	19,683	22,616
Inventory	22,365	19,945	18,156	17,070	16,786	18,236	21,985	22,669
Prepaid expenses and other current assets	9,291	9,273	6,954	7,002	6,860	6,475	10,237	14,104
Total current assets	378,394	381,011	379,058	375,972	381,403	386,784	413,198	433,706
Restricted cash	2,604	2,647	2,686	2,920	2,597	2,596	2,594	2,577
Property and equipment, net	17,926	17,517	18,328	19,056	20,162	21,441	22,295	19,683
Intangible assets, net	6,034	6,003	6,476	7,129	7,516	7,511	8,527	9,692
Operating lease right-of-use assets	18,251	19,860	20,380	20,891	21,223	27,165	32,935	29,298
Other non-current assets	1,802	2,004	2,282	1,345	1,298	1,200	377	378
Total assets	425,011	429,042	429,210	427,313	434,199	446,697	488,601	504,657
Current liabilities:
Accounts payable	5,048	4,786	3,238	2,585	3,836	2,106	5,983	4,262
Accrued compensation and benefits	13,659	9,775	7,525	4,880	10,658	10,503	10,638	8,139
Accrued expenses and other current liabilities	6,041	6,672	6,777		5,133
Deferred revenue	9,468	9,827	10,421	10,682	8,644	8,976	12,157	9,194
Operating lease liabilities	4,241	4,093	3,986	3,875	2,687	767	2,696	1,886
Total current liabilities	38,457	35,153	31,947	26,937	30,958	28,571	36,669	31,773
Deferred revenue, net of current portion	1,227	1,126	1,304	1,419	1,415	1,591	1,531	1,222
Operating lease liabilities, net of current portion	37,223	38,306	39,378	40,409	41,417	42,196	43,135	43,563
Other non-current liabilities	1,177	1,105	1,225	1,216	1,469	1,570	1,810	1,691
Total liabilities	78,084	75,690	73,854	69,981	75,259	73,928	83,145	78,249
Commitments and contingencies (Note 16)
Stockholders’ equity:
Common stock	38	38	37	37	37	37	37	37
Additional paid-in capital	783,142	778,615	772,473	768,141	763,688	759,312	756,139	750,742
Accumulated other comprehensive loss	(1,757)	(3,214)	(2,825)	(2,581)	(2,623)	(2,999)	(2,203)	(756)
Accumulated deficit	(434,496)	(422,087)	(414,329)	(408,265)	(402,162)	(383,581)	(348,517)	(323,615)
Total stockholders’ equity	346,927	353,352	355,356	357,332	358,940	372,769	405,456	426,408	440,952	206,125
Total liabilities and stockholders’ equity	425,011	429,042	429,210	427,313	434,199	446,697	488,601	504,657
Adjustments
Current assets:
Cash and cash equivalents	0	0	0	0	0	0	258	226
Marketable securities	0	0
Accounts receivable, net of allowance for expected credit losses	0	0	(54)	0	0	0	0	0
Inventory	3,758	912	177	(441)	178	(192)	700	1,757
Prepaid expenses and other current assets	(57)	(57)	(57)	(57)	(57)	(400)	(258)	(226)
Total current assets	3,701	855	66	(498)	121	(592)	700	1,757
Restricted cash	0	(45)	(85)	(322)	0	0	0	0
Property and equipment, net	0	(230)	(280)	(366)	(351)	(246)	(319)	(44)
Intangible assets, net	0	0	0	0	0	0	0	0
Operating lease right-of-use assets	0	0	0	0	0	0	0	617
Other non-current assets	(145)	(401)	(645)	(407)	(387)	(290)	(300)	(300)
Total assets	3,556	179	(944)	(1,593)	(617)	(1,128)	81	2,030
Current liabilities:
Accounts payable	0	0	0	0	0	0	(75)	(75)
Accrued compensation and benefits	511	730	650	531	377	257	139	57
Accrued expenses and other current liabilities	14	14	14		14
Deferred revenue	0	0	0	0	0	0	0	0
Operating lease liabilities	0	0	0	0	0	0	0	0
Total current liabilities	525	744	664	545	391	347	64	(18)
Deferred revenue, net of current portion	0	0	0	0	0	0	0	0
Operating lease liabilities, net of current portion	0	0	0	0	0	0	0	0
Other non-current liabilities	0	0	0	0	0	0	0	0
Total liabilities	525	744	664	545	391	347	64	(18)
Commitments and contingencies (Note 16)
Stockholders’ equity:
Common stock	0	0	0	0	0	0	0	0
Additional paid-in capital	0	0	(1)	(18)	(59)	(58)	(52)	(25)
Accumulated other comprehensive loss	85	85	85	85	85	0	0	0
Accumulated deficit	2,946	(650)	(1,692)	(2,205)	(1,034)	(1,417)	69	2,073
Total stockholders’ equity	3,031	(565)	(1,608)	(2,138)	(1,008)	(1,475)	17	2,048	$ 1,840	$ (300)
Total liabilities and stockholders’ equity	$ 3,556	$ 179	$ (944)	$ (1,593)	$ (617)	$ (1,128)	$ 81	$ 2,030

Restatement of Financial Stat_5

Restatement of Financial Statements - Statement Of Operations (Details) - USD ($) $ / shares in Units, $ in Thousands	3 Months Ended								6 Months Ended		9 Months Ended		12 Months Ended
	Dec. 31, 2023	Sep. 30, 2023	Jun. 30, 2023	Mar. 31, 2023	Dec. 31, 2022	Sep. 30, 2022	Jun. 30, 2022	Mar. 31, 2022	Jun. 30, 2023	Jun. 30, 2022	Sep. 30, 2023	Sep. 30, 2022	Dec. 31, 2023	Dec. 31, 2022	Dec. 31, 2021
Revenues:
Total revenues	$ 31,549	$ 31,559	$ 30,804	$ 28,456	$ 25,824	$ 26,646	$ 23,500	$ 29,552	$ 59,260	$ 53,052	$ 90,819	$ 79,698	$ 122,368	$ 105,522	$ 110,556
Costs of goods sold and services:
Total costs of goods sold and services	12,143	12,150	11,388	12,463	13,180	16,341	15,183	15,455	23,851	30,638	36,001	46,979	48,144	60,159	45,784
Gross profit	19,406	19,409	19,416	15,993	12,644	10,305	8,317	14,097	35,409	22,414	54,818	32,719	74,224	45,363	64,772
Operating expenses:
Research and development	7,210	7,739	6,128	4,987	5,843	6,829	7,257	6,880	11,115	14,137	18,854	20,966	26,064	26,809	28,383
Selling, general and administrative	23,436	23,550	21,308	20,817	18,385	20,479	27,816	25,171	42,125	52,987	65,675	73,466	89,111	91,851	92,910
Other lease costs	1,016	928	968	800	802	609			1,768		2,696	609	3,712	1,411	0
Impairment and restructuring	1,361	0	0	(33)	9,006	20,343	207		(33)	207	(33)	20,550	1,328	29,556	0
Total operating expenses	33,023	32,217	28,404	26,571	34,036	48,260	35,280	32,051	54,975	67,331	87,192	115,591	120,215	149,627	121,293
Loss from operations	(13,617)	(12,808)	(8,988)	(10,578)	(21,392)	(37,955)	(26,963)	(17,954)	(19,566)	(44,917)	(32,374)	(82,872)	(45,991)	(104,264)	(56,521)
Interest income (expense), net	4,319	4,185	3,886	3,449	2,815	1,712	552	52	7,335	604	11,520	2,316	15,839	5,131	(403)
Other income (expense), net	626	2,110	(214)	(5)	533	(235)	(358)	(217)	(219)	(575)	1,891	(810)	2,517	(277)	1,340
Loss before income taxes	(8,672)	(6,513)	(5,316)	(7,134)	(18,044)	(36,478)	(26,769)	(18,119)	(12,450)	(44,888)	(18,963)	(81,366)	(27,635)	(99,410)	(55,584)
Income tax (expense) benefit	(141)	(203)	(235)	(140)	(154)	(72)	(137)	199	(375)	62	(578)	(10)	(719)	(164)	36
Net loss	$ (8,813)	$ (6,716)	$ (5,551)	$ (7,274)	$ (18,198)	$ (36,550)	$ (26,906)	$ (17,920)	$ (12,825)	$ (44,826)	$ (19,541)	$ (81,376)	$ (28,354)	$ (99,574)	$ (55,548)
Net loss per common share, basic (in dollars per share)	$ (0.23)	$ (0.18)	$ (0.15)	$ (0.19)	$ (0.49)	$ (0.99)	$ (0.73)	$ (0.49)	$ (0.34)	$ (1.22)	$ (0.52)	$ (2.20)	$ (0.75)	$ (2.69)	$ (1.54)
Net loss per common share, diluted (in dollars per share)	$ (0.23)	$ (0.18)	$ (0.15)	$ (0.19)	$ (0.49)	$ (0.99)	$ (0.73)	$ (0.49)	$ (0.34)	$ (1.22)	$ (0.52)	$ (2.20)	$ (0.75)	$ (2.69)	$ (1.54)
Weighted-average common shares outstanding, basic (in shares)	37,890,000	37,657,000	37,494,000	37,327,000	37,160,000	37,005,000	36,922,000	36,851,000	37,411,000	36,887,000	37,494,000	36,927,000	37,594,000	36,991,000	35,997,000
Weighted-average common shares outstanding, diluted (in shares)	37,890,000	37,657,000	37,494,000	37,327,000	37,160,000	37,005,000	36,922,000	36,851,000	37,411,000	36,887,000	37,494,000	36,927,000	37,594,000	36,991,000	35,997,000
Common stock, par value	$ 0.001	$ 0.001	$ 0.001	$ 0.001	$ 0.001	$ 0.001	$ 0.001	$ 0.001	$ 0.001	$ 0.001	$ 0.001	$ 0.001	$ 0.001	$ 0.001
Common stock, shares issued	38,014,000	37,839,000	37,566,000	37,424,000	37,280,000	37,094,000	36,975,000	36,899,000	37,566,000	36,975,000	37,839,000	37,094,000	38,014,000	37,280,000
Common stock, shares outstanding	38,014,000	37,839,000	37,566,000	37,424,000	37,280,000	37,094,000	36,975,000	36,899,000	37,566,000	36,975,000	37,839,000	37,094,000	38,014,000	37,280,000
Marketable securities	$ 146,902	$ 126,449			$ 0						$ 126,449		$ 146,902	$ 0
Product revenue
Revenues:
Total revenues	20,821	19,940	$ 19,549	$ 19,360	16,674	$ 17,693	$ 14,785	$ 20,656	$ 38,909	$ 35,441	58,849	$ 53,134	79,670	69,808	$ 81,062
Costs of goods sold and services:
Total costs of goods sold and services	7,314	7,209	6,701	7,879	9,491	11,347	10,546	11,457	14,580	22,003	21,789	33,350	29,103	42,841	32,114
Service revenue
Revenues:
Total revenues														34,495
Service and other revenue
Revenues:
Total revenues	10,230	10,883	10,470	8,506	8,767	8,370	8,548	8,810	18,976	17,358	29,859	25,728	40,089	34,495	23,629
Costs of goods sold and services:
Total costs of goods sold and services	4,829	4,941	4,687	4,584	3,689	4,994	4,637	3,998	9,271	8,635	14,212	13,629	19,041	17,318	13,670
Collaboration and license revenue
Revenues:
Total revenues	146	237	629	368	170	301	92	86	997	178	1,234	479	1,380	649	648
Grant revenue
Revenues:
Total revenues	352	499	156	222	213	282	75		378	75	877	357	1,229	570	5,217
As Previously Reported
Revenues:
Total revenues	31,549	31,334	31,029	28,456	25,824	26,646	23,500	29,552	59,485	53,052	90,819	79,698	122,368	105,522	110,556
Costs of goods sold and services:
Total costs of goods sold and services	14,750	13,551	11,891	11,530	13,232	15,702	14,789	14,993	23,421	29,782	36,972	45,484	51,722	58,716	48,828
Gross profit	16,799	17,783	19,138	16,926	12,592	10,944	8,711	14,559	36,064	23,270	53,847	34,214	70,646	46,806	61,728
Operating expenses:
Research and development	6,991	7,200	5,946	4,720	5,600	6,631	6,625	7,034	10,666	13,659	17,866	20,290	24,857	25,890	27,978
Selling, general and administrative	24,172	23,595	21,591	20,883	19,272	19,966	27,045	25,712	42,474	52,757	66,069	72,723	90,241	91,995	92,336
Other lease costs	1,016	758	1,162	776	669	609			1,938		2,696	609	3,712	1,278
Impairment and restructuring	1,570	0	0	(33)	9,006	20,341	0		(33)	0	(33)	20,341	1,537	29,347
Total operating expenses	33,749	31,553	28,699	26,346	34,547	47,547	33,670	32,746	55,045	66,416	86,598	113,963	120,347	148,510	120,314
Loss from operations	(16,950)	(13,770)	(9,561)	(9,420)	(21,955)	(36,603)	(24,959)	(18,187)	(18,981)	(43,146)	(32,751)	(79,749)	(49,701)	(101,704)	(58,586)
Interest income (expense), net	4,319	4,185	3,886	3,449	2,815	1,712	552	52	7,335	604	11,520	2,316	15,839	5,131	(403)
Other income (expense), net	363	2,030	(154)	8	614	(101)	(358)	(217)	(146)	(575)	1,884	(676)	2,247	(62)	1,265
Loss before income taxes	(12,268)	(7,555)	(5,829)	(5,963)	(18,526)	(34,992)	(24,765)	(18,352)	(11,792)	(43,117)	(19,347)	(78,109)	(31,615)	(96,635)	(57,724)
Income tax (expense) benefit	(141)	(203)	(235)	(140)	(55)	(72)	(137)	199	(375)	62	(578)	(10)	(719)	(65)	36
Net loss	$ (12,409)	$ (7,758)	$ (6,064)	$ (6,103)	$ (18,581)	$ (35,064)	$ (24,902)	$ (18,153)	$ (12,167)	$ (43,055)	$ (19,925)	$ (78,119)	$ (32,334)	$ (96,700)	$ (57,688)
Net loss per common share, basic (in dollars per share)	$ (0.33)	$ (0.21)	$ (0.16)	$ (0.16)	$ (0.50)	$ (0.95)	$ (0.67)	$ (0.49)	$ (0.33)	$ (1.17)	$ (0.53)	$ (2.12)	$ (0.86)	$ (2.61)	$ (1.60)
Net loss per common share, diluted (in dollars per share)	$ (0.33)	$ (0.21)	$ (0.16)	$ (0.16)	$ (0.50)	$ (0.95)	$ (0.67)	$ (0.49)	$ (0.33)	$ (1.17)	$ (0.53)	$ (2.12)	$ (0.86)	$ (2.61)	$ (1.60)
Weighted-average common shares outstanding, basic (in shares)	37,890,000	37,657,000	37,494,000	37,327,000	37,160,000	37,005,000	36,922,000	36,851,000	37,411,000	36,887,000	37,494,000	36,927,000	37,594,000	36,991,000	35,997,000
Weighted-average common shares outstanding, diluted (in shares)	37,890,000	37,657,000	37,494,000	37,327,000	37,160,000	37,005,000	36,922,000	36,851,000	37,411,000	36,887,000	37,494,000	36,927,000	37,594,000	36,991,000	35,997,000
Marketable securities	$ 146,902	$ 126,449									$ 126,449		$ 146,902
As Previously Reported | Product revenue
Revenues:
Total revenues	20,821	19,660	$ 19,692	$ 19,287	$ 16,674	$ 17,693	$ 14,785	$ 20,656	$ 38,979	$ 35,441	58,639	$ 53,134	79,460	$ 69,808	$ 81,062
Costs of goods sold and services:
Total costs of goods sold and services	10,025	8,342	7,236	7,033	9,631	10,511	9,921	10,746	14,269	20,667	22,611	31,178	32,636	40,809	34,149
As Previously Reported | Service and other revenue
Revenues:
Total revenues	10,230	10,938	10,552	8,579	8,767	8,370	8,548	8,810	19,131	17,358	30,069	25,728	40,299	34,495	23,629
Costs of goods sold and services:
Total costs of goods sold and services	4,725	5,209	4,655	4,497	3,601	5,191	4,868	4,247	9,152	9,115	14,361	14,306	19,086	17,907	14,679
As Previously Reported | Collaboration and license revenue
Revenues:
Total revenues	146	237	629	368	170	301	92	86	997	178	1,234	479	1,380	649	648
As Previously Reported | Grant revenue
Revenues:
Total revenues	352	499	156	222	213	282	75		378	75	877	357	1,229	570	5,217
Adjustments
Revenues:
Total revenues	0	225	(225)	0	0	0	0	0	(225)	0	0	0	0	0	0
Costs of goods sold and services:
Total costs of goods sold and services	(2,607)	(1,401)	(503)	933	(52)	639	394	462	430	856	(971)	1,495	(3,578)	1,443	(3,044)
Gross profit	2,607	1,626	278	(933)	52	(639)	(394)	(462)	(655)	(856)	971	(1,495)	3,578	(1,443)	3,044
Operating expenses:
Research and development	219	539	182	267	243	198	632	(154)	449	478	988	676	1,207	919	405
Selling, general and administrative	(736)	(45)	(283)	(66)	(887)	513	771	(541)	(349)	230	(394)	743	(1,130)	(144)	574
Other lease costs	0	170	(194)	24	133	0			(170)		0	0	0	133
Impairment and restructuring	(209)	0	0	0	0	2	207		0	207	0	209	(209)	209
Total operating expenses	(726)	664	(295)	225	(511)	713	1,610	(695)	(70)	915	594	1,628	(132)	1,117	979
Loss from operations	3,333	962	573	(1,158)	563	(1,352)	(2,004)	233	(585)	(1,771)	377	(3,123)	3,710	(2,560)	2,065
Interest income (expense), net	0	0	0	0	0	0	0	0	0	0	0	0	0	0	0
Other income (expense), net	263	80	(60)	(13)	(81)	(134)	0	0	(73)	0	7	(134)	270	(215)	75
Loss before income taxes	3,596	1,042	513	(1,171)	482	(1,486)	(2,004)	233	(658)	(1,771)	384	(3,257)	3,980	(2,775)	2,140
Income tax (expense) benefit	0	0	0	0	(99)	0	0	0	0	0	0	0	0	(99)	0
Net loss	$ 3,596	$ 1,042	$ 513	$ (1,171)	$ 383	$ (1,486)	$ (2,004)	$ 233	$ (658)	$ (1,771)	$ 384	$ (3,257)	$ 3,980	$ (2,874)	$ 2,140
Net loss per common share, basic (in dollars per share)	$ 0.10	$ 0.03	$ 0.01	$ (0.03)	$ 0.01	$ (0.04)	$ (0.06)	$ 0	$ (0.01)	$ (0.05)	$ 0.01	$ (0.08)	$ 0.11	$ (0.08)	$ 0.06
Net loss per common share, diluted (in dollars per share)	$ 0.10	$ 0.03	$ 0.01	$ (0.03)	$ 0.01	$ (0.04)	$ (0.06)	$ 0	$ (0.01)	$ (0.05)	$ 0.01	$ (0.08)	$ 0.11	$ (0.08)	$ 0.06
Weighted-average common shares outstanding, basic (in shares)	0	0	0	0	0	0	0	0	0	0	0	0	0	0	0
Weighted-average common shares outstanding, diluted (in shares)	0	0	0	0	0	0	0	0	0	0	0	0	0	0	0
Marketable securities	$ 0	$ 0									$ 0		$ 0
Adjustments | Product revenue
Revenues:
Total revenues	0	280	$ (143)	$ 73	$ 0	$ 0	$ 0	$ 0	$ (70)	$ 0	210	$ 0	210	$ 0	$ 0
Costs of goods sold and services:
Total costs of goods sold and services	(2,711)	(1,133)	(535)	846	(140)	836	625	711	311	1,336	(822)	2,172	(3,533)	2,032	(2,035)
Adjustments | Service and other revenue
Revenues:
Total revenues	0	(55)	(82)	(73)	0	0	0	0	(155)	0	(210)	0	(210)	0	0
Costs of goods sold and services:
Total costs of goods sold and services	104	(268)	32	87	88	(197)	(231)	(249)	119	(480)	(149)	(677)	(45)	(589)	(1,009)
Adjustments | Collaboration and license revenue
Revenues:
Total revenues	0	0	0	0	0	0	0	$ 0	0	0	0	0	0	0	0
Adjustments | Grant revenue
Revenues:
Total revenues	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0		$ 0	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0

Restatement of Financial Stat_6

Restatement of Financial Statements - Statement Of Comprehensive Loss (Details) - USD ($) $ in Thousands	3 Months Ended								6 Months Ended		9 Months Ended		12 Months Ended
	Dec. 31, 2023	Sep. 30, 2023	Jun. 30, 2023	Mar. 31, 2023	Dec. 31, 2022	Sep. 30, 2022	Jun. 30, 2022	Mar. 31, 2022	Jun. 30, 2023	Jun. 30, 2022	Sep. 30, 2023	Sep. 30, 2022	Dec. 31, 2023	Dec. 31, 2022	Dec. 31, 2021
Reclassification [Line Items]
Net loss	$ (8,813)	$ (6,716)	$ (5,551)	$ (7,274)	$ (18,198)	$ (36,550)	$ (26,906)	$ (17,920)	$ (12,825)	$ (44,826)	$ (19,541)	$ (81,376)	$ (28,354)	$ (99,574)	$ (55,548)
Unrealized gains (losses) on marketable securities		(241)									(241)		325	0	0
Foreign currency translation adjustment		(148)	(244)	42		(796)	(1,447)	(1,197)	(202)	(2,644)	(350)	(3,440)	541	(2,979)	(1,993)
Total other comprehensive income (loss)		(389)	(244)	42		(796)	(1,447)	(1,197)	(202)	(2,644)	(591)	(3,440)	866	(2,979)	(1,993)
Comprehensive loss		(7,105)	(5,795)	(7,232)		(37,346)	(28,353)	(19,117)	(13,027)	(47,470)	(20,132)	(84,816)	(27,488)	(102,553)	(57,541)
As Previously Reported
Reclassification [Line Items]
Net loss	(12,409)	(7,758)	(6,064)	(6,103)	(18,581)	(35,064)	(24,902)	(18,153)	(12,167)	(43,055)	(19,925)	(78,119)	(32,334)	(96,700)	(57,688)
Unrealized gains (losses) on marketable securities		(241)									(241)		325
Foreign currency translation adjustment		(148)	(244)	42		(796)	(1,447)	(1,197)	(202)	(2,644)	(350)	(3,440)	541	(3,064)	(1,993)
Total other comprehensive income (loss)		(389)	(244)	42		(796)	(1,447)	(1,197)	(202)	(2,644)	(591)	(3,440)	866	(3,064)	(1,993)
Comprehensive loss		(8,147)	(6,308)	(6,061)		(35,860)	(26,349)	(19,350)	(12,369)	(45,699)	(20,516)	(81,559)	(31,468)	(99,764)	(59,681)
Adjustments
Reclassification [Line Items]
Net loss	$ 3,596	1,042	513	(1,171)	$ 383	(1,486)	(2,004)	233	(658)	(1,771)	384	(3,257)	3,980	(2,874)	2,140
Unrealized gains (losses) on marketable securities		0									0		0
Foreign currency translation adjustment		0	0	0		0	0	0	0	0	0	0	0	85	0
Total other comprehensive income (loss)		0	0	0		0	0	0	0	0	0	0	0	85	0
Comprehensive loss		$ 1,042	$ 513	$ (1,171)		$ (1,486)	$ (2,004)	$ 233	$ (658)	$ (1,771)	$ 384	$ (3,257)	$ 3,980	$ (2,789)	$ 2,140

Restatement of Financial Stat_7

Restatement of Financial Statements - Statement of Stockholders' Equity (Details) - USD ($) shares in Thousands, $ in Thousands	3 Months Ended								6 Months Ended		9 Months Ended		12 Months Ended
	Dec. 31, 2023	Sep. 30, 2023	Jun. 30, 2023	Mar. 31, 2023	Dec. 31, 2022	Sep. 30, 2022	Jun. 30, 2022	Mar. 31, 2022	Jun. 30, 2023	Jun. 30, 2022	Sep. 30, 2023	Sep. 30, 2022	Dec. 31, 2023	Dec. 31, 2022	Dec. 31, 2021
Reclassification [Line Items]
Beginning Balance	$ 352,787	$ 353,748	$ 355,194	$ 357,932	$ 371,294	$ 405,473	$ 428,456	$ 442,792	$ 357,932	$ 442,792	$ 357,932	$ 442,792	$ 357,932	$ 442,792	$ 205,825
Issuance of common stock under stock plans, including tax effects		1,800	139	551		407	211	979	690	1,190	2,490	1,597	2,691	2,310	8,815
Sale of common stock in underwritten public offering, net															269,718
Stock-based compensation expense		4,344	4,210	3,943		2,760	5,159	3,802	8,153	8,961	12,497	11,721	16,823	15,383	15,975
Unrealized gains (losses) on marketable securities		(241)									(241)		325	0	0
Foreign currency translation adjustment		(148)	(244)	42		(796)	(1,447)	(1,197)	(202)	(2,644)	(350)	(3,440)	541	(2,979)	(1,993)
Net loss	(8,813)	(6,716)	(5,551)	(7,274)	(18,198)	(36,550)	(26,906)	(17,920)	(12,825)	(44,826)	(19,541)	(81,376)	(28,354)	(99,574)	(55,548)
Ending Balance	$ 349,958	$ 352,787	$ 353,748	$ 355,194	$ 357,932	$ 371,294	$ 405,473	$ 428,456	$ 353,748	$ 405,473	$ 352,787	$ 371,294	$ 349,958	$ 357,932	$ 442,792
Common Stock
Reclassification [Line Items]
Beginning balance (in shares)	37,839	37,566	37,424	37,280	37,094	36,975	36,899	36,768	37,280	36,768	37,280	36,768	37,280	36,768	31,797
Beginning Balance	$ 38	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 32
Issuance of common stock under stock plans, including tax effects (in shares)		273	142	144		119	76	131	286	207	559	326	734	512	864
Issuance of common stock under stock plans, including tax effects		$ 1									$ 1		$ 1		$ 1
Sale of common stock in underwritten public offering, net (in shares)															4,107
Sale of common stock in underwritten public offering, net															$ 4
Ending Balance (in shares)	38,014	37,839	37,566	37,424	37,280	37,094	36,975	36,899	37,566	36,975	37,839	37,094	38,014	37,280	36,768
Ending Balance	$ 38	$ 38	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 38	$ 37	$ 38	$ 37	$ 37
Additional paid-in capital
Reclassification [Line Items]
Beginning Balance	778,615	772,472	768,123	763,629	759,254	756,087	750,717	745,936	763,629	745,936	763,629	745,936	763,629	745,936	451,433
Issuance of common stock under stock plans, including tax effects		1,799	139	551		407	211	979	690	1,190	2,489	1,597	2,690	2,310	8,814
Sale of common stock in underwritten public offering, net															269,714
Stock-based compensation expense		4,344	4,210	3,943		2,760	5,159	3,802	8,153	8,961	12,497	11,721	16,823	15,383	15,975
Ending Balance	783,142	778,615	772,472	768,123	763,629	759,254	756,087	750,717	772,472	756,087	778,615	759,254	783,142	763,629	745,936
Accumulated other comprehensive income (loss)
Reclassification [Line Items]
Beginning Balance	(3,129)	(2,740)	(2,496)	(2,538)	(2,999)	(2,203)	(756)	441	(2,538)	441	(2,538)	441	(2,538)	441	2,434
Unrealized gains (losses) on marketable securities		(241)									(241)		325
Foreign currency translation adjustment		(148)	(244)	42		(796)	(1,447)	(1,197)	(202)	(2,644)	(350)	(3,440)	541	(2,979)	(1,993)
Ending Balance	(1,672)	(3,129)	(2,740)	(2,496)	(2,538)	(2,999)	(2,203)	(756)	(2,740)	(2,203)	(3,129)	(2,999)	(1,672)	(2,538)	441
Accumulated deficit
Reclassification [Line Items]
Beginning Balance	(422,737)	(416,021)	(410,470)	(403,196)	(384,998)	(348,448)	(321,542)	(303,622)	(403,196)	(303,622)	(403,196)	(303,622)	(403,196)	(303,622)	(248,074)
Net loss		(6,716)	(5,551)	(7,274)		(36,550)	(26,906)	(17,920)	(12,825)	(44,826)	(19,541)	(81,376)	(28,354)	(99,574)	(55,548)
Ending Balance	(431,550)	(422,737)	(416,021)	(410,470)	(403,196)	(384,998)	(348,448)	(321,542)	(416,021)	(348,448)	(422,737)	(384,998)	(431,550)	(403,196)	(303,622)
As Previously Reported
Reclassification [Line Items]
Beginning Balance	353,352	355,356	357,332	358,940	372,769	405,456	426,408	440,952	358,940	440,952	358,940	440,952	358,940	440,952	206,125
Issuance of common stock under stock plans, including tax effects		1,800	139	551		407	211	979	690	1,190	2,490	1,597	2,691	2,310	8,815
Sale of common stock in underwritten public offering, net															269,718
Stock-based compensation expense		4,343	4,193	3,902		2,766	5,186	3,827	8,095	9,013	12,438	11,779	16,764	15,442	15,975
Unrealized gains (losses) on marketable securities		(241)									(241)		325
Foreign currency translation adjustment		(148)	(244)	42		(796)	(1,447)	(1,197)	(202)	(2,644)	(350)	(3,440)	541	(3,064)	(1,993)
Net loss	(12,409)	(7,758)	(6,064)	(6,103)	(18,581)	(35,064)	(24,902)	(18,153)	(12,167)	(43,055)	(19,925)	(78,119)	(32,334)	(96,700)	(57,688)
Ending Balance	$ 346,927	$ 353,352	$ 355,356	$ 357,332	$ 358,940	$ 372,769	$ 405,456	$ 426,408	$ 355,356	$ 405,456	$ 353,352	$ 372,769	$ 346,927	$ 358,940	$ 440,952
As Previously Reported | Common Stock
Reclassification [Line Items]
Beginning balance (in shares)	37,839	37,566	37,424	37,280	37,094	36,975	36,899	36,768	37,280	36,768	37,280	36,768	37,280	36,768	31,797
Beginning Balance	$ 38	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 32
Issuance of common stock under stock plans, including tax effects (in shares)		273	142	144		119	76	131	286	207	559	326	734	512	864
Issuance of common stock under stock plans, including tax effects		$ 1									$ 1		$ 1		$ 1
Sale of common stock in underwritten public offering, net (in shares)															4,107
Sale of common stock in underwritten public offering, net															$ 4
Ending Balance (in shares)	38,014	37,839	37,566	37,424	37,280	37,094	36,975	36,899	37,566	36,975	37,839	37,094	38,014	37,280	36,768
Ending Balance	$ 38	$ 38	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 38	$ 37	$ 38	$ 37	$ 37
As Previously Reported | Additional paid-in capital
Reclassification [Line Items]
Beginning Balance	778,615	772,473	768,141	763,688	759,312	756,139	750,742	745,936	763,688	745,936	763,688	745,936	763,688	745,936	451,433
Issuance of common stock under stock plans, including tax effects		1,799	139	551		407	211	979	690	1,190	2,489	1,597	2,690	2,310	8,814
Sale of common stock in underwritten public offering, net															269,714
Stock-based compensation expense		4,343	4,193	3,902		2,766	5,186	3,827	8,095	9,013	12,438	11,779	16,764	15,442	15,975
Ending Balance	783,142	778,615	772,473	768,141	763,688	759,312	756,139	750,742	772,473	756,139	778,615	759,312	783,142	763,688	745,936
As Previously Reported | Accumulated other comprehensive income (loss)
Reclassification [Line Items]
Beginning Balance	(3,214)	(2,825)	(2,581)	(2,623)	(2,999)	(2,203)	(756)	441	(2,623)	441	(2,623)	441	(2,623)	441	2,434
Unrealized gains (losses) on marketable securities		(241)									(241)		325
Foreign currency translation adjustment		(148)	(244)	42		(796)	(1,447)	(1,197)	(202)	(2,644)	(350)	(3,440)	541	(3,064)	(1,993)
Ending Balance	(1,757)	(3,214)	(2,825)	(2,581)	(2,623)	(2,999)	(2,203)	(756)	(2,825)	(2,203)	(3,214)	(2,999)	(1,757)	(2,623)	441
As Previously Reported | Accumulated deficit
Reclassification [Line Items]
Beginning Balance	(422,087)	(414,329)	(408,265)	(402,162)	(383,581)	(348,517)	(323,615)	(305,462)	(402,162)	(305,462)	(402,162)	(305,462)	(402,162)	(305,462)	(247,774)
Net loss		(7,758)	(6,064)	(6,103)		(35,064)	(24,902)	(18,153)	(12,167)	(43,055)	(19,925)	(78,119)	(32,334)	(96,700)	(57,688)
Ending Balance	(434,496)	(422,087)	(414,329)	(408,265)	(402,162)	(383,581)	(348,517)	(323,615)	(414,329)	(348,517)	(422,087)	(383,581)	(434,496)	(402,162)	(305,462)
Adjustments
Reclassification [Line Items]
Beginning Balance	(565)	(1,608)	(2,138)	(1,008)	(1,475)	17	2,048	1,840	(1,008)	1,840	(1,008)	1,840	(1,008)	1,840	(300)
Issuance of common stock under stock plans, including tax effects		0	0	0		0	0	0	0	0	0	0	0	0	0
Sale of common stock in underwritten public offering, net															0
Stock-based compensation expense		1	17	41		(6)	(27)	(25)	58	(52)	59	(58)	59	(59)	0
Unrealized gains (losses) on marketable securities		0									0		0
Foreign currency translation adjustment		0	0	0		0	0	0	0	0	0	0	0	85	0
Net loss	3,596	1,042	513	(1,171)	383	(1,486)	(2,004)	233	(658)	(1,771)	384	(3,257)	3,980	(2,874)	2,140
Ending Balance	$ 3,031	$ (565)	$ (1,608)	$ (2,138)	$ (1,008)	$ (1,475)	$ 17	$ 2,048	$ (1,608)	$ 17	$ (565)	$ (1,475)	$ 3,031	$ (1,008)	$ 1,840
Adjustments | Common Stock
Reclassification [Line Items]
Beginning balance (in shares)	0	0	0	0	0	0	0	0	0	0	0	0	0	0	0
Beginning Balance	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0
Ending Balance (in shares)	0	0	0	0	0	0	0	0	0	0	0	0	0	0	0
Ending Balance	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0
Adjustments | Additional paid-in capital
Reclassification [Line Items]
Beginning Balance	0	(1)	(18)	(59)	(58)	(52)	(25)	0	(59)	0	(59)	0	(59)	0	0
Stock-based compensation expense		1	17	41		(6)	(27)	(25)	58	(52)	59	(58)	59	(59)
Ending Balance	0	0	(1)	(18)	(59)	(58)	(52)	(25)	(1)	(52)	0	(58)	0	(59)	0
Adjustments | Accumulated other comprehensive income (loss)
Reclassification [Line Items]
Beginning Balance	85	85	85	85	0	0	0	0	85	0	85	0	85	0	0
Foreign currency translation adjustment														85
Ending Balance	85	85	85	85	85	0	0	0	85	0	85	0	85	85	0
Adjustments | Accumulated deficit
Reclassification [Line Items]
Beginning Balance	(650)	(1,692)	(2,205)	(1,034)	(1,417)	69	2,073	1,840	(1,034)	1,840	(1,034)	1,840	(1,034)	1,840	(300)
Net loss		1,042	513	(1,171)		(1,486)	(2,004)	233	(658)	(1,771)	384	(3,257)	3,980	(2,874)	2,140
Ending Balance	$ 2,946	$ (650)	$ (1,692)	$ (2,205)	$ (1,034)	$ (1,417)	$ 69	$ 2,073	$ (1,692)	$ 69	$ (650)	$ (1,417)	$ 2,946	$ (1,034)	$ 1,840

Restatement of Financial Stat_8

Restatement of Financial Statements - Statement Of Cash Flows (Details) - USD ($) $ in Thousands	3 Months Ended		6 Months Ended		9 Months Ended		12 Months Ended
	Mar. 31, 2023	Mar. 31, 2022	Jun. 30, 2023	Jun. 30, 2022	Sep. 30, 2023	Sep. 30, 2022	Dec. 31, 2023	Dec. 31, 2022	Dec. 31, 2021
Cash flows from operating activities:
Net loss	$ (7,274)	$ (17,920)	$ (12,825)	$ (44,826)	$ (19,541)	$ (81,376)	$ (28,354)	$ (99,574)	$ (55,548)
Adjustments to reconcile net loss to net cash used in operating activities:
Depreciation and amortization expense	1,454	1,373	2,883	2,820	4,843	4,231	6,275	5,409	4,880
Credit losses on accounts receivable	178	(171)	324	581	311	102	336	(301)	213
Operating lease right-of-use asset amortization			1,002		1,518		2,015	715	499
Stock-based compensation expense	3,943	3,802	8,153	8,961	12,497	11,721	16,823	15,383	15,975
Impairment			54	207	130		1,361	25,801	0
Other operating activity			602				(150)	(340)	692
Changes in assets and liabilities:
Accounts receivable	(3,741)	1,319	(5,750)	3,479	(5,615)	5,045	(6,695)	5,156	(6,853)
Inventory	530	(206)	(1,180)	1,515	(3,700)	6,146	(8,944)	7,243	(10,125)
Prepaid expenses and other current assets	(422)	(2,071)	(527)	961	(2,829)	(282)	(2,371)	(738)	(166)
Other non-current assets	(33)	1	(707)	2	(702)	(857)	(717)	(822)	(2)
Accounts payable	(1,271)	(5,306)	(631)	(3,220)	948	(7,010)	1,189	(5,287)	2,339
Accrued compensation and benefits, accrued expenses, and other current liabilities	(5,809)	(4,864)	(1,053)	(4,248)	1,229	(2,914)	4,410	(3,599)	606
Deferred revenue	2,041	2,956	1,666	6,228	894	3,108	635	2,599	1,462
Operating lease liabilities	179	(87)	(730)	(3,128)	(1,690)	(1,156)	(2,645)	(266)	(1,230)
Other non-current liabilities	(203)	(271)	(72)	(9)	(107)	128	(53)	10	(363)
Net cash used in operating activities	(9,824)	(21,755)	(8,791)	(29,716)	(12,657)	(44,521)	(18,849)	(48,611)	(47,621)
Cash flows from investing activities:
Purchases of marketable debt securities					(125,200)		(175,613)	0	0
Proceeds from marketable debt securities							31,000	0	0
Purchases of property and equipment	(136)	(1,335)	(947)	(6,342)	(1,878)	(10,019)	(3,841)	(11,614)	(13,675)
Proceeds from RADx grant on assets purchased		520		520		520	0	520	7,278
Net cash used in investing activities	(136)	(815)	(947)	(5,822)	(127,078)	(9,499)	(148,454)	(11,094)	(6,397)
Cash flows from financing activities:
Proceeds from common stock issued under stock plans	551	594	777	594	2,632	881	2,889	2,311	8,815
Payments on notes payable							0	0	(7,738)
Net cash provided by financing activities	551	979	690	1,190	2,490	1,597	2,691	2,311	270,795
Net increase (decrease) in cash, cash equivalents, and restricted cash	(9,409)	(21,591)	(9,048)	(34,348)	(137,245)	(52,423)	(164,612)	(57,394)	216,777
Effect of exchange rate changes on cash, cash equivalents, and restricted cash	24	(558)	(163)	(776)	(229)	(507)	301	(538)	(92)
Cash, cash equivalents, and restricted cash at beginning of period	341,337	399,269	341,337	399,269	341,337	399,269	341,337	399,269	182,584
Cash, cash equivalents, and restricted cash at end of period	331,952	377,120	332,126	364,145	203,863	346,339	177,026	341,337	399,269
Supplemental disclosure of cash flow information:
Cash paid for taxes	246		502		719	263	808	684	0
Purchases of property and equipment in accounts payable and accruals	147					288	419	152	229
Cash paid for interest							0	0	389
Operating lease right-of-use assets obtained in exchange for lease liabilities		18,156				22,239	0	22,494	0
Shares received as consideration under product sales agreement (Note 4, 7)							775	0	0
As Previously Reported
Cash flows from operating activities:
Net loss	(6,103)	(18,153)	(12,167)	(43,055)	(19,925)	(78,119)	(32,334)	(96,700)	(57,688)
Adjustments to reconcile net loss to net cash used in operating activities:
Depreciation and amortization expense	1,439	1,358	2,845	2,790	4,788	4,186	6,364	5,349	4,851
Credit losses on accounts receivable	178	(171)	324	581	311	102	336	(301)	213
Operating lease right-of-use asset amortization			1,002		1,518		2,015	715	499
Stock-based compensation expense	3,902	3,827	8,095	9,013	12,438	11,779	16,764	15,442	15,975
Impairment			0	0	0		1,570	25,592
Other operating activity			548				150	(439)	692
Changes in assets and liabilities:
Accounts receivable	(3,741)	1,319	(5,750)	3,479	(5,615)	5,045	(6,695)	5,156	(6,853)
Inventory	(89)	(484)	(1,181)	180	(2,966)	3,919	(5,364)	5,386	(8,090)
Prepaid expenses and other current assets	(422)	(2,070)	(527)	930	(2,829)	(262)	(2,371)	(568)	(393)
Other non-current assets	(33)	1	(965)	2	(716)	(859)	(775)	(909)	(2)
Accounts payable	(1,271)	(5,306)	(631)	(3,220)	948	(7,085)	1,189	(5,362)	2,414
Accrued compensation and benefits, accrued expenses, and other current liabilities	(5,983)	(4,921)	(1,326)	(4,387)	876	(3,021)	4,276	(3,976)	606
Deferred revenue	2,041	2,956	1,666	6,228	894	3,108	635	2,599	1,462
Operating lease liabilities	179	(87)	(730)	(3,128)	(1,690)	(1,156)	(2,645)	(266)	(1,230)
Other non-current liabilities	(203)	(271)	(72)	(9)	(107)	128	(53)	10	(363)
Net cash used in operating activities	(9,502)	(21,695)	(8,869)	(29,635)	(12,918)	(44,182)	(18,902)	(48,272)	(47,907)
Cash flows from investing activities:
Purchases of marketable debt securities					(125,200)		(175,613)		0
Proceeds from marketable debt securities							31,000		0
Purchases of property and equipment	(136)	(1,394)	(784)	(6,454)	(1,572)	(10,131)	(3,788)	(11,726)	(13,616)
Proceeds from RADx grant on assets purchased		520		520		520		520	7,278
Net cash used in investing activities	(136)	(874)	(784)	(5,934)	(126,772)	(9,611)	(148,401)	(11,206)	(6,338)
Cash flows from financing activities:
Proceeds from common stock issued under stock plans	551	594	777	594	2,632	881	2,889	2,311	8,815
Payments on notes payable									(7,738)
Net cash provided by financing activities	551	979	690	1,190	2,490	1,597	2,691	2,311	270,795
Net increase (decrease) in cash, cash equivalents, and restricted cash	(9,087)	(21,590)	(8,963)	(34,379)	(137,200)	(52,196)	(164,612)	(57,167)	216,550
Effect of exchange rate changes on cash, cash equivalents, and restricted cash	24	(558)	(163)	(776)	(229)	(507)	301	(538)	(92)
Cash, cash equivalents, and restricted cash at beginning of period	341,337	399,042	341,337	399,042	341,337	399,042	341,337	399,042	182,584
Cash, cash equivalents, and restricted cash at end of period	332,274	376,894	332,211	363,887	203,908	346,339	177,026	341,337	399,042
Supplemental disclosure of cash flow information:
Cash paid for taxes	246		502		719	263	808	684
Purchases of property and equipment in accounts payable and accruals	147					198	419	152	229
Cash paid for interest									389
Operating lease right-of-use assets obtained in exchange for lease liabilities		18,156				22,239		22,494
Shares received as consideration under product sales agreement (Note 4, 7)							775
Adjustments
Cash flows from operating activities:
Net loss	(1,171)	233	(658)	(1,771)	384	(3,257)	3,980	(2,874)	2,140
Adjustments to reconcile net loss to net cash used in operating activities:
Depreciation and amortization expense	15	15	38	30	55	45	(89)	60	29
Credit losses on accounts receivable	0	0	0	0	0	0	0	0	0
Operating lease right-of-use asset amortization			0		0		0	0	0
Stock-based compensation expense	41	(25)	58	(52)	59	(58)	59	(59)	0
Impairment			54	207	130		(209)	209
Other operating activity			54				(300)	99	0
Changes in assets and liabilities:
Accounts receivable	0	0	0	0	0	0	0	0	0
Inventory	619	278	1	1,335	(734)	2,227	(3,580)	1,857	(2,035)
Prepaid expenses and other current assets	0	(1)	0	31	0	(20)	0	(170)	227
Other non-current assets	0	0	258	0	14	2	58	87	0
Accounts payable	0	0	0	0	0	75	0	75	(75)
Accrued compensation and benefits, accrued expenses, and other current liabilities	174	57	273	139	353	107	134	377	0
Deferred revenue	0	0	0	0	0	0	0	0	0
Operating lease liabilities	0	0	0	0	0	0	0	0	0
Other non-current liabilities	0	0	0	0	0	0	0	0	0
Net cash used in operating activities	(322)	(60)	78	(81)	261	(339)	53	(339)	286
Cash flows from investing activities:
Purchases of marketable debt securities					0		0		0
Proceeds from marketable debt securities							0		0
Purchases of property and equipment	0	59	(163)	112	(306)	112	(53)	112	(59)
Proceeds from RADx grant on assets purchased		0		0		0		0	0
Net cash used in investing activities	0	59	(163)	112	(306)	112	(53)	112	(59)
Cash flows from financing activities:
Proceeds from common stock issued under stock plans	0	0	0	0	0	0	0	0	0
Payments on notes payable									0
Net cash provided by financing activities	0	0	0	0	0	0	0	0	0
Net increase (decrease) in cash, cash equivalents, and restricted cash	(322)	(1)	(85)	31	(45)	(227)	0	(227)	227
Effect of exchange rate changes on cash, cash equivalents, and restricted cash	0	0	0	0	0	0	0	0	0
Cash, cash equivalents, and restricted cash at beginning of period	0	227	0	227	0	227	0	227	0
Cash, cash equivalents, and restricted cash at end of period	(322)	226	(85)	$ 258	(45)	0	0	0	227
Supplemental disclosure of cash flow information:
Cash paid for taxes	0		$ 0		$ 0	0	0	0
Purchases of property and equipment in accounts payable and accruals	$ 0					90	0	0	0
Cash paid for interest									0
Operating lease right-of-use assets obtained in exchange for lease liabilities		$ 0				$ 0		0
Shares received as consideration under product sales agreement (Note 4, 7)							0
Underwritten public offering
Cash flows from financing activities:
Sale of common stock in underwritten public offering, net							$ 0	$ 0	269,718
Underwritten public offering | As Previously Reported
Cash flows from financing activities:
Sale of common stock in underwritten public offering, net									269,718
Underwritten public offering | Adjustments
Cash flows from financing activities:
Sale of common stock in underwritten public offering, net									$ 0

Organization and Nature of Bu_2

Organization and Nature of Business (Details) $ in Thousands	3 Months Ended								6 Months Ended		9 Months Ended		12 Months Ended
	Dec. 31, 2023 USD ($) project customer	Sep. 30, 2023 USD ($)	Jun. 30, 2023 USD ($)	Mar. 31, 2023 USD ($)	Dec. 31, 2022 USD ($)	Sep. 30, 2022 USD ($)	Jun. 30, 2022 USD ($)	Mar. 31, 2022 USD ($)	Jun. 30, 2023 USD ($)	Jun. 30, 2022 USD ($)	Sep. 30, 2023 USD ($)	Sep. 30, 2022 USD ($)	Dec. 31, 2023 USD ($) project customer	Dec. 31, 2022 USD ($)	Dec. 31, 2021 USD ($)
Organization and Nature of Business
Number of projects completed | project	2,200												2,200
Net loss	$ (8,813)	$ (6,716)	$ (5,551)	$ (7,274)	$ (18,198)	$ (36,550)	$ (26,906)	$ (17,920)	$ (12,825)	$ (44,826)	$ (19,541)	$ (81,376)	$ (28,354)	$ (99,574)	$ (55,548)
Accumulated deficit	(431,550)	(422,737)	(416,021)	(410,470)	(403,196)	(384,998)	(348,448)	(321,542)	(416,021)	(348,448)	(422,737)	(384,998)	(431,550)	(403,196)
Cash and cash equivalents	174,422	201,261	$ 329,525	$ 329,354	338,740	$ 343,743	$ 361,551	$ 374,543	$ 329,525	$ 361,551	201,261	$ 343,743	174,422	338,740
Marketable securities	$ 146,902	$ 126,449			$ 0						$ 126,449		$ 146,902	$ 0
Minimum
Organization and Nature of Business
Number of customers served | customer	480												480

Significant Accounting Polici_4

Significant Accounting Policies - Foreign Currency (Details) - USD ($) $ in Millions	12 Months Ended
	Dec. 31, 2023	Dec. 31, 2022	Dec. 31, 2021
Accounting Policies [Abstract]
Foreign exchange losses	$ (0.2)	$ (1)	$ (0.4)

Significant Accounting Polici_5

Significant Accounting Policies - Contract Assets and Liabilities (Details)	12 Months Ended
	Dec. 31, 2023
Minimum
Service Revenue
Period of payment	30 days
Maximum
Service Revenue
Period of payment	45 days

Significant Accounting Polici_6

Significant Accounting Policies - Research and Development Expenses (Details) - USD ($) $ in Millions	12 Months Ended	24 Months Ended
	Dec. 31, 2023	Dec. 31, 2022
Accounting Policies [Abstract]
Reimbursed amount	$ 0.4	$ 0

Significant Accounting Polici_7

Significant Accounting Policies - Cash and Cash Equivalents (Details) - USD ($) $ in Thousands	Dec. 31, 2023	Sep. 30, 2023	Jun. 30, 2023	Mar. 31, 2023	Dec. 31, 2022	Sep. 30, 2022	Jun. 30, 2022	Mar. 31, 2022
Accounting Policies [Abstract]
Cash	$ 12,162				$ 32,643
Money market funds	155,367				306,097
Marketable securities	6,893				0
Total cash and cash equivalents	$ 174,422	$ 201,261	$ 329,525	$ 329,354	$ 338,740	$ 343,743	$ 361,551	$ 374,543

Significant Accounting Polici_8

Significant Accounting Policies - Restricted Cash (Details) - USD ($) $ in Thousands	Dec. 31, 2023	Sep. 30, 2023	Jun. 30, 2023	Mar. 31, 2023	Dec. 31, 2022	Sep. 30, 2022	Jun. 30, 2022	Mar. 31, 2022	Dec. 31, 2021	Dec. 31, 2020
Cash and Cash equivalents
Cash and cash equivalents	$ 174,422	$ 201,261	$ 329,525	$ 329,354	$ 338,740	$ 343,743	$ 361,551	$ 374,543
Restricted cash	2,604				2,597
Cash, cash equivalents, and restricted cash	$ 177,026	$ 203,863	$ 332,126	$ 331,952	$ 341,337	$ 346,339	$ 364,145	$ 377,120	$ 399,269	$ 182,584

Significant accounting polici_9

Significant accounting policies - Property and Equipment (Details)	Dec. 31, 2023
Laboratory and manufacturing equipment
Property and equipment
Property, Plant and Equipment, Useful Life	5 years
Office furniture and equipment
Property and equipment
Property, Plant and Equipment, Useful Life	7 years
Computers and software
Property and equipment
Property, Plant and Equipment, Useful Life	3 years

Significant accounting polic_10

Significant accounting policies - Credit, product and supplier concentrations and off-balance-sheet risk (Details) $ in Thousands	3 Months Ended								6 Months Ended		9 Months Ended		12 Months Ended
	Dec. 31, 2023 USD ($)	Sep. 30, 2023 USD ($)	Jun. 30, 2023 USD ($)	Mar. 31, 2023 USD ($)	Dec. 31, 2022 USD ($)	Sep. 30, 2022 USD ($)	Jun. 30, 2022 USD ($)	Mar. 31, 2022 USD ($)	Jun. 30, 2023 USD ($)	Jun. 30, 2022 USD ($)	Sep. 30, 2023 USD ($)	Sep. 30, 2022 USD ($)	Dec. 31, 2023 USD ($) customer	Dec. 31, 2022 USD ($) customer	Dec. 31, 2021 USD ($) customer
Concentration Risk [Line Items]
Revenue | $	$ 31,549	$ 31,559	$ 30,804	$ 28,456	$ 25,824	$ 26,646	$ 23,500	$ 29,552	$ 59,260	$ 53,052	$ 90,819	$ 79,698	$ 122,368	$ 105,522	$ 110,556
Accounts Receivable [Member] | Customer Concentration Risk [Member] | Customers outside the United States
Concentration Risk [Line Items]
Concentration risk (in percent)													38%	41%
Revenue Benchmark [Member] | Customer Concentration Risk [Member] | One customer
Concentration Risk [Line Items]
Concentration risk (in percent)													10%	10%
Revenue | $													$ 14,000	$ 13,700
Number Of Customers With High Percent Of Total Revenue | customer													1	1
Revenue Benchmark [Member] | Customer Concentration Risk [Member] | Customer
Concentration Risk [Line Items]
Concentration risk (in percent)															10%
Number Of Customers With High Percent Of Total Revenue | customer															0

Significant accounting polic_11

Significant accounting policies - Stock-Based Compensation (Details)	12 Months Ended
	Dec. 31, 2023
Accounting Policies [Abstract]
Expected dividend yield (as a percent)	0%

Significant accounting polic_12

Significant accounting policies - Advertising (Details) - USD ($) $ in Millions	12 Months Ended
	Dec. 31, 2023	Dec. 31, 2022	Dec. 31, 2021
Accounting Policies [Abstract]
Advertising Expense	$ 0.3	$ 0.3	$ 0.5

Revenue and Related Matters - D

Revenue and Related Matters - Disaggregated revenue (Details) - USD ($) $ in Thousands	3 Months Ended								6 Months Ended		9 Months Ended		12 Months Ended
	Dec. 31, 2023	Sep. 30, 2023	Jun. 30, 2023	Mar. 31, 2023	Dec. 31, 2022	Sep. 30, 2022	Jun. 30, 2022	Mar. 31, 2022	Jun. 30, 2023	Jun. 30, 2022	Sep. 30, 2023	Sep. 30, 2022	Dec. 31, 2023	Dec. 31, 2022	Dec. 31, 2021
Service Revenue
Total revenues	$ 31,549	$ 31,559	$ 30,804	$ 28,456	$ 25,824	$ 26,646	$ 23,500	$ 29,552	$ 59,260	$ 53,052	$ 90,819	$ 79,698	$ 122,368	$ 105,522	$ 110,556
Product revenue
Service Revenue
Total revenues	20,821	19,940	19,549	19,360	16,674	17,693	14,785	20,656	38,909	35,441	58,849	53,134	79,670	69,808	81,062
Product revenue | North America
Service Revenue
Total revenues													41,496	35,148	47,135
Product revenue | EMEA
Service Revenue
Total revenues													25,600	22,876	24,300
Product revenue | Asia Pacific
Service Revenue
Total revenues													12,574	11,784	9,627
Instruments
Service Revenue
Total revenues													15,705	25,004	25,973
Instruments | North America
Service Revenue
Total revenues													6,374	9,254	12,138
Instruments | EMEA
Service Revenue
Total revenues													4,384	8,362	8,178
Instruments | Asia Pacific
Service Revenue
Total revenues													4,947	7,388	5,657
Consumable and other products
Service Revenue
Total revenues													63,965	44,804	55,089
Consumable and other products | North America
Service Revenue
Total revenues													35,122	25,894	34,997
Consumable and other products | EMEA
Service Revenue
Total revenues													21,216	14,514	16,122
Consumable and other products | Asia Pacific
Service Revenue
Total revenues													7,627	4,396	3,970
Service revenue
Service Revenue
Total revenues														34,495
Service revenue | North America
Service Revenue
Total revenues														29,218
Service revenue | EMEA
Service Revenue
Total revenues														4,514
Service revenue | Asia Pacific
Service Revenue
Total revenues														763
Research services
Service Revenue
Total revenues													27,828	23,653	14,825
Research services | North America
Service Revenue
Total revenues													24,706	22,493	12,101
Research services | EMEA
Service Revenue
Total revenues													2,000	1,013	2,600
Research services | Asia Pacific
Service Revenue
Total revenues													1,122	147	124
Other services
Service Revenue
Total revenues													2,382	2,002	2,176
Other services | North America
Service Revenue
Total revenues													1,436	1,144	1,372
Other services | EMEA
Service Revenue
Total revenues													951	722	695
Other services | Asia Pacific
Service Revenue
Total revenues													(5)	136	109
Service and other revenue
Service Revenue
Total revenues	10,230	10,883	10,470	8,506	8,767	8,370	8,548	8,810	18,976	17,358	29,859	25,728	40,089	34,495	23,629
Service and other revenue | North America
Service Revenue
Total revenues													32,407		17,807
Service and other revenue | EMEA
Service Revenue
Total revenues													5,952		5,334
Service and other revenue | Asia Pacific
Service Revenue
Total revenues													1,730		488
Service-type warranties
Service Revenue
Total revenues													9,879	8,840	6,628
Service-type warranties | North America
Service Revenue
Total revenues													6,265	5,581	4,334
Service-type warranties | EMEA
Service Revenue
Total revenues													3,001	2,779	2,039
Service-type warranties | Asia Pacific
Service Revenue
Total revenues													613	480	255
Collaboration and license revenue
Service Revenue
Total revenues	146	237	629	368	170	301	92	$ 86	997	178	1,234	479	1,380	649	648
Collaboration and license revenue | North America
Service Revenue
Total revenues													1,380	274	360
Collaboration and license revenue | EMEA
Service Revenue
Total revenues													0	323	288
Collaboration and license revenue | Asia Pacific
Service Revenue
Total revenues													0	52	0
Grant revenue
Service Revenue
Total revenues	$ 352	$ 499	$ 156	$ 222	$ 213	$ 282	$ 75		$ 378	$ 75	$ 877	$ 357	$ 1,229	$ 570	$ 5,217

Revenue and Related Matters - S

Revenue and Related Matters - Service Revenue (Details) $ in Thousands

3 Months Ended

6 Months Ended

9 Months Ended

12 Months Ended

Dec. 31, 2023 USD ($)

Sep. 30, 2023 USD ($)

Jun. 30, 2023 USD ($)

Mar. 31, 2023 USD ($)

Dec. 31, 2022 USD ($)

Sep. 30, 2022 USD ($)

Jun. 30, 2022 USD ($)

Mar. 31, 2022 USD ($)

Jun. 30, 2023 USD ($)

Jun. 30, 2022 USD ($)

Sep. 30, 2023 USD ($)

Sep. 30, 2022 USD ($)

Dec. 31, 2023 USD ($) customer

Dec. 31, 2022 USD ($) customer

Dec. 31, 2021 USD ($)

Service Revenue

Revenue

$ 31,549

$ 31,559

$ 30,804

$ 28,456

$ 25,824

$ 26,646

$ 23,500

$ 29,552

$ 59,260

$ 53,052

$ 90,819

$ 79,698

$ 122,368

$ 105,522

$ 110,556

One customer | Customer Concentration Risk [Member] | Revenue Benchmark [Member]

Service Revenue

Revenue

$ 14,000

$ 13,700

Number Of Customers With High Percent Of Total Revenue | customer

1

1

Revenue as a percentage of total revenue

10%

10%

Service revenue

Service Revenue

Revenue

$ 34,495

Collaboration agreement | Eli Lilly

Service Revenue

Revenue

$ 6,000

$ 10,900

Master collaboration agreement | Eli Lilly

Service Revenue

Non-refundable up-front payment received

5,000

Statement of works agreement | Eli Lilly

Service Revenue

Collaborative arrangement payment received per quarter

$ 1,500

Minimum

Service Revenue

Period of payment

30 days

Maximum

Service Revenue

Period of payment

45 days

Revenue and Related Matters - C

Revenue and Related Matters - Collaboration and license arrangements (Details) - USD ($) $ in Thousands, shares in Millions	3 Months Ended								6 Months Ended		9 Months Ended		12 Months Ended
	Dec. 31, 2023	Sep. 30, 2023	Jun. 30, 2023	Mar. 31, 2023	Dec. 31, 2022	Sep. 30, 2022	Jun. 30, 2022	Mar. 31, 2022	Jun. 30, 2023	Jun. 30, 2022	Sep. 30, 2023	Sep. 30, 2022	Dec. 31, 2023	Dec. 31, 2022	Dec. 31, 2021
Collaboration and license arrangements
Revenue	$ 31,549	$ 31,559	$ 30,804	$ 28,456	$ 25,824	$ 26,646	$ 23,500	$ 29,552	$ 59,260	$ 53,052	$ 90,819	$ 79,698	$ 122,368	$ 105,522	$ 110,556
UltraDx
Collaboration and license arrangements
Revenue													1,800	1,900
Consideration on collaboration activities						1,900
Shares amount received						1,000						1,000
Number of contingent consideration collaboration			1
Collaboration and license revenue
Collaboration and license arrangements
Revenue	$ 146	$ 237	$ 629	$ 368	$ 170	$ 301	$ 92	$ 86	$ 997	$ 178	$ 1,234	$ 479	$ 1,380	$ 649	$ 648

Revenue and Related Matters - G

Revenue and Related Matters - Grant revenue (Details) - USD ($) $ in Thousands				3 Months Ended								6 Months Ended		9 Months Ended		12 Months Ended			24 Months Ended
	Sep. 21, 2022	Mar. 24, 2022	Sep. 29, 2020	Dec. 31, 2023	Sep. 30, 2023	Jun. 30, 2023	Mar. 31, 2023	Dec. 31, 2022	Sep. 30, 2022	Jun. 30, 2022	Mar. 31, 2022	Jun. 30, 2023	Jun. 30, 2022	Sep. 30, 2023	Sep. 30, 2022	Dec. 31, 2023	Dec. 31, 2022	Dec. 31, 2021	Dec. 31, 2023	Jun. 30, 2020
Total revenues				$ 31,549	$ 31,559	$ 30,804	$ 28,456	$ 25,824	$ 26,646	$ 23,500	$ 29,552	$ 59,260	$ 53,052	$ 90,819	$ 79,698	$ 122,368	$ 105,522	$ 110,556
Research and development				7,210	7,739	6,128	4,987	5,843	6,829	7,257	6,880	11,115	14,137	18,854	20,966	26,064	26,809	28,383
Deferred revenue							(2,041)				$ (2,956)	(1,666)	(6,228)	(894)	(3,108)	(635)	(2,599)	(1,462)
Commitments and contingencies (Note 16)
RADx WP2
Contract value			$ 18,200
Grants Received																				$ 500
Research and development																		3,400
Commitments and contingencies (Note 16)				0												0			0
ADDF
Contract value		$ 2,300
Grants Received				2,300												2,300			2,300
Research and development																500	600
National Institutes of Health
Contract value	$ 1,700
Grants Received				600												600			600
Research and development																600
Grant revenue
Total revenues				$ 352	$ 499	$ 156	$ 222	$ 213	$ 282	$ 75		$ 378	$ 75	$ 877	$ 357	1,229	570	5,217
Grant revenue | RADx WP2
Total revenues																		$ 5,200	$ 0
Grant revenue | ADDF
Total revenues																500	$ 600
Deferred revenue																1,100
Grant revenue | National Institutes of Health
Total revenues																$ 700

Revenue and Related Matters -_2

Revenue and Related Matters - Contract Assets and Liabilities (Details) $ in Millions	Dec. 31, 2023 USD ($)
Revenue from Contract with Customer [Abstract]
Contract with Customer, Asset, after Allowance for Credit Loss	$ 0

Revenue and Related Matters -_3

Revenue and Related Matters - Deferred Revenue (Details) - USD ($) $ in Millions	12 Months Ended
	Dec. 31, 2023	Dec. 31, 2022
Revenue from Contract with Customer [Abstract]
Deferred Revenue, Revenue Recognized	$ 7.7	$ 5.5

Revenue and Related Matters - R

Revenue and Related Matters - Remaining Performance Obligations (Details) $ in Millions	Dec. 31, 2023 USD ($)
Transaction Price Allocated to Remaining Performance Obligations
Amount of transaction price allocated to performance obligations	$ 10.7
Undelivered licenses of intellectual property
Transaction Price Allocated to Remaining Performance Obligations
Amount of transaction price allocated to performance obligations	10.7
Revenue, Remaining Performance Obligation, Expected Timing of Satisfaction, Start Date [Axis]: 2024-01-01
Transaction Price Allocated to Remaining Performance Obligations
Amount of transaction price allocated to performance obligations	$ 9.5
Performance obligation satisfaction period	12 months

Revenue and Related Matters -_4

Revenue and Related Matters - Costs to obtain a contract (Details) - USD ($) $ in Thousands	12 Months Ended			36 Months Ended
	Dec. 31, 2023	Dec. 31, 2022	Dec. 31, 2021	Dec. 31, 2023
Change in the balance of costs to obtain a contract
Balance at beginning of period	$ 377	$ 440	$ 248	$ 248
Capitalization of costs to obtain a contract	528	1,387	905
Recognition of costs to obtain a contract	(617)	(1,450)	(713)
Balance at end of period	$ 288	$ 377	$ 440	288
Impairment loss				$ 0

Allowance for Credit Losses (De

Allowance for Credit Losses (Details) - USD ($) $ in Thousands	12 Months Ended
	Dec. 31, 2023	Dec. 31, 2022	Dec. 31, 2021
Allowance for Credit Loss [Abstract]
Beginning Balance	$ 118	$ 419	$ 370
Provision for expected credit losses	(729)	(752)	(213)
Write-offs and recoveries collected	(393)	(1,053)	(164)
Ending Balance	$ 454	$ 118	$ 419

Marketable Securities (Details)

Marketable Securities (Details) - USD ($) $ in Thousands	Dec. 31, 2023	Sep. 30, 2023	Dec. 31, 2022
Marketable Securities [Line Items]
Amortized cost	$ 153,470
Unrealized Gains	352
Unrealized Losses	(27)
Fair Value	153,795
Marketable securities	146,902	$ 126,449	$ 0
Debt Securities, Available-for-Sale, Continuous Unrealized Loss Position, Less than 12 Months	56,365
Unrealized Loses	(27)
Amortized cost due within one year	95,188
Fair value due within one year	95,232
Amortized cost due in one to two years	58,282
Fair value due in one to two years	58,563
Accrued interest receivable	1,000
Commercial paper
Marketable Securities [Line Items]
Amortized cost	53,482
Unrealized Gains	23
Unrealized Losses	(12)
Fair Value	53,493
Debt Securities, Available-for-Sale, Continuous Unrealized Loss Position, Less than 12 Months	32,137
Unrealized Loses	(12)
U.S. Treasuries
Marketable Securities [Line Items]
Amortized cost	4,896
Unrealized Gains	1
Fair Value	4,897
Marketable securities	146,902
U.S. Government agencies
Marketable Securities [Line Items]
Amortized cost	28,366
Unrealized Gains	39
Unrealized Losses	(7)
Fair Value	28,398
Debt Securities, Available-for-Sale, Continuous Unrealized Loss Position, Less than 12 Months	15,861
Unrealized Loses	(7)
Corporate bonds
Marketable Securities [Line Items]
Amortized cost	66,726
Unrealized Gains	289
Unrealized Losses	(8)
Fair Value	67,007
Debt Securities, Available-for-Sale, Continuous Unrealized Loss Position, Less than 12 Months	8,367
Unrealized Loses	(8)
Cash and cash equivalents
Marketable Securities [Line Items]
Marketable securities	$ 6,893

Fair value of Financial Instr_3

Fair value of Financial Instruments (As Restated) (Details) - USD ($) $ in Thousands	Dec. 31, 2023	Sep. 30, 2023	Dec. 31, 2022
Financial Instruments, Financial Assets, Balance Sheet Groupings [Abstract]
Cash equivalents - money market funds	$ 162,260
Marketable securities	146,902	$ 126,449	$ 0
Total financial assets	309,162		306,097
Money market funds
Financial Instruments, Financial Assets, Balance Sheet Groupings [Abstract]
Cash equivalents - money market funds	155,367
Marketable securities			306,097
Commercial paper
Financial Instruments, Financial Assets, Balance Sheet Groupings [Abstract]
Cash equivalents - money market funds	1,996
Marketable securities	51,498
U.S. Treasuries
Financial Instruments, Financial Assets, Balance Sheet Groupings [Abstract]
Cash equivalents - money market funds	4,897
U.S. Government agencies
Financial Instruments, Financial Assets, Balance Sheet Groupings [Abstract]
Marketable securities	28,398
Corporate bonds
Financial Instruments, Financial Assets, Balance Sheet Groupings [Abstract]
Marketable securities	67,006
Level 1
Financial Instruments, Financial Assets, Balance Sheet Groupings [Abstract]
Cash equivalents - money market funds	155,367
Total financial assets	155,367		306,097
Level 1 | Money market funds
Financial Instruments, Financial Assets, Balance Sheet Groupings [Abstract]
Cash equivalents - money market funds	155,367
Marketable securities			$ 306,097
Level 2
Financial Instruments, Financial Assets, Balance Sheet Groupings [Abstract]
Cash equivalents - money market funds	6,893
Marketable securities	146,902
Total financial assets	153,795
Level 2 | Commercial paper
Financial Instruments, Financial Assets, Balance Sheet Groupings [Abstract]
Cash equivalents - money market funds	1,996
Marketable securities	51,498
Level 2 | U.S. Treasuries
Financial Instruments, Financial Assets, Balance Sheet Groupings [Abstract]
Cash equivalents - money market funds	4,897
Level 2 | U.S. Government agencies
Financial Instruments, Financial Assets, Balance Sheet Groupings [Abstract]
Marketable securities	28,398
Level 2 | Corporate bonds
Financial Instruments, Financial Assets, Balance Sheet Groupings [Abstract]
Marketable securities	$ 67,006

Fair value of Financial Instr_4

Fair value of Financial Instruments (As Restated) - Changes in Level 3 Financial Instruments (Details) $ in Millions	Jun. 26, 2023 USD ($)
Changes in the Company's Level 3 financial instruments
Receipt of ordinary shares of UltraDx (Note 13)	$ 0.8

Fair value of Financial Instr_5

Fair value of Financial Instruments (As Restated) - Changes in Carrying Value (Details) - USD ($) $ in Thousands	Dec. 31, 2023	Dec. 31, 2022
Fair Value, Assets and Liabilities Measured on Recurring and Nonrecurring Basis [Line Items]
Carrying value of the Company's Level 3 financial assets	$ 309,162	$ 306,097
Nonrecurring | Fair Value, Inputs, Level 3
Fair Value, Assets and Liabilities Measured on Recurring and Nonrecurring Basis [Line Items]
Carrying value of the Company's Level 3 financial assets	$ 800

Inventory (Details)

Inventory (Details) - USD ($) $ in Thousands	Dec. 31, 2023	Sep. 30, 2023	Jun. 30, 2023	Mar. 31, 2023	Dec. 31, 2022	Sep. 30, 2022	Jun. 30, 2022	Mar. 31, 2022
Inventory Disclosure [Abstract]
Raw Materials	$ 5,114				$ 5,509
Work in process	5,439				3,415
Finished goods	15,570				8,040
Total inventory	$ 26,123	$ 20,857	$ 18,333	$ 16,629	$ 16,964	$ 18,044	$ 22,685	$ 24,426

Property and Equipment (Details

Property and Equipment (Details) - USD ($) $ in Thousands	3 Months Ended							6 Months Ended		9 Months Ended		12 Months Ended			24 Months Ended	36 Months Ended
	Dec. 31, 2023	Sep. 30, 2023	Jun. 30, 2023	Mar. 31, 2023	Dec. 31, 2022	Sep. 30, 2022	Jun. 30, 2022	Jun. 30, 2023	Jun. 30, 2022	Sep. 30, 2023	Sep. 30, 2022	Dec. 31, 2023	Dec. 31, 2022	Dec. 31, 2021	Dec. 31, 2022	Dec. 31, 2023	Mar. 31, 2022
Property and equipment
Property and equipment, gross	$ 32,047				$ 30,820							$ 32,047	$ 30,820		$ 30,820	$ 32,047
Less: accumulated depreciation	(14,121)				(11,009)							(14,121)	(11,009)		(11,009)	(14,121)
Total	17,926	$ 17,287	$ 18,048	$ 18,690	19,811	$ 21,195	$ 21,976	$ 18,048	$ 21,976	$ 17,287	$ 21,195	17,926	19,811		$ 19,811	17,926	$ 19,639
Depreciation expense												$ 4,700	3,500	$ 2,800
Disposals related to equipment no longer in use												0.8 million			no
Impairment and restructuring	1,361	$ 0	$ 0	$ (33)	9,006	$ 20,343	$ 207	$ (33)	$ 207	$ (33)	$ 20,550	$ 1,328	29,556	$ 0
Capitalized software development costs												1,100			$ 0
Laboratory and manufacturing equipment
Property and equipment
Property and equipment, gross	13,141				11,753							13,141	11,753		11,753	13,141
Office furniture and equipment.
Property and equipment
Property and equipment, gross	1,905				1,798							1,905	1,798		1,798	1,905
Computers and software
Property and equipment
Property and equipment, gross	3,927				3,581							3,927	3,581		3,581	3,927
Depreciation expense																0
Leasehold Improvements [Member]
Property and equipment
Property and equipment, gross	$ 13,074				$ 13,688							$ 13,074	$ 13,688		$ 13,688	$ 13,074

Accrued Expenses and Other Cu_3

Accrued Expenses and Other Current Liabilities (Details) - USD ($) $ in Thousands	Dec. 31, 2023	Sep. 30, 2023	Jun. 30, 2023	Dec. 31, 2022
Accrued Expenses and Other Current Liabilities
Accrued professional services	$ 1,596			$ 1,409
Accrued royalties	1,689			815
Accrued tax liabilities	822			186
Other accrued expenses	1,948			2,737
Total accrued expenses and other current liabilities	$ 6,055	$ 6,686	$ 6,791	$ 5,147

Stock-Based Compensation - Stoc

Stock-Based Compensation - Stock options (Details) - USD ($) $ / shares in Units, $ in Thousands	12 Months Ended
	Dec. 31, 2023	Dec. 31, 2022	Dec. 31, 2021
Aggregate intrinsic value
Exercised	$ 1,900	$ 3,400	$ 24,300
Intrinsic value of stock options exercised	$ 1,900	$ 3,400	$ 24,300
Employee Stock Option
Stock-based compensation
Options expiration period (in years)	10 years
Number outstanding
Outstanding at the beginning of the period (in shares)	2,188,000
Granted (in shares)	1,187,000
Exercised (in shares)	(161,000)
Cancelled (in shares)	(440,000)
Outstanding at the end of the period (in shares)	2,774,000	2,188,000
Exercisable at the end of the period (in shares)	1,117,000
Vested and expected to vest at the end of the period (in shares)	2,774,000
Weighted-average exercise price
Outstanding at the beginning of the period (in dollars per share)	$ 20.69
Granted (in dollars per share)	16.23
Exercised (in dollars per share)	11.77
Cancelled (in dollars per share)	18.65
Outstanding at the end of the period (in dollars per share)	19.62	$ 20.69
Exercisable at the end of the period (in dollars per share)	22.81
Vested and expected to vest at the end of the period (in dollars per share)	$ 19.62
Weighted-average remaining contractual life
Outstanding (in years)	7 years 10 months 24 days	8 years 1 month 6 days
Exercisable at the end of the period (in years)	6 years 4 months 24 days
Vested and expected to vest at the end of the period (in years)	7 years 10 months 24 days
Aggregate intrinsic value
Outstanding	$ 26,941	$ 4,273
Exercisable at the end of the period	9,176
Vested and expected to vest at the end of the period	$ 26,941
Employee Stock Option | Share-Based Payment Arrangement, Tranche One [Member]
Stock-based compensation
Vesting period (in years)	4 years
Vesting percentage 1 (as a percent)	25%
Employee Stock Option | Share-Based Payment Arrangement, Tranche Two [Member]
Stock-based compensation
Vesting period (in years)	3 years
Vesting percentage 1 (as a percent)	75%

Stock-Based Compensation - Rest

Stock-Based Compensation - Restricted stock units (Details) - Restricted Stock Units (RSUs) [Member] - USD ($) $ / shares in Units, shares in Thousands, $ in Millions	12 Months Ended
	Dec. 31, 2023	Dec. 31, 2022	Dec. 31, 2021
Number of restricted stock units
Unvested restricted common stock at the beginning of the period (in shares)	1,188
Granted (in shares)	868
Vested (in shares)	(453)
Cancelled (in shares)	(275)
Unvested restricted common stock at the end of the period (in shares)	1,328	1,188
Expected to convert	1,328
Weighted-average grant date fair value per share
Unvested restricted common stock at the beginning of the period (in dollars per share)	$ 21.18
Granted (in dollars per share)	15.90	$ 18.32	$ 58.20
Vested (in dollars per share)	23
Cancelled (in dollars per share)	17.50
Unvested restricted common stock at the end of the period (in dollars per share)	17.87	$ 21.18
Expected to convert (in dollars per share)	$ 17.87
Aggregate fair value of restricted stock awards	$ 10.4	$ 9.8	$ 7.3

Stock-Based Compensation - St_2

Stock-Based Compensation - Stock-based compensation plans (Details) - shares shares in Thousands	12 Months Ended
	Dec. 31, 2023	Dec. 31, 2022	Dec. 31, 2021	Dec. 31, 2019	Dec. 31, 2017
2017 Plan
Stock-based compensation
Shares authorized under the plan (in shares)					1,000
Shares outstanding (in shares)	4,000
Shares available for grant under the plan (in shares)	2,000
Annual increase in the shares available for grant under the plan (as a percent of shares of common stock outstanding)	4%
2017 ESPP
Stock-based compensation
Shares authorized under the plan (in shares)				600
Shares available for grant under the plan (in shares)	1,700
Employee stock purchase plan (in shares)	121	57	29
Annual increase in the shares available for grant under the plan (as a percent of shares of common stock outstanding)				1%
Employee Stock Option | 2017 Plan
Stock-based compensation
Shares authorized under the plan (in shares)					2,500

Stock-Based Compensation - Shar

Stock-Based Compensation - Share-based compensation expense (Details) - USD ($) $ in Thousands	12 Months Ended
	Dec. 31, 2023	Dec. 31, 2022	Dec. 31, 2021
Stock-based compensation
Share-based compensation expense	$ 16,823	$ 15,383	$ 15,975
Cost of product revenue
Stock-based compensation
Share-based compensation expense	839	596	471
Cost of service and other revenue
Stock-based compensation
Share-based compensation expense	1,124	819	403
Research and development
Stock-based compensation
Share-based compensation expense	1,713	1,629	1,807
General and Administrative Expense [Member]
Stock-based compensation
Share-based compensation expense	13,147	$ 12,339	$ 13,294
Restricted stock units and stock options
Stock-based compensation
Total unrecognized compensation cost related to unvested stock awards	$ 35,400
Period of recognition of unrecognized compensation cost	2 years 8 months 12 days

Stock-Based Compensation - Blac

Stock-Based Compensation - Black-Scholes valuation model Assumptions (Details) - $ / shares	12 Months Ended
	Dec. 31, 2023	Dec. 31, 2022	Dec. 31, 2021
Fair value assumptions:
Expected dividend yield	0%
Employee Stock Option
Fair value assumptions:
Risk-free interest rate, Minimum	3.50%	1.40%	0.40%
Risk-free interest rate, Maximum	4.70%	4.10%	1.30%
Expected dividend yield	0%	0%	0%
Expected term (in years)			6 years
Expected volatility, Minimum	71.10%	55%	49.20%
Expected volatility, Maximum	83.10%	70.80%	55.60%
Weighted-average grant date fair value	$ 10.63	$ 9.88	$ 29.96
Employee Stock Option | Minimum
Fair value assumptions:
Expected term (in years)	5 years	5 years
Employee Stock Option | Maximum
Fair value assumptions:
Expected term (in years)	5 years 2 months 12 days	5 years 9 months 18 days
Employee Stock Purchase Plan
Fair value assumptions:
Risk-free interest rate, Minimum	5.20%	0.70%	0.10%
Risk-free interest rate, Maximum	5.50%	3.90%
Expected dividend yield	0%	0%	0%
Expected term (in years)	6 months	6 months	6 months
Expected volatility, Minimum	72.80%	51.90%	46.30%
Expected volatility, Maximum	82.50%	117.30%	59.80%
Weighted-average grant date fair value	$ 3.19	$ 3.53	$ 8.06

Net Loss Per Share - Basic and

Net Loss Per Share - Basic and diluted (Details) - USD ($) $ / shares in Units, $ in Thousands	3 Months Ended								6 Months Ended		9 Months Ended		12 Months Ended
	Dec. 31, 2023	Sep. 30, 2023	Jun. 30, 2023	Mar. 31, 2023	Dec. 31, 2022	Sep. 30, 2022	Jun. 30, 2022	Mar. 31, 2022	Jun. 30, 2023	Jun. 30, 2022	Sep. 30, 2023	Sep. 30, 2022	Dec. 31, 2023	Dec. 31, 2022	Dec. 31, 2021
Earnings Per Share [Abstract]
Net loss	$ (8,813)	$ (6,716)	$ (5,551)	$ (7,274)	$ (18,198)	$ (36,550)	$ (26,906)	$ (17,920)	$ (12,825)	$ (44,826)	$ (19,541)	$ (81,376)	$ (28,354)	$ (99,574)	$ (55,548)
Basic weighted average common shares outstanding	37,890,000	37,657,000	37,494,000	37,327,000	37,160,000	37,005,000	36,922,000	36,851,000	37,411,000	36,887,000	37,494,000	36,927,000	37,594,000	36,991,000	35,997,000
Diluted weighted average common shares outstanding	37,890,000	37,657,000	37,494,000	37,327,000	37,160,000	37,005,000	36,922,000	36,851,000	37,411,000	36,887,000	37,494,000	36,927,000	37,594,000	36,991,000	35,997,000
Basic net (loss) income per share	$ (0.23)	$ (0.18)	$ (0.15)	$ (0.19)	$ (0.49)	$ (0.99)	$ (0.73)	$ (0.49)	$ (0.34)	$ (1.22)	$ (0.52)	$ (2.20)	$ (0.75)	$ (2.69)	$ (1.54)
Diluted net (loss) income per share	$ (0.23)	$ (0.18)	$ (0.15)	$ (0.19)	$ (0.49)	$ (0.99)	$ (0.73)	$ (0.49)	$ (0.34)	$ (1.22)	$ (0.52)	$ (2.20)	$ (0.75)	$ (2.69)	$ (1.54)

Net Loss Per Share - Common sha

Net Loss Per Share - Common share equivalents have been excluded from the calculation of diluted net loss per share (Details) - shares shares in Thousands	12 Months Ended
	Dec. 31, 2023	Dec. 31, 2022	Dec. 31, 2021
Common stock and RSUs
Net loss per share
Number of common share equivalents excluded in the calculation of diluted net loss per share	1,512	702	531
Estimated ESPP purchases
Net loss per share
Number of common share equivalents excluded in the calculation of diluted net loss per share	23	52	8
Employee Stock Option
Net loss per share
Number of common share equivalents excluded in the calculation of diluted net loss per share	2,783	446	2,305
Warrant [Member]
Net loss per share
Number of common share equivalents excluded in the calculation of diluted net loss per share	4,318	1,200	2,844

Income Taxes (As Restated) - Co

Income Taxes (As Restated) - Components of loss before income taxes (Details) - USD ($) $ in Thousands	3 Months Ended								6 Months Ended		9 Months Ended		12 Months Ended
	Dec. 31, 2023	Sep. 30, 2023	Jun. 30, 2023	Mar. 31, 2023	Dec. 31, 2022	Sep. 30, 2022	Jun. 30, 2022	Mar. 31, 2022	Jun. 30, 2023	Jun. 30, 2022	Sep. 30, 2023	Sep. 30, 2022	Dec. 31, 2023	Dec. 31, 2022	Dec. 31, 2021
Components of loss before income taxes
United States													$ (30,355)	$ (92,360)	$ (54,414)
Foreign													2,720	(7,050)	(1,170)
Loss before income taxes	$ (8,672)	$ (6,513)	$ (5,316)	$ (7,134)	$ (18,044)	$ (36,478)	$ (26,769)	$ (18,119)	$ (12,450)	$ (44,888)	$ (18,963)	$ (81,366)	$ (27,635)	$ (99,410)	$ (55,584)

Income Taxes (As Restated) - Pr

Income Taxes (As Restated) - Provision for income tax benefit (Details) - USD ($) $ in Thousands	3 Months Ended								6 Months Ended		9 Months Ended		12 Months Ended
	Dec. 31, 2023	Sep. 30, 2023	Jun. 30, 2023	Mar. 31, 2023	Dec. 31, 2022	Sep. 30, 2022	Jun. 30, 2022	Mar. 31, 2022	Jun. 30, 2023	Jun. 30, 2022	Sep. 30, 2023	Sep. 30, 2022	Dec. 31, 2023	Dec. 31, 2022	Dec. 31, 2021
Current:
Federal													$ 0	$ 0	$ 0
State													(161)	(87)	(33)
Foreign													(850)	(372)	(340)
Total deferred income tax benefit													(1,011)	(459)	(373)
Deferred
Federal													0	10	5
State													0	13	(6)
Foreign													292	272	410
Total deferred income tax benefit (provision)													292	295	409
Total income tax (expense) benefit	$ (141)	$ (203)	$ (235)	$ (140)	$ (154)	$ (72)	$ (137)	$ 199	$ (375)	$ 62	$ (578)	$ (10)	$ (719)	$ (164)	$ 36

Income Taxes (As Restated) - Re

Income Taxes (As Restated) - Reconciliation of the federal statutory income tax rate to the effective tax rate (Details)	12 Months Ended
	Dec. 31, 2023	Dec. 31, 2022	Dec. 31, 2021
Reconciliation of the federal statutory income tax rate to the effective tax rate
Federal statutory income tax rate (in percent)	21%	21%	21%
State taxes, net of federal benefit (in percent)	(0.30%)	2.80%	6.60%
Tax credits (in percent)	4.90%	1.30%	2.10%
Share-based compensation (in percent)	(1.60%)	(2.40%)	7.70%
Permanent items (in percent)	(0.60%)	(0.30%)	(1.90%)
Disallowed officer compensation (in percent)	(2.60%)	0%	0%
Deferred tax rate change (in percent)	0%	0%	0.20%
Change in valuation allowance (in percent)	(24.30%)	(20.80%)	(35.20%)
Impairment of goodwill	0%	(1.60%)	0%
Other (in percent)	0.90%	(0.20%)	(0.40%)
Effective income tax rate (in percent)	(2.60%)	(0.20%)	0.10%

Income Taxes (As Restated) - De

Income Taxes (As Restated) - Deferred tax assets and liabilities (Details) - USD ($) $ in Thousands	Dec. 31, 2023	Dec. 31, 2022	Dec. 31, 2021
Deferred tax assets:
Net operating loss carryforwards	$ 77,731	$ 77,689
Tax credits	8,645	7,303
Deferred revenue	2,499	2,385
Amortization	727	847
Stock-based compensation	3,186	2,773
Inventory	1,391	1,507
Capitalized R&D Costs	9,353	5,743
Lease liability	9,948	10,856
Other deferred tax assets	2,357	1,536
Total deferred tax assets	115,837	110,639
Less: valuation allowances	(110,082)	(103,372)	$ (82,654)
Net deferred tax assets	5,755	7,267
Right-of-Use Assets	(4,372)	(5,219)
Depreciation	(1,307)	(1,756)
Amortization acquired intangibles	(1,253)	(1,570)
Other deferred tax liabilities	0	(189)
Net deferred tax liabilities	$ (1,177)	$ (1,467)

Income Taxes (As Restated) - Va

Income Taxes (As Restated) - Valuation allowance deferred tax (Details) - USD ($) $ in Thousands	12 Months Ended
	Dec. 31, 2023	Dec. 31, 2022
Income Tax Disclosure [Abstract]
Balance, beginning of year	$ 103,372	$ 82,654
Change in valuation allowance	6,710	20,718
Balance, end of year	$ 110,082	$ 103,372

Income Taxes (As Restated) (Det

Income Taxes (As Restated) (Details) - USD ($) $ in Thousands	Dec. 31, 2023	Dec. 31, 2022	Dec. 31, 2021	Dec. 31, 2018	Dec. 31, 2017
Income Taxes
Capitalized R&D Costs	$ 9,353	$ 5,743
Tax reserve accrued	0	$ 0	$ 0
Domestic Tax Authority [Member]
Income Taxes
Net operating loss (NOL) carryforwards	308,800
Operating loss carryforwards, not subject to expiry				$ 200,300
Domestic Tax Authority [Member] | Through Tax Year 2037
Income Taxes
Operating loss carryforwards, subject to expiry					$ 108,500
Domestic Tax Authority [Member] | Through Tax Year 2042
Income Taxes
Tax credit carryforwards	6,700
Domestic Tax Authority [Member] | Through Tax Year 2043
Income Taxes
Operating loss carryforwards, subject to expiry	6,700
State and Local Jurisdiction [Member]
Income Taxes
Net operating loss (NOL) carryforwards	214,300
Operating loss carryforwards, not subject to expiry	10,500
State and Local Jurisdiction [Member] | Through Tax Year 2038
Income Taxes
Operating loss carryforwards, subject to expiry	2,500
State and Local Jurisdiction [Member] | Through Tax Year 2042
Income Taxes
Operating loss carryforwards, subject to expiry	$ 203,800

Goodwill and intangible asset_2

Goodwill and intangible assets (Details) - USD ($) $ in Thousands	3 Months Ended	9 Months Ended	12 Months Ended
	Sep. 30, 2022	Sep. 30, 2022	Dec. 31, 2023	Dec. 31, 2022	Dec. 31, 2021	Jun. 30, 2022	Mar. 31, 2022
Rollforward of goodwill balance
Goodwill Impairment	$ 8,200	$ 8,220
Goodwill			$ 0	$ 0		$ 8,675	$ 9,323
Purchased intangible assets
Gross Carrying Value			16,400	16,400
Accumulated Amortization			(9,280)	(7,402)
Cumulative Translation Adjustment			(1,086)	(1,482)
Net Carrying Value			6,034	7,516
Amortization expense			1,600	$ 1,800	$ 2,000
Estimated amortization expense
2024			1,631
2025			1,493
2026			1,465
2027			1,443
2028			2
Thereafter			0
Total amortization expense			$ 6,034
Know How
Purchased intangible assets
Estimated Useful Life (in years)			8 years 6 months	8 years 6 months
Gross Carrying Value			$ 13,000	$ 13,000
Accumulated Amortization			(6,326)	(4,763)
Cumulative Translation Adjustment			(1,050)	(1,433)
Net Carrying Value			$ 5,624	$ 6,804
Weighted Average Life Remaining (in years)			4 years	5 years
Developed technology
Purchased intangible assets
Estimated Useful Life (in years)			7 years	7 years
Gross Carrying Value			$ 1,650	$ 1,650
Accumulated Amortization			(1,581)	(1,458)
Cumulative Translation Adjustment			0	0
Net Carrying Value			$ 69	$ 192
Weighted Average Life Remaining (in years)			1 year 1 month 6 days	2 years 1 month 6 days
Customer relationships
Purchased intangible assets
Gross Carrying Value			$ 1,360	$ 1,360
Accumulated Amortization			(1,067)	(938)
Cumulative Translation Adjustment			(9)	(12)
Net Carrying Value			$ 284	$ 410
Weighted Average Life Remaining (in years)			4 years 1 month 6 days	5 years 1 month 6 days
Non-compete agreements
Purchased intangible assets
Estimated Useful Life (in years)			5 years 6 months	5 years 6 months
Gross Carrying Value			$ 340	$ 340
Accumulated Amortization			(256)	(193)
Cumulative Translation Adjustment			(27)	(37)
Net Carrying Value			$ 57	$ 110
Weighted Average Life Remaining (in years)			1 year	2 years
Trade names
Purchased intangible assets
Estimated Useful Life (in years)			3 years	3 years
Gross Carrying Value			$ 50	$ 50
Accumulated Amortization			(50)	(50)
Cumulative Translation Adjustment			0	0
Net Carrying Value			$ 0	$ 0
Minimum | Customer relationships
Purchased intangible assets
Estimated Useful Life (in years)			8 years 6 months	8 years 6 months
Maximum | Customer relationships
Purchased intangible assets
Estimated Useful Life (in years)			10 years	10 years

Leases (Details)

Leases (Details) - USD ($) $ in Thousands	12 Months Ended
	Dec. 31, 2023	Dec. 31, 2022	Dec. 31, 2021	Sep. 30, 2023	Jun. 30, 2023	Mar. 31, 2023	Sep. 30, 2022	Jun. 30, 2022	Mar. 31, 2022
Leases
Operating lease right-of-use assets	$ 18,251	$ 21,223		$ 19,860	$ 20,380	$ 20,891	$ 27,165	$ 32,935	$ 29,915
Operating lease liabilities, net of current portion	37,223	41,417		$ 38,306	$ 39,378	$ 40,409	$ 42,196	$ 43,135	$ 43,563
Operating lease costs	5,209	5,488	$ 2,726
Restructuring Plan Q3 2022
Leases
Impairment related to restructuring	1,300	16,300
ROU Assets Impairment expense	1,100	12,000	$ 0
Leasehold Improvement Impairment expense	0	4,300
Carrying Value of ROU Assets and Related Leasehold Improvements	$ 10,100	$ 12,800

Leases - Lease Cost (Details)

Leases - Lease Cost (Details) - USD ($) $ in Thousands	12 Months Ended
	Dec. 31, 2023	Dec. 31, 2022	Dec. 31, 2021
Lessee Disclosure [Abstract]
Operating lease costs	$ 5,209	$ 5,488	$ 2,726
Short-term lease cost	3,996	3,417	2,699
Total lease cost	$ 9,205	$ 8,905	$ 5,425

Leases - Supplemental cash flow

Leases - Supplemental cash flow information (Details) - USD ($) $ in Thousands	3 Months Ended	9 Months Ended	12 Months Ended
	Mar. 31, 2022	Sep. 30, 2022	Dec. 31, 2023	Dec. 31, 2022	Dec. 31, 2021
Lessee Disclosure [Abstract]
Cash paid for amounts included in the measurement of operating lease liabilities			$ 8,935	$ 6,539
Right-of-use asset obtained in exchange for lease liabilities	$ 18,156	$ 22,239	$ 0	$ 22,494	$ 0
Weighted average remaining lease term (years)			6 years 9 months 18 days	7 years 9 months 18 days
Weighted average discount rate			7.86%	7.83%

Leases - Undiscounted future le

Leases - Undiscounted future lease payments for non-cancelable operating leases (Details) $ in Thousands	Dec. 31, 2023 USD ($)
Maturity of lease liabilities
2024	$ 7,094
2025	7,254
2026	7,408
2027	7,641
2028	7,880
Thereafter	15,741
Total lease payments	53,018
Less: imputed interest	11,554
Total operating lease liabilities	$ 41,464

Leases - Balance Sheet informat

Leases - Balance Sheet information (Details) - USD ($) $ in Thousands	Dec. 31, 2023	Sep. 30, 2023	Jun. 30, 2023	Mar. 31, 2023	Dec. 31, 2022	Sep. 30, 2022	Jun. 30, 2022	Mar. 31, 2022
Lessee Disclosure [Abstract]
Operating lease right-of-use assets	$ 18,251	$ 19,860	$ 20,380	$ 20,891	$ 21,223	$ 27,165	$ 32,935	$ 29,915
Operating lease liabilities	4,241	4,093	3,986	3,875	2,687	767	2,696	1,886
Operating lease liabilities, net of current portion	37,223	$ 38,306	$ 39,378	$ 40,409	$ 41,417	$ 42,196	$ 43,135	$ 43,563
Total operating lease liabilities	$ 41,464

Commitments and Contingencies -

Commitments and Contingencies - Purchase Commitments (Details) item in Thousands, $ in Thousands	3 Months Ended								6 Months Ended		9 Months Ended		12 Months Ended			36 Months Ended
	Dec. 31, 2023 USD ($) item	Sep. 30, 2023 USD ($)	Jun. 30, 2023 USD ($)	Mar. 31, 2023 USD ($)	Dec. 31, 2022 USD ($)	Sep. 30, 2022 USD ($)	Jun. 30, 2022 USD ($)	Mar. 31, 2022 USD ($)	Jun. 30, 2023 USD ($)	Jun. 30, 2022 USD ($)	Sep. 30, 2023 USD ($)	Sep. 30, 2022 USD ($)	Dec. 31, 2023 USD ($) item	Dec. 31, 2022 USD ($) item	Dec. 31, 2021 USD ($)	Dec. 31, 2024 item
Commitments and contingencies
Purchase commitments expects to incur in next year	$ 3,100												$ 3,100
Cost of revenue	12,143	$ 12,150	$ 11,388	$ 12,463	$ 13,180	$ 16,341	$ 15,183	$ 15,455	$ 23,851	$ 30,638	$ 36,001	$ 46,979	48,144	$ 60,159	$ 45,784
Accounts payable				$ (1,271)				$ (5,306)	$ (631)	$ (3,220)	$ 948	$ (7,010)	1,189	(5,287)	$ 2,339
Stratec Supply Agreement
Commitments and contingencies
Purchase commitments expects to incur in next year	1,000												1,000
Other Commitment	$ 3,700												3,700
Cost of revenue													1,600	$ 500
Accounts payable													$ 2,100
Stratec Supply Agreement | Forecast
Commitments and contingencies
Number of discs purchased | item																515
Stratec Supply Agreement | Stratec Consumables GmBH (Stratec)
Commitments and contingencies
Number of discs shipped | item													218	75
Number of discs required to be shipped | item	222												222

Commitments and Contingencies_2

Commitments and Contingencies - License agreements and Lease commitments (Details) - License agreements - USD ($) $ in Thousands	1 Months Ended	12 Months Ended
	Aug. 31, 2022	Dec. 31, 2023	Dec. 31, 2022	Dec. 31, 2021
Tufts
License agreements
Royalty expense		$ 1,700	$ 1,400	$ 1,600
Harvard University
License agreements
Royalty expense		$ 0	$ 0
Upfront Fee	$ 600

Related Party Transactions (Det

Related Party Transactions (Details) - USD ($) $ in Thousands	3 Months Ended								6 Months Ended		9 Months Ended		12 Months Ended
	Dec. 31, 2023	Sep. 30, 2023	Jun. 30, 2023	Mar. 31, 2023	Dec. 31, 2022	Sep. 30, 2022	Jun. 30, 2022	Mar. 31, 2022	Jun. 30, 2023	Jun. 30, 2022	Sep. 30, 2023	Sep. 30, 2022	Dec. 31, 2023	Dec. 31, 2022	Dec. 31, 2021
Related party transactions
Cost of revenue	$ 12,143	$ 12,150	$ 11,388	$ 12,463	$ 13,180	$ 16,341	$ 15,183	$ 15,455	$ 23,851	$ 30,638	$ 36,001	$ 46,979	$ 48,144	$ 60,159	$ 45,784
Accounts receivable	25,414	24,083	24,369	22,546	19,017	18,330	19,683	22,616	24,369	19,683	24,083	18,330	25,414	19,017
Accounts payable	5,048	4,786	3,238	2,585	3,836	2,106	5,908	4,187	3,238	5,908	4,786	2,106	5,048	3,836
Selling, general and administrative	23,436	23,550	21,308	20,817	18,385	20,479	27,816	25,171	42,125	52,987	65,675	73,466	89,111	91,851	92,910
Commitment to sponsor agreement
Product revenue
Related party transactions
Cost of revenue	7,314	7,209	6,701	7,879	9,491	11,347	10,546	11,457	14,580	22,003	21,789	33,350	29,103	42,841	32,114
Service and other revenue
Related party transactions
Cost of revenue	4,829	$ 4,941	$ 4,687	$ 4,584	$ 3,689	$ 4,994	$ 4,637	$ 3,998	$ 9,271	$ 8,635	$ 14,212	$ 13,629	19,041	17,318	$ 13,670
Harvard University
Related party transactions
Cost of revenue													300
Related party revenue													1,300	700
UltraDx
Related party transactions
Cost of revenue													300	$ 700
Accounts payable	$ 0												$ 0

Restructuring (Details)

Restructuring (Details) $ in Thousands

3 Months Ended

6 Months Ended

9 Months Ended

12 Months Ended

Dec. 31, 2023 USD ($)

Sep. 30, 2023 USD ($)

Jun. 30, 2023 USD ($)

Mar. 31, 2023 USD ($)

Dec. 31, 2022 USD ($)

Sep. 30, 2022 USD ($)

Jun. 30, 2022 USD ($)

Jun. 30, 2023 USD ($)

Jun. 30, 2022 USD ($)

Sep. 30, 2023 USD ($)

Sep. 30, 2022 USD ($)

Dec. 31, 2023 USD ($) position

Dec. 31, 2022 USD ($)

Dec. 31, 2021 USD ($)

Restructuring

Total headcount | position

119

Restructuring

$ 3,800

Payment of severance

3,500

Goodwill Impairment

$ 8,200

$ 8,220

Impairment and restructuring

$ 1,361

$ 0

$ 0

$ (33)

$ 9,006

$ 20,343

$ 207

$ (33)

$ 207

$ (33)

$ 20,550

$ 1,328

29,556

$ 0

Other Restructuring Charges

Restructuring

Impairment and restructuring

25,600

Other Restructuring Charges | ROU assets and related leasehold improvements

Restructuring

Impairment and restructuring

$ 1,300

16,300

Other Restructuring Charges | Software and Software Development Costs [Member]

Restructuring

Impairment and restructuring

1,100

Other Restructuring Charges | Goodwill [Member]

Restructuring

Impairment and restructuring

$ 8,200

Restructuring - Roll forward (D

Restructuring - Roll forward (Details) - Severance and Employee Benefit Costs $ in Thousands	12 Months Ended
	Dec. 31, 2023 USD ($)
Restructuring
Balance at Beginning of period	$ 328
Accrual adjustments	(33)
Cash payments	(16)
Foreign currency translation	8
Balance at End of period	$ 287

Employee benefit plans (Details

Employee benefit plans (Details) - USD ($) $ in Millions	12 Months Ended
	Dec. 31, 2023	Dec. 31, 2022	Dec. 31, 2021
Retirement Benefits [Abstract]
Contribution made to the 401 (k) Plan	$ 0.8	$ 1.2	$ 1.1

Variable Interest Entities (A_2

Variable Interest Entities (As Restated) (Details) - USD ($) $ in Thousands, shares in Millions	3 Months Ended	12 Months Ended
	Jun. 30, 2023	Dec. 31, 2023	Sep. 30, 2023	Mar. 31, 2023	Dec. 31, 2022	Sep. 30, 2022	Jun. 30, 2022	Mar. 31, 2022
Other non-current assets	$ 1,637	$ 1,657	$ 1,603	$ 938	$ 911	$ 910	$ 77	$ 78
UltraDx
Number of contingent consideration collaboration	1
Ownership interest		5%
Variable interest entities
Other non-current assets		$ 800
Variable interest entities | UltraDx
Number of contingent consideration collaboration	1

Quarterly Financial Data (Una_3

Quarterly Financial Data (Unaudited and Restated) - Balance Sheet (Details) - USD ($) $ in Thousands	Dec. 31, 2023	Sep. 30, 2023	Jun. 30, 2023	Mar. 31, 2023	Dec. 31, 2022	Sep. 30, 2022	Jun. 30, 2022	Mar. 31, 2022	Dec. 31, 2021	Dec. 31, 2020
Current assets:
Cash and cash equivalents	$ 174,422	$ 201,261	$ 329,525	$ 329,354	$ 338,740	$ 343,743	$ 361,551	$ 374,543
Marketable securities	146,902	126,449			0
Accounts receivable, net of allowance for expected credit losses	25,414	24,083	24,369	22,546	19,017	18,330	19,683	22,616
Inventory	26,123	20,857	18,333	16,629	16,964	18,044	22,685	24,426
Prepaid expenses and other current assets	9,234	9,216	6,897	6,945	6,803	6,075	9,979	13,878
Total current assets	382,095	381,866	379,124	375,474	381,524	386,192	413,898	435,463
Restricted cash	2,604	2,602	2,601	2,598	2,597	2,596	2,594	2,577
Property and equipment, net	17,926	17,287	18,048	18,690	19,811	21,195	21,976	19,639
Intangible assets, net	6,034	6,003	6,476	7,129	7,516	7,511	8,527	9,692
Goodwill	0				0		8,675	9,323
Operating lease right-of-use assets	18,251	19,860	20,380	20,891	21,223	27,165	32,935	29,915
Other non-current assets	1,657	1,603	1,637	938	911	910	77	78
Current liabilities:
Accounts payable	5,048	4,786	3,238	2,585	3,836	2,106	5,908	4,187
Accrued compensation and benefits	14,170	10,505	8,175	5,411	11,035	10,760	10,777	8,196
Other accrued expenses				4,638		6,041	4,799	8,024
Accrued expenses and other current liabilities	6,055	6,686	6,791		5,147
Deferred revenue	9,468	9,827	10,421	10,682	8,644	8,976	12,157	9,194
Operating lease liabilities	4,241	4,093	3,986	3,875	2,687	767	2,696	1,886
Other current liabilities				291		268	396	268
Total current liabilities	38,982	35,897	32,611	27,482	31,349	28,918	36,733	31,755
Total assets	428,567	429,221	428,266	425,720	433,582	445,569	488,682	506,687
Deferred revenue, net of current portion	1,227	1,126	1,304	1,419	1,415	1,591	1,531	1,222
Operating lease liabilities, net of current portion	37,223	38,306	39,378	40,409	41,417	42,196	43,135	43,563
Other non-current liabilities	1,177	1,105	1,225	1,216	1,469	1,570	1,810	1,691
Total liabilities	78,609	76,434	74,518	70,526	75,650	74,275	83,209	78,231
Commitments and contingencies (Note 16)
Stockholders’ equity:
Common stock	38	38	37	37	37	37	37	37
Additional paid-in capital	783,142	778,615	772,472	768,123	763,629	759,254	756,087	750,717
Accumulated other comprehensive loss	(1,672)	(3,129)	(2,740)	(2,496)	(2,538)	(2,999)	(2,203)	(756)
Accumulated deficit	(431,550)	(422,737)	(416,021)	(410,470)	(403,196)	(384,998)	(348,448)	(321,542)
Total stockholders’ equity	349,958	352,787	353,748	355,194	357,932	371,294	405,473	428,456	$ 442,792	$ 205,825
Total liabilities and stockholders’ equity	428,567	429,221	428,266	425,720	433,582	445,569	488,682	506,687
As Previously Reported
Current assets:
Cash and cash equivalents	174,422	201,261	329,525	329,354	338,740	343,743	361,293	374,317
Marketable securities	146,902	126,449
Accounts receivable, net of allowance for expected credit losses	25,414	24,083	24,423	22,546	19,017	18,330	19,683	22,616
Inventory	22,365	19,945	18,156	17,070	16,786	18,236	21,985	22,669
Prepaid expenses and other current assets	9,291	9,273	6,954	7,002	6,860	6,475	10,237	14,104
Total current assets	378,394	381,011	379,058	375,972	381,403	386,784	413,198	433,706
Restricted cash	2,604	2,647	2,686	2,920	2,597	2,596	2,594	2,577
Property and equipment, net	17,926	17,517	18,328	19,056	20,162	21,441	22,295	19,683
Intangible assets, net	6,034	6,003	6,476	7,129	7,516	7,511	8,527	9,692
Goodwill							8,675	9,323
Operating lease right-of-use assets	18,251	19,860	20,380	20,891	21,223	27,165	32,935	29,298
Other non-current assets	1,802	2,004	2,282	1,345	1,298	1,200	377	378
Current liabilities:
Accounts payable	5,048	4,786	3,238	2,585	3,836	2,106	5,983	4,262
Accrued compensation and benefits	13,659	9,775	7,525	4,880	10,658	10,503	10,638	8,139
Other accrued expenses				4,624		5,951	4,799	8,024
Accrued expenses and other current liabilities	6,041	6,672	6,777		5,133
Deferred revenue	9,468	9,827	10,421	10,682	8,644	8,976	12,157	9,194
Operating lease liabilities	4,241	4,093	3,986	3,875	2,687	767	2,696	1,886
Other current liabilities				291		268	396	268
Total current liabilities	38,457	35,153	31,947	26,937	30,958	28,571	36,669	31,773
Total assets	425,011	429,042	429,210	427,313	434,199	446,697	488,601	504,657
Deferred revenue, net of current portion	1,227	1,126	1,304	1,419	1,415	1,591	1,531	1,222
Operating lease liabilities, net of current portion	37,223	38,306	39,378	40,409	41,417	42,196	43,135	43,563
Other non-current liabilities	1,177	1,105	1,225	1,216	1,469	1,570	1,810	1,691
Total liabilities	78,084	75,690	73,854	69,981	75,259	73,928	83,145	78,249
Commitments and contingencies (Note 16)
Stockholders’ equity:
Common stock	38	38	37	37	37	37	37	37
Additional paid-in capital	783,142	778,615	772,473	768,141	763,688	759,312	756,139	750,742
Accumulated other comprehensive loss	(1,757)	(3,214)	(2,825)	(2,581)	(2,623)	(2,999)	(2,203)	(756)
Accumulated deficit	(434,496)	(422,087)	(414,329)	(408,265)	(402,162)	(383,581)	(348,517)	(323,615)
Total stockholders’ equity	346,927	353,352	355,356	357,332	358,940	372,769	405,456	426,408	440,952	206,125
Total liabilities and stockholders’ equity	425,011	429,042	429,210	427,313	434,199	446,697	488,601	504,657
Adjustments
Current assets:
Cash and cash equivalents	0	0	0	0	0	0	258	226
Marketable securities	0	0
Accounts receivable, net of allowance for expected credit losses	0	0	(54)	0	0	0	0	0
Inventory	3,758	912	177	(441)	178	(192)	700	1,757
Prepaid expenses and other current assets	(57)	(57)	(57)	(57)	(57)	(400)	(258)	(226)
Total current assets	3,701	855	66	(498)	121	(592)	700	1,757
Restricted cash	0	(45)	(85)	(322)	0	0	0	0
Property and equipment, net	0	(230)	(280)	(366)	(351)	(246)	(319)	(44)
Intangible assets, net	0	0	0	0	0	0	0	0
Goodwill							0	0
Operating lease right-of-use assets	0	0	0	0	0	0	0	617
Other non-current assets	(145)	(401)	(645)	(407)	(387)	(290)	(300)	(300)
Current liabilities:
Accounts payable	0	0	0	0	0	0	(75)	(75)
Accrued compensation and benefits	511	730	650	531	377	257	139	57
Other accrued expenses				14		90	0	0
Accrued expenses and other current liabilities	14	14	14		14
Deferred revenue	0	0	0	0	0	0	0	0
Operating lease liabilities	0	0	0	0	0	0	0	0
Other current liabilities				0		0	0	0
Total current liabilities	525	744	664	545	391	347	64	(18)
Total assets	3,556	179	(944)	(1,593)	(617)	(1,128)	81	2,030
Deferred revenue, net of current portion	0	0	0	0	0	0	0	0
Operating lease liabilities, net of current portion	0	0	0	0	0	0	0	0
Other non-current liabilities	0	0	0	0	0	0	0	0
Total liabilities	525	744	664	545	391	347	64	(18)
Commitments and contingencies (Note 16)
Stockholders’ equity:
Common stock	0	0	0	0	0	0	0	0
Additional paid-in capital	0	0	(1)	(18)	(59)	(58)	(52)	(25)
Accumulated other comprehensive loss	85	85	85	85	85	0	0	0
Accumulated deficit	2,946	(650)	(1,692)	(2,205)	(1,034)	(1,417)	69	2,073
Total stockholders’ equity	3,031	(565)	(1,608)	(2,138)	(1,008)	(1,475)	17	2,048	$ 1,840	$ (300)
Total liabilities and stockholders’ equity	$ 3,556	$ 179	$ (944)	$ (1,593)	$ (617)	$ (1,128)	$ 81	$ 2,030

Quarterly Financial Data (Una_4

Quarterly Financial Data (Unaudited and Restated) - Balance Sheet (Parenthetical) (Details) - $ / shares	Dec. 31, 2023	Sep. 30, 2023	Jun. 30, 2023	Mar. 31, 2023	Dec. 31, 2022	Sep. 30, 2022	Jun. 30, 2022	Mar. 31, 2022
Statement of Financial Position [Abstract]
Common stock, par value	$ 0.001	$ 0.001	$ 0.001	$ 0.001	$ 0.001	$ 0.001	$ 0.001	$ 0.001
Common stock, authorized shares	120,000,000	120,000,000	120,000,000	120,000,000	120,000,000	120,000,000	120,000,000	120,000,000
Common stock, shares issued	38,014,000	37,839,000	37,566,000	37,424,000	37,280,000	37,094,000	36,975,000	36,899,000
Common stock, shares outstanding	38,014,000	37,839,000	37,566,000	37,424,000	37,280,000	37,094,000	36,975,000	36,899,000

Quarterly Financial Data (Una_5

Quarterly Financial Data (Unaudited and Restated) - Statement Of Operations (Details) - USD ($) $ / shares in Units, $ in Thousands	3 Months Ended								6 Months Ended		9 Months Ended		12 Months Ended
	Dec. 31, 2023	Sep. 30, 2023	Jun. 30, 2023	Mar. 31, 2023	Dec. 31, 2022	Sep. 30, 2022	Jun. 30, 2022	Mar. 31, 2022	Jun. 30, 2023	Jun. 30, 2022	Sep. 30, 2023	Sep. 30, 2022	Dec. 31, 2023	Dec. 31, 2022	Dec. 31, 2021
Revenues:
Total revenues	$ 31,549	$ 31,559	$ 30,804	$ 28,456	$ 25,824	$ 26,646	$ 23,500	$ 29,552	$ 59,260	$ 53,052	$ 90,819	$ 79,698	$ 122,368	$ 105,522	$ 110,556
Costs of goods sold and services:
Total costs of goods sold and services	12,143	12,150	11,388	12,463	13,180	16,341	15,183	15,455	23,851	30,638	36,001	46,979	48,144	60,159	45,784
Gross profit	19,406	19,409	19,416	15,993	12,644	10,305	8,317	14,097	35,409	22,414	54,818	32,719	74,224	45,363	64,772
Operating expenses:
Research and development	7,210	7,739	6,128	4,987	5,843	6,829	7,257	6,880	11,115	14,137	18,854	20,966	26,064	26,809	28,383
Selling, general and administrative	23,436	23,550	21,308	20,817	18,385	20,479	27,816	25,171	42,125	52,987	65,675	73,466	89,111	91,851	92,910
Other lease costs	1,016	928	968	800	802	609			1,768		2,696	609	3,712	1,411	0
Impairment and restructuring	1,361	0	0	(33)	9,006	20,343	207		(33)	207	(33)	20,550	1,328	29,556	0
Total operating expenses	33,023	32,217	28,404	26,571	34,036	48,260	35,280	32,051	54,975	67,331	87,192	115,591	120,215	149,627	121,293
Loss from operations	(13,617)	(12,808)	(8,988)	(10,578)	(21,392)	(37,955)	(26,963)	(17,954)	(19,566)	(44,917)	(32,374)	(82,872)	(45,991)	(104,264)	(56,521)
Interest income (expense), net	4,319	4,185	3,886	3,449	2,815	1,712	552	52	7,335	604	11,520	2,316	15,839	5,131	(403)
Other income (expense), net	626	2,110	(214)	(5)	533	(235)	(358)	(217)	(219)	(575)	1,891	(810)	2,517	(277)	1,340
Loss before income taxes	(8,672)	(6,513)	(5,316)	(7,134)	(18,044)	(36,478)	(26,769)	(18,119)	(12,450)	(44,888)	(18,963)	(81,366)	(27,635)	(99,410)	(55,584)
Income tax (expense) benefit	(141)	(203)	(235)	(140)	(154)	(72)	(137)	199	(375)	62	(578)	(10)	(719)	(164)	36
Net loss	$ (8,813)	$ (6,716)	$ (5,551)	$ (7,274)	$ (18,198)	$ (36,550)	$ (26,906)	$ (17,920)	$ (12,825)	$ (44,826)	$ (19,541)	$ (81,376)	$ (28,354)	$ (99,574)	$ (55,548)
Net loss per common share, basic (in dollars per share)	$ (0.23)	$ (0.18)	$ (0.15)	$ (0.19)	$ (0.49)	$ (0.99)	$ (0.73)	$ (0.49)	$ (0.34)	$ (1.22)	$ (0.52)	$ (2.20)	$ (0.75)	$ (2.69)	$ (1.54)
Net loss per common share, diluted (in dollars per share)	$ (0.23)	$ (0.18)	$ (0.15)	$ (0.19)	$ (0.49)	$ (0.99)	$ (0.73)	$ (0.49)	$ (0.34)	$ (1.22)	$ (0.52)	$ (2.20)	$ (0.75)	$ (2.69)	$ (1.54)
Weighted-average common shares outstanding, basic (in shares)	37,890,000	37,657,000	37,494,000	37,327,000	37,160,000	37,005,000	36,922,000	36,851,000	37,411,000	36,887,000	37,494,000	36,927,000	37,594,000	36,991,000	35,997,000
Weighted-average common shares outstanding, diluted (in shares)	37,890,000	37,657,000	37,494,000	37,327,000	37,160,000	37,005,000	36,922,000	36,851,000	37,411,000	36,887,000	37,494,000	36,927,000	37,594,000	36,991,000	35,997,000
Product revenue
Revenues:
Total revenues	$ 20,821	$ 19,940	$ 19,549	$ 19,360	$ 16,674	$ 17,693	$ 14,785	$ 20,656	$ 38,909	$ 35,441	$ 58,849	$ 53,134	$ 79,670	$ 69,808	$ 81,062
Costs of goods sold and services:
Total costs of goods sold and services	7,314	7,209	6,701	7,879	9,491	11,347	10,546	11,457	14,580	22,003	21,789	33,350	29,103	42,841	32,114
Service revenue
Revenues:
Total revenues														34,495
Service and other revenue
Revenues:
Total revenues	10,230	10,883	10,470	8,506	8,767	8,370	8,548	8,810	18,976	17,358	29,859	25,728	40,089	34,495	23,629
Costs of goods sold and services:
Total costs of goods sold and services	4,829	4,941	4,687	4,584	3,689	4,994	4,637	3,998	9,271	8,635	14,212	13,629	19,041	17,318	13,670
Collaboration and license revenue
Revenues:
Total revenues	146	237	629	368	170	301	92	86	997	178	1,234	479	1,380	649	648
Grant revenue
Revenues:
Total revenues	352	499	156	222	213	282	75		378	75	877	357	1,229	570	5,217
As Previously Reported
Revenues:
Total revenues	31,549	31,334	31,029	28,456	25,824	26,646	23,500	29,552	59,485	53,052	90,819	79,698	122,368	105,522	110,556
Costs of goods sold and services:
Total costs of goods sold and services	14,750	13,551	11,891	11,530	13,232	15,702	14,789	14,993	23,421	29,782	36,972	45,484	51,722	58,716	48,828
Gross profit	16,799	17,783	19,138	16,926	12,592	10,944	8,711	14,559	36,064	23,270	53,847	34,214	70,646	46,806	61,728
Operating expenses:
Research and development	6,991	7,200	5,946	4,720	5,600	6,631	6,625	7,034	10,666	13,659	17,866	20,290	24,857	25,890	27,978
Selling, general and administrative	24,172	23,595	21,591	20,883	19,272	19,966	27,045	25,712	42,474	52,757	66,069	72,723	90,241	91,995	92,336
Other lease costs	1,016	758	1,162	776	669	609			1,938		2,696	609	3,712	1,278
Impairment and restructuring	1,570	0	0	(33)	9,006	20,341	0		(33)	0	(33)	20,341	1,537	29,347
Total operating expenses	33,749	31,553	28,699	26,346	34,547	47,547	33,670	32,746	55,045	66,416	86,598	113,963	120,347	148,510	120,314
Loss from operations	(16,950)	(13,770)	(9,561)	(9,420)	(21,955)	(36,603)	(24,959)	(18,187)	(18,981)	(43,146)	(32,751)	(79,749)	(49,701)	(101,704)	(58,586)
Interest income (expense), net	4,319	4,185	3,886	3,449	2,815	1,712	552	52	7,335	604	11,520	2,316	15,839	5,131	(403)
Other income (expense), net	363	2,030	(154)	8	614	(101)	(358)	(217)	(146)	(575)	1,884	(676)	2,247	(62)	1,265
Loss before income taxes	(12,268)	(7,555)	(5,829)	(5,963)	(18,526)	(34,992)	(24,765)	(18,352)	(11,792)	(43,117)	(19,347)	(78,109)	(31,615)	(96,635)	(57,724)
Income tax (expense) benefit	(141)	(203)	(235)	(140)	(55)	(72)	(137)	199	(375)	62	(578)	(10)	(719)	(65)	36
Net loss	$ (12,409)	$ (7,758)	$ (6,064)	$ (6,103)	$ (18,581)	$ (35,064)	$ (24,902)	$ (18,153)	$ (12,167)	$ (43,055)	$ (19,925)	$ (78,119)	$ (32,334)	$ (96,700)	$ (57,688)
Net loss per common share, basic (in dollars per share)	$ (0.33)	$ (0.21)	$ (0.16)	$ (0.16)	$ (0.50)	$ (0.95)	$ (0.67)	$ (0.49)	$ (0.33)	$ (1.17)	$ (0.53)	$ (2.12)	$ (0.86)	$ (2.61)	$ (1.60)
Net loss per common share, diluted (in dollars per share)	$ (0.33)	$ (0.21)	$ (0.16)	$ (0.16)	$ (0.50)	$ (0.95)	$ (0.67)	$ (0.49)	$ (0.33)	$ (1.17)	$ (0.53)	$ (2.12)	$ (0.86)	$ (2.61)	$ (1.60)
Weighted-average common shares outstanding, basic (in shares)	37,890,000	37,657,000	37,494,000	37,327,000	37,160,000	37,005,000	36,922,000	36,851,000	37,411,000	36,887,000	37,494,000	36,927,000	37,594,000	36,991,000	35,997,000
Weighted-average common shares outstanding, diluted (in shares)	37,890,000	37,657,000	37,494,000	37,327,000	37,160,000	37,005,000	36,922,000	36,851,000	37,411,000	36,887,000	37,494,000	36,927,000	37,594,000	36,991,000	35,997,000
As Previously Reported | Product revenue
Revenues:
Total revenues	$ 20,821	$ 19,660	$ 19,692	$ 19,287	$ 16,674	$ 17,693	$ 14,785	$ 20,656	$ 38,979	$ 35,441	$ 58,639	$ 53,134	$ 79,460	$ 69,808	$ 81,062
Costs of goods sold and services:
Total costs of goods sold and services	10,025	8,342	7,236	7,033	9,631	10,511	9,921	10,746	14,269	20,667	22,611	31,178	32,636	40,809	34,149
As Previously Reported | Service and other revenue
Revenues:
Total revenues	10,230	10,938	10,552	8,579	8,767	8,370	8,548	8,810	19,131	17,358	30,069	25,728	40,299	34,495	23,629
Costs of goods sold and services:
Total costs of goods sold and services	4,725	5,209	4,655	4,497	3,601	5,191	4,868	4,247	9,152	9,115	14,361	14,306	19,086	17,907	14,679
As Previously Reported | Collaboration and license revenue
Revenues:
Total revenues	146	237	629	368	170	301	92	86	997	178	1,234	479	1,380	649	648
As Previously Reported | Grant revenue
Revenues:
Total revenues	352	499	156	222	213	282	75		378	75	877	357	1,229	570	5,217
Adjustments
Revenues:
Total revenues	0	225	(225)	0	0	0	0	0	(225)	0	0	0	0	0	0
Costs of goods sold and services:
Total costs of goods sold and services	(2,607)	(1,401)	(503)	933	(52)	639	394	462	430	856	(971)	1,495	(3,578)	1,443	(3,044)
Gross profit	2,607	1,626	278	(933)	52	(639)	(394)	(462)	(655)	(856)	971	(1,495)	3,578	(1,443)	3,044
Operating expenses:
Research and development	219	539	182	267	243	198	632	(154)	449	478	988	676	1,207	919	405
Selling, general and administrative	(736)	(45)	(283)	(66)	(887)	513	771	(541)	(349)	230	(394)	743	(1,130)	(144)	574
Other lease costs	0	170	(194)	24	133	0			(170)		0	0	0	133
Impairment and restructuring	(209)	0	0	0	0	2	207		0	207	0	209	(209)	209
Total operating expenses	(726)	664	(295)	225	(511)	713	1,610	(695)	(70)	915	594	1,628	(132)	1,117	979
Loss from operations	3,333	962	573	(1,158)	563	(1,352)	(2,004)	233	(585)	(1,771)	377	(3,123)	3,710	(2,560)	2,065
Interest income (expense), net	0	0	0	0	0	0	0	0	0	0	0	0	0	0	0
Other income (expense), net	263	80	(60)	(13)	(81)	(134)	0	0	(73)	0	7	(134)	270	(215)	75
Loss before income taxes	3,596	1,042	513	(1,171)	482	(1,486)	(2,004)	233	(658)	(1,771)	384	(3,257)	3,980	(2,775)	2,140
Income tax (expense) benefit	0	0	0	0	(99)	0	0	0	0	0	0	0	0	(99)	0
Net loss	$ 3,596	$ 1,042	$ 513	$ (1,171)	$ 383	$ (1,486)	$ (2,004)	$ 233	$ (658)	$ (1,771)	$ 384	$ (3,257)	$ 3,980	$ (2,874)	$ 2,140
Net loss per common share, basic (in dollars per share)	$ 0.10	$ 0.03	$ 0.01	$ (0.03)	$ 0.01	$ (0.04)	$ (0.06)	$ 0	$ (0.01)	$ (0.05)	$ 0.01	$ (0.08)	$ 0.11	$ (0.08)	$ 0.06
Net loss per common share, diluted (in dollars per share)	$ 0.10	$ 0.03	$ 0.01	$ (0.03)	$ 0.01	$ (0.04)	$ (0.06)	$ 0	$ (0.01)	$ (0.05)	$ 0.01	$ (0.08)	$ 0.11	$ (0.08)	$ 0.06
Weighted-average common shares outstanding, basic (in shares)	0	0	0	0	0	0	0	0	0	0	0	0	0	0	0
Weighted-average common shares outstanding, diluted (in shares)	0	0	0	0	0	0	0	0	0	0	0	0	0	0	0
Adjustments | Product revenue
Revenues:
Total revenues	$ 0	$ 280	$ (143)	$ 73	$ 0	$ 0	$ 0	$ 0	$ (70)	$ 0	$ 210	$ 0	$ 210	$ 0	$ 0
Costs of goods sold and services:
Total costs of goods sold and services	(2,711)	(1,133)	(535)	846	(140)	836	625	711	311	1,336	(822)	2,172	(3,533)	2,032	(2,035)
Adjustments | Service and other revenue
Revenues:
Total revenues	0	(55)	(82)	(73)	0	0	0	0	(155)	0	(210)	0	(210)	0	0
Costs of goods sold and services:
Total costs of goods sold and services	104	(268)	32	87	88	(197)	(231)	(249)	119	(480)	(149)	(677)	(45)	(589)	(1,009)
Adjustments | Collaboration and license revenue
Revenues:
Total revenues	0	0	0	0	0	0	0	$ 0	0	0	0	0	0	0	0
Adjustments | Grant revenue
Revenues:
Total revenues	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0		$ 0	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0

Quarterly Financial Data (Una_6

Quarterly Financial Data (Unaudited and Restated) - Statement Of Comprehensive Loss (Details) - USD ($) $ in Thousands	3 Months Ended								6 Months Ended		9 Months Ended		12 Months Ended
	Dec. 31, 2023	Sep. 30, 2023	Jun. 30, 2023	Mar. 31, 2023	Dec. 31, 2022	Sep. 30, 2022	Jun. 30, 2022	Mar. 31, 2022	Jun. 30, 2023	Jun. 30, 2022	Sep. 30, 2023	Sep. 30, 2022	Dec. 31, 2023	Dec. 31, 2022	Dec. 31, 2021
Reclassification [Line Items]
Net loss	$ (8,813)	$ (6,716)	$ (5,551)	$ (7,274)	$ (18,198)	$ (36,550)	$ (26,906)	$ (17,920)	$ (12,825)	$ (44,826)	$ (19,541)	$ (81,376)	$ (28,354)	$ (99,574)	$ (55,548)
Other comprehensive income (loss), net of tax:
Unrealized gains (losses) on marketable securities		(241)									(241)		325	0	0
Foreign currency translation adjustment		(148)	(244)	42		(796)	(1,447)	(1,197)	(202)	(2,644)	(350)	(3,440)	541	(2,979)	(1,993)
Total other comprehensive income (loss)		(389)	(244)	42		(796)	(1,447)	(1,197)	(202)	(2,644)	(591)	(3,440)	866	(2,979)	(1,993)
Comprehensive loss		(7,105)	(5,795)	(7,232)		(37,346)	(28,353)	(19,117)	(13,027)	(47,470)	(20,132)	(84,816)	(27,488)	(102,553)	(57,541)
As Previously Reported
Reclassification [Line Items]
Net loss	(12,409)	(7,758)	(6,064)	(6,103)	(18,581)	(35,064)	(24,902)	(18,153)	(12,167)	(43,055)	(19,925)	(78,119)	(32,334)	(96,700)	(57,688)
Other comprehensive income (loss), net of tax:
Unrealized gains (losses) on marketable securities		(241)									(241)		325
Foreign currency translation adjustment		(148)	(244)	42		(796)	(1,447)	(1,197)	(202)	(2,644)	(350)	(3,440)	541	(3,064)	(1,993)
Total other comprehensive income (loss)		(389)	(244)	42		(796)	(1,447)	(1,197)	(202)	(2,644)	(591)	(3,440)	866	(3,064)	(1,993)
Comprehensive loss		(8,147)	(6,308)	(6,061)		(35,860)	(26,349)	(19,350)	(12,369)	(45,699)	(20,516)	(81,559)	(31,468)	(99,764)	(59,681)
Adjustments
Reclassification [Line Items]
Net loss	$ 3,596	1,042	513	(1,171)	$ 383	(1,486)	(2,004)	233	(658)	(1,771)	384	(3,257)	3,980	(2,874)	2,140
Other comprehensive income (loss), net of tax:
Unrealized gains (losses) on marketable securities		0									0		0
Foreign currency translation adjustment		0	0	0		0	0	0	0	0	0	0	0	85	0
Total other comprehensive income (loss)		0	0	0		0	0	0	0	0	0	0	0	85	0
Comprehensive loss		$ 1,042	$ 513	$ (1,171)		$ (1,486)	$ (2,004)	$ 233	$ (658)	$ (1,771)	$ 384	$ (3,257)	$ 3,980	$ (2,789)	$ 2,140

Quarterly Financial Data (Una_7

Quarterly Financial Data (Unaudited and Restated) - Statement Of Stockholders' Equity (Details) - USD ($) shares in Thousands, $ in Thousands	3 Months Ended								6 Months Ended		9 Months Ended		12 Months Ended
	Dec. 31, 2023	Sep. 30, 2023	Jun. 30, 2023	Mar. 31, 2023	Dec. 31, 2022	Sep. 30, 2022	Jun. 30, 2022	Mar. 31, 2022	Jun. 30, 2023	Jun. 30, 2022	Sep. 30, 2023	Sep. 30, 2022	Dec. 31, 2023	Dec. 31, 2022	Dec. 31, 2021
Increase (Decrease) in Stockholders' Equity
Beginning Balance	$ 352,787	$ 353,748	$ 355,194	$ 357,932	$ 371,294	$ 405,473	$ 428,456	$ 442,792	$ 357,932	$ 442,792	$ 357,932	$ 442,792	$ 357,932	$ 442,792	$ 205,825
Issuance of common stock under stock plans, including tax effects		1,800	139	551		407	211	979	690	1,190	2,490	1,597	2,691	2,310	8,815
Sale of common stock in underwritten public offering, net															269,718
Stock-based compensation expense		4,344	4,210	3,943		2,760	5,159	3,802	8,153	8,961	12,497	11,721	16,823	15,383	15,975
Unrealized gains on marketable securities, net of tax		(241)									(241)		325	0	0
Foreign currency translation adjustment		(148)	(244)	42		(796)	(1,447)	(1,197)	(202)	(2,644)	(350)	(3,440)	541	(2,979)	(1,993)
Net loss	(8,813)	(6,716)	(5,551)	(7,274)	(18,198)	(36,550)	(26,906)	(17,920)	(12,825)	(44,826)	(19,541)	(81,376)	(28,354)	(99,574)	(55,548)
Ending Balance	$ 349,958	$ 352,787	$ 353,748	$ 355,194	$ 357,932	$ 371,294	$ 405,473	$ 428,456	$ 353,748	$ 405,473	$ 352,787	$ 371,294	$ 349,958	$ 357,932	$ 442,792
Common Stock
Increase (Decrease) in Stockholders' Equity
Beginning balance (in shares)	37,839	37,566	37,424	37,280	37,094	36,975	36,899	36,768	37,280	36,768	37,280	36,768	37,280	36,768	31,797
Beginning Balance	$ 38	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 32
Issuance of common stock under stock plans, including tax effects (in shares)		273	142	144		119	76	131	286	207	559	326	734	512	864
Issuance of common stock under stock plans, including tax effects		$ 1									$ 1		$ 1		$ 1
Sale of common stock in underwritten public offering, net (in shares)															4,107
Sale of common stock in underwritten public offering, net															$ 4
Ending Balance (in shares)	38,014	37,839	37,566	37,424	37,280	37,094	36,975	36,899	37,566	36,975	37,839	37,094	38,014	37,280	36,768
Ending Balance	$ 38	$ 38	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 38	$ 37	$ 38	$ 37	$ 37
Additional paid-in capital
Increase (Decrease) in Stockholders' Equity
Beginning Balance	778,615	772,472	768,123	763,629	759,254	756,087	750,717	745,936	763,629	745,936	763,629	745,936	763,629	745,936	451,433
Issuance of common stock under stock plans, including tax effects		1,799	139	551		407	211	979	690	1,190	2,489	1,597	2,690	2,310	8,814
Sale of common stock in underwritten public offering, net															269,714
Stock-based compensation expense		4,344	4,210	3,943		2,760	5,159	3,802	8,153	8,961	12,497	11,721	16,823	15,383	15,975
Ending Balance	783,142	778,615	772,472	768,123	763,629	759,254	756,087	750,717	772,472	756,087	778,615	759,254	783,142	763,629	745,936
Accumulated other comprehensive income (loss)
Increase (Decrease) in Stockholders' Equity
Beginning Balance	(3,129)	(2,740)	(2,496)	(2,538)	(2,999)	(2,203)	(756)	441	(2,538)	441	(2,538)	441	(2,538)	441	2,434
Unrealized gains on marketable securities, net of tax		(241)									(241)		325
Foreign currency translation adjustment		(148)	(244)	42		(796)	(1,447)	(1,197)	(202)	(2,644)	(350)	(3,440)	541	(2,979)	(1,993)
Ending Balance	(1,672)	(3,129)	(2,740)	(2,496)	(2,538)	(2,999)	(2,203)	(756)	(2,740)	(2,203)	(3,129)	(2,999)	(1,672)	(2,538)	441
Accumulated deficit
Increase (Decrease) in Stockholders' Equity
Beginning Balance	(422,737)	(416,021)	(410,470)	(403,196)	(384,998)	(348,448)	(321,542)	(303,622)	(403,196)	(303,622)	(403,196)	(303,622)	(403,196)	(303,622)	(248,074)
Net loss		(6,716)	(5,551)	(7,274)		(36,550)	(26,906)	(17,920)	(12,825)	(44,826)	(19,541)	(81,376)	(28,354)	(99,574)	(55,548)
Ending Balance	(431,550)	(422,737)	(416,021)	(410,470)	(403,196)	(384,998)	(348,448)	(321,542)	(416,021)	(348,448)	(422,737)	(384,998)	(431,550)	(403,196)	(303,622)
As Previously Reported
Increase (Decrease) in Stockholders' Equity
Beginning Balance	353,352	355,356	357,332	358,940	372,769	405,456	426,408	440,952	358,940	440,952	358,940	440,952	358,940	440,952	206,125
Issuance of common stock under stock plans, including tax effects		1,800	139	551		407	211	979	690	1,190	2,490	1,597	2,691	2,310	8,815
Sale of common stock in underwritten public offering, net															269,718
Stock-based compensation expense		4,343	4,193	3,902		2,766	5,186	3,827	8,095	9,013	12,438	11,779	16,764	15,442	15,975
Unrealized gains on marketable securities, net of tax		(241)									(241)		325
Foreign currency translation adjustment		(148)	(244)	42		(796)	(1,447)	(1,197)	(202)	(2,644)	(350)	(3,440)	541	(3,064)	(1,993)
Net loss	(12,409)	(7,758)	(6,064)	(6,103)	(18,581)	(35,064)	(24,902)	(18,153)	(12,167)	(43,055)	(19,925)	(78,119)	(32,334)	(96,700)	(57,688)
Ending Balance	$ 346,927	$ 353,352	$ 355,356	$ 357,332	$ 358,940	$ 372,769	$ 405,456	$ 426,408	$ 355,356	$ 405,456	$ 353,352	$ 372,769	$ 346,927	$ 358,940	$ 440,952
As Previously Reported | Common Stock
Increase (Decrease) in Stockholders' Equity
Beginning balance (in shares)	37,839	37,566	37,424	37,280	37,094	36,975	36,899	36,768	37,280	36,768	37,280	36,768	37,280	36,768	31,797
Beginning Balance	$ 38	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 32
Issuance of common stock under stock plans, including tax effects (in shares)		273	142	144		119	76	131	286	207	559	326	734	512	864
Issuance of common stock under stock plans, including tax effects		$ 1									$ 1		$ 1		$ 1
Sale of common stock in underwritten public offering, net (in shares)															4,107
Sale of common stock in underwritten public offering, net															$ 4
Ending Balance (in shares)	38,014	37,839	37,566	37,424	37,280	37,094	36,975	36,899	37,566	36,975	37,839	37,094	38,014	37,280	36,768
Ending Balance	$ 38	$ 38	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 37	$ 38	$ 37	$ 38	$ 37	$ 37
As Previously Reported | Additional paid-in capital
Increase (Decrease) in Stockholders' Equity
Beginning Balance	778,615	772,473	768,141	763,688	759,312	756,139	750,742	745,936	763,688	745,936	763,688	745,936	763,688	745,936	451,433
Issuance of common stock under stock plans, including tax effects		1,799	139	551		407	211	979	690	1,190	2,489	1,597	2,690	2,310	8,814
Sale of common stock in underwritten public offering, net															269,714
Stock-based compensation expense		4,343	4,193	3,902		2,766	5,186	3,827	8,095	9,013	12,438	11,779	16,764	15,442	15,975
Ending Balance	783,142	778,615	772,473	768,141	763,688	759,312	756,139	750,742	772,473	756,139	778,615	759,312	783,142	763,688	745,936
As Previously Reported | Accumulated other comprehensive income (loss)
Increase (Decrease) in Stockholders' Equity
Beginning Balance	(3,214)	(2,825)	(2,581)	(2,623)	(2,999)	(2,203)	(756)	441	(2,623)	441	(2,623)	441	(2,623)	441	2,434
Unrealized gains on marketable securities, net of tax		(241)									(241)		325
Foreign currency translation adjustment		(148)	(244)	42		(796)	(1,447)	(1,197)	(202)	(2,644)	(350)	(3,440)	541	(3,064)	(1,993)
Ending Balance	(1,757)	(3,214)	(2,825)	(2,581)	(2,623)	(2,999)	(2,203)	(756)	(2,825)	(2,203)	(3,214)	(2,999)	(1,757)	(2,623)	441
As Previously Reported | Accumulated deficit
Increase (Decrease) in Stockholders' Equity
Beginning Balance	(422,087)	(414,329)	(408,265)	(402,162)	(383,581)	(348,517)	(323,615)	(305,462)	(402,162)	(305,462)	(402,162)	(305,462)	(402,162)	(305,462)	(247,774)
Net loss		(7,758)	(6,064)	(6,103)		(35,064)	(24,902)	(18,153)	(12,167)	(43,055)	(19,925)	(78,119)	(32,334)	(96,700)	(57,688)
Ending Balance	(434,496)	(422,087)	(414,329)	(408,265)	(402,162)	(383,581)	(348,517)	(323,615)	(414,329)	(348,517)	(422,087)	(383,581)	(434,496)	(402,162)	(305,462)
Adjustments
Increase (Decrease) in Stockholders' Equity
Beginning Balance	(565)	(1,608)	(2,138)	(1,008)	(1,475)	17	2,048	1,840	(1,008)	1,840	(1,008)	1,840	(1,008)	1,840	(300)
Issuance of common stock under stock plans, including tax effects		0	0	0		0	0	0	0	0	0	0	0	0	0
Sale of common stock in underwritten public offering, net															0
Stock-based compensation expense		1	17	41		(6)	(27)	(25)	58	(52)	59	(58)	59	(59)	0
Unrealized gains on marketable securities, net of tax		0									0		0
Foreign currency translation adjustment		0	0	0		0	0	0	0	0	0	0	0	85	0
Net loss	3,596	1,042	513	(1,171)	383	(1,486)	(2,004)	233	(658)	(1,771)	384	(3,257)	3,980	(2,874)	2,140
Ending Balance	$ 3,031	$ (565)	$ (1,608)	$ (2,138)	$ (1,008)	$ (1,475)	$ 17	$ 2,048	$ (1,608)	$ 17	$ (565)	$ (1,475)	$ 3,031	$ (1,008)	$ 1,840
Adjustments | Common Stock
Increase (Decrease) in Stockholders' Equity
Beginning balance (in shares)	0	0	0	0	0	0	0	0	0	0	0	0	0	0	0
Beginning Balance	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0
Ending Balance (in shares)	0	0	0	0	0	0	0	0	0	0	0	0	0	0	0
Ending Balance	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0	$ 0
Adjustments | Additional paid-in capital
Increase (Decrease) in Stockholders' Equity
Beginning Balance	0	(1)	(18)	(59)	(58)	(52)	(25)	0	(59)	0	(59)	0	(59)	0	0
Stock-based compensation expense		1	17	41		(6)	(27)	(25)	58	(52)	59	(58)	59	(59)
Ending Balance	0	0	(1)	(18)	(59)	(58)	(52)	(25)	(1)	(52)	0	(58)	0	(59)	0
Adjustments | Accumulated other comprehensive income (loss)
Increase (Decrease) in Stockholders' Equity
Beginning Balance	85	85	85	85	0	0	0	0	85	0	85	0	85	0	0
Foreign currency translation adjustment														85
Ending Balance	85	85	85	85	85	0	0	0	85	0	85	0	85	85	0
Adjustments | Accumulated deficit
Increase (Decrease) in Stockholders' Equity
Beginning Balance	(650)	(1,692)	(2,205)	(1,034)	(1,417)	69	2,073	1,840	(1,034)	1,840	(1,034)	1,840	(1,034)	1,840	(300)
Net loss		1,042	513	(1,171)		(1,486)	(2,004)	233	(658)	(1,771)	384	(3,257)	3,980	(2,874)	2,140
Ending Balance	$ 2,946	$ (650)	$ (1,692)	$ (2,205)	$ (1,034)	$ (1,417)	$ 69	$ 2,073	$ (1,692)	$ 69	$ (650)	$ (1,417)	$ 2,946	$ (1,034)	$ 1,840

Quarterly Financial Data (Una_8

Quarterly Financial Data (Unaudited and Restated) - Statement Of Cash Flows (Details) - USD ($) $ in Thousands	3 Months Ended			6 Months Ended		9 Months Ended		12 Months Ended
	Mar. 31, 2023	Sep. 30, 2022	Mar. 31, 2022	Jun. 30, 2023	Jun. 30, 2022	Sep. 30, 2023	Sep. 30, 2022	Dec. 31, 2023	Dec. 31, 2022	Dec. 31, 2021
Cash flows from operating activities:
Net loss	$ (7,274)		$ (17,920)	$ (12,825)	$ (44,826)	$ (19,541)	$ (81,376)	$ (28,354)	$ (99,574)	$ (55,548)
Adjustments to reconcile net loss to net cash used in operating activities:
Depreciation and amortization expense	1,454		1,373	2,883	2,820	4,843	4,231	6,275	5,409	4,880
Credit losses on accounts receivable	178		(171)	324	581	311	102	336	(301)	213
Non-cash operating lease expense	334						1,099
Reduction in the carrying amount of right-of-use assets			(269)		925
Foreign currency losses						359
Unrealized losses on marketable securities						(241)
Amortization of (discount) premium on marketable securities						(1,249)		(1,964)	0	0
Accretion of marketable securities						(1,249)		(1,964)	0	0
Operating lease right-of-use asset amortization				1,002		1,518		2,015	715	499
Stock-based compensation expense	3,943		3,802	8,153	8,961	12,497	11,721	16,823	15,383	15,975
Goodwill Impairment		$ 8,200					8,220
Impairment Expense							8,904
Other non-cash items							39
Deferred income taxes	270					242
Other operating activity				602				(150)	(340)	692
Loss on disposal of fixed assets					6	46
Changes in assets and liabilities:
Accounts receivable	(3,741)		1,319	(5,750)	3,479	(5,615)	5,045	(6,695)	5,156	(6,853)
Inventory	530		(206)	(1,180)	1,515	(3,700)	6,146	(8,944)	7,243	(10,125)
Prepaid expenses and other current assets	(422)		(2,071)	(527)	961	(2,829)	(282)	(2,371)	(738)	(166)
Other non-current assets	(33)		1	(707)	2	(702)	(857)	(717)	(822)	(2)
Accounts payable	(1,271)		(5,306)	(631)	(3,220)	948	(7,010)	1,189	(5,287)	2,339
Accrued compensation and benefits, accrued expenses, and other current liabilities	(5,809)		(4,864)	(1,053)	(4,248)	1,229	(2,914)	4,410	(3,599)	606
Contract acquisition costs			(41)		30		331
Deferred revenue	2,041		2,956	1,666	6,228	894	3,108	635	2,599	1,462
Operating lease liabilities	179		(87)	(730)	(3,128)	(1,690)	(1,156)	(2,645)	(266)	(1,230)
Other non-current liabilities	(203)		(271)	(72)	(9)	(107)	128	(53)	10	(363)
Net cash used in operating activities	(9,824)		(21,755)	(8,791)	(29,716)	(12,657)	(44,521)	(18,849)	(48,611)	(47,621)
Cash flows from investing activities:
Purchases of marketable debt securities						(125,200)		(175,613)	0	0
Purchases of property and equipment	(136)		(1,335)	(947)	(6,342)	(1,878)	(10,019)	(3,841)	(11,614)	(13,675)
Proceeds from RADx grant on assets purchased			520		520		520	0	520	7,278
Net cash used in investing activities	(136)		(815)	(947)	(5,822)	(127,078)	(9,499)	(148,454)	(11,094)	(6,397)
Cash flows from financing activities:
Proceeds from stock options exercised	13		385		596		716
Proceeds from common stock issued under stock plans	551		594	777	594	2,632	881	2,889	2,311	8,815
Payments for employee taxes withheld on stock-based compensation awards	(13)			(87)		(142)		(198)	0	0
Payments on notes payable								0	0	(7,738)
Net cash provided by financing activities	551		979	690	1,190	2,490	1,597	2,691	2,311	270,795
Net increase (decrease) in cash, cash equivalents, and restricted cash	(9,409)		(21,591)	(9,048)	(34,348)	(137,245)	(52,423)	(164,612)	(57,394)	216,777
Effect of exchange rate changes on cash, cash equivalents, and restricted cash	24		(558)	(163)	(776)	(229)	(507)	301	(538)	(92)
Cash, cash equivalents, and restricted cash at beginning of period	341,337	364,145	399,269	341,337	399,269	341,337	399,269	341,337	399,269	182,584
Cash, cash equivalents, and restricted cash at end of period	331,952	346,339	377,120	332,126	364,145	203,863	346,339	177,026	341,337	399,269
Supplemental disclosure of cash flow information:
Cash paid for taxes	246			502		719	263	808	684	0
Purchases of property and equipment in accounts payable and accruals	147						288	419	152	229
Shares received as consideration under product sales agreement (Note 4, 7)				1,000		775
Cash paid for interest								0	0	389
Right-of-use asset obtained in exchange for lease liabilities			18,156				22,239	0	22,494	0
Cash and cash equivalents	329,354	343,743	374,543	329,525	361,551	201,261	343,743	174,422	338,740
Restricted cash	2,598	2,596	2,577	2,601	2,594	2,602	2,596	2,604	2,597
Shares received as consideration under product sales agreement (Note 4, 7)								775	0	0
As Previously Reported
Cash flows from operating activities:
Net loss	(6,103)		(18,153)	(12,167)	(43,055)	(19,925)	(78,119)	(32,334)	(96,700)	(57,688)
Adjustments to reconcile net loss to net cash used in operating activities:
Depreciation and amortization expense	1,439		1,358	2,845	2,790	4,788	4,186	6,364	5,349	4,851
Credit losses on accounts receivable	178		(171)	324	581	311	102	336	(301)	213
Non-cash operating lease expense	334						1,099
Reduction in the carrying amount of right-of-use assets			348		925
Foreign currency losses						359
Unrealized losses on marketable securities						(241)
Amortization of (discount) premium on marketable securities						(1,249)		(1,964)
Accretion of marketable securities						(1,249)		(1,964)
Operating lease right-of-use asset amortization				1,002		1,518		2,015	715	499
Stock-based compensation expense	3,902		3,827	8,095	9,013	12,438	11,779	16,764	15,442	15,975
Goodwill Impairment							8,220
Impairment Expense							8,695
Other non-cash items							39
Deferred income taxes	270					242
Other operating activity				548				150	(439)	692
Loss on disposal of fixed assets					6	46
Changes in assets and liabilities:
Accounts receivable	(3,741)		1,319	(5,750)	3,479	(5,615)	5,045	(6,695)	5,156	(6,853)
Inventory	(89)		(484)	(1,181)	180	(2,966)	3,919	(5,364)	5,386	(8,090)
Prepaid expenses and other current assets	(422)		(2,070)	(527)	930	(2,829)	(262)	(2,371)	(568)	(393)
Other non-current assets	(33)		1	(965)	2	(716)	(859)	(775)	(909)	(2)
Accounts payable	(1,271)		(5,306)	(631)	(3,220)	948	(7,085)	1,189	(5,362)	2,414
Accrued compensation and benefits, accrued expenses, and other current liabilities	(5,983)		(4,921)	(1,326)	(4,387)	876	(3,021)	4,276	(3,976)	606
Contract acquisition costs			(41)		30		0
Deferred revenue	2,041		2,956	1,666	6,228	894	3,108	635	2,599	1,462
Operating lease liabilities	179		(87)	(730)	(3,128)	(1,690)	(1,156)	(2,645)	(266)	(1,230)
Other non-current liabilities	(203)		(271)	(72)	(9)	(107)	128	(53)	10	(363)
Net cash used in operating activities	(9,502)		(21,695)	(8,869)	(29,635)	(12,918)	(44,182)	(18,902)	(48,272)	(47,907)
Cash flows from investing activities:
Purchases of marketable debt securities						(125,200)		(175,613)		0
Purchases of property and equipment	(136)		(1,394)	(784)	(6,454)	(1,572)	(10,131)	(3,788)	(11,726)	(13,616)
Proceeds from RADx grant on assets purchased			520		520		520		520	7,278
Net cash used in investing activities	(136)		(874)	(784)	(5,934)	(126,772)	(9,611)	(148,401)	(11,206)	(6,338)
Cash flows from financing activities:
Proceeds from stock options exercised	13		385		596		716
Proceeds from common stock issued under stock plans	551		594	777	594	2,632	881	2,889	2,311	8,815
Payments for employee taxes withheld on stock-based compensation awards	(13)			(87)		(142)		(198)
Payments on notes payable										(7,738)
Net cash provided by financing activities	551		979	690	1,190	2,490	1,597	2,691	2,311	270,795
Net increase (decrease) in cash, cash equivalents, and restricted cash	(9,087)		(21,590)	(8,963)	(34,379)	(137,200)	(52,196)	(164,612)	(57,167)	216,550
Effect of exchange rate changes on cash, cash equivalents, and restricted cash	24		(558)	(163)	(776)	(229)	(507)	301	(538)	(92)
Cash, cash equivalents, and restricted cash at beginning of period	341,337	363,887	399,042	341,337	399,042	341,337	399,042	341,337	399,042	182,584
Cash, cash equivalents, and restricted cash at end of period	332,274	346,339	376,894	332,211	363,887	203,908	346,339	177,026	341,337	399,042
Supplemental disclosure of cash flow information:
Cash paid for taxes	246			502		719	263	808	684
Purchases of property and equipment in accounts payable and accruals	147						198	419	152	229
Shares received as consideration under product sales agreement (Note 4, 7)				1,000		775
Cash paid for interest										389
Right-of-use asset obtained in exchange for lease liabilities			18,156				22,239		22,494
Cash and cash equivalents	329,354	343,743	374,317	329,525	361,293	201,261	343,743	174,422	338,740
Restricted cash	2,920	2,596	2,577	2,686	2,594	2,647	2,596	2,604	2,597
Shares received as consideration under product sales agreement (Note 4, 7)								775
Adjustments
Cash flows from operating activities:
Net loss	(1,171)		233	(658)	(1,771)	384	(3,257)	3,980	(2,874)	2,140
Adjustments to reconcile net loss to net cash used in operating activities:
Depreciation and amortization expense	15		15	38	30	55	45	(89)	60	29
Credit losses on accounts receivable	0		0	0	0	0	0	0	0	0
Non-cash operating lease expense	0						0
Reduction in the carrying amount of right-of-use assets			(617)		0
Foreign currency losses						0
Unrealized losses on marketable securities						0
Amortization of (discount) premium on marketable securities						0		0
Accretion of marketable securities						0		0
Operating lease right-of-use asset amortization				0		0		0	0	0
Stock-based compensation expense	41		(25)	58	(52)	59	(58)	59	(59)	0
Goodwill Impairment							0
Impairment Expense							209
Other non-cash items							0
Deferred income taxes	0					0
Other operating activity				54				(300)	99	0
Loss on disposal of fixed assets					0	0
Changes in assets and liabilities:
Accounts receivable	0		0	0	0	0	0	0	0	0
Inventory	619		278	1	1,335	(734)	2,227	(3,580)	1,857	(2,035)
Prepaid expenses and other current assets	0		(1)	0	31	0	(20)	0	(170)	227
Other non-current assets	0		0	258	0	14	2	58	87	0
Accounts payable	0		0	0	0	0	75	0	75	(75)
Accrued compensation and benefits, accrued expenses, and other current liabilities	174		57	273	139	353	107	134	377	0
Contract acquisition costs			0		0		331
Deferred revenue	0		0	0	0	0	0	0	0	0
Operating lease liabilities	0		0	0	0	0	0	0	0	0
Other non-current liabilities	0		0	0	0	0	0	0	0	0
Net cash used in operating activities	(322)		(60)	78	(81)	261	(339)	53	(339)	286
Cash flows from investing activities:
Purchases of marketable debt securities						0		0		0
Purchases of property and equipment	0		59	(163)	112	(306)	112	(53)	112	(59)
Proceeds from RADx grant on assets purchased			0		0		0		0	0
Net cash used in investing activities	0		59	(163)	112	(306)	112	(53)	112	(59)
Cash flows from financing activities:
Proceeds from stock options exercised	0		0		0		0
Proceeds from common stock issued under stock plans	0		0	0	0	0	0	0	0	0
Payments for employee taxes withheld on stock-based compensation awards	0			0		0		0
Payments on notes payable										0
Net cash provided by financing activities	0		0	0	0	0	0	0	0	0
Net increase (decrease) in cash, cash equivalents, and restricted cash	(322)		(1)	(85)	31	(45)	(227)	0	(227)	227
Effect of exchange rate changes on cash, cash equivalents, and restricted cash	0		0	0	0	0	0	0	0	0
Cash, cash equivalents, and restricted cash at beginning of period	0	258	227	0	227	0	227	0	227	0
Cash, cash equivalents, and restricted cash at end of period	(322)	0	226	(85)	258	(45)	0	0	0	227
Supplemental disclosure of cash flow information:
Cash paid for taxes	0			0		0	0	0	0
Purchases of property and equipment in accounts payable and accruals	0						90	0	0	0
Shares received as consideration under product sales agreement (Note 4, 7)				0		0
Cash paid for interest										0
Right-of-use asset obtained in exchange for lease liabilities			0				0		0
Cash and cash equivalents	0	0	226	0	258	0	0	0	0
Restricted cash	$ (322)	$ 0	$ 0	$ (85)	$ 0	$ (45)	$ 0	0	0
Shares received as consideration under product sales agreement (Note 4, 7)								0
Underwritten public offering
Cash flows from financing activities:
Sale of common stock in underwritten public offering, net								$ 0	$ 0	269,718
Underwritten public offering | As Previously Reported
Cash flows from financing activities:
Sale of common stock in underwritten public offering, net										269,718
Underwritten public offering | Adjustments
Cash flows from financing activities:
Sale of common stock in underwritten public offering, net										$ 0