quarter, but prompt actions by our East Texas operations team allowed production to return within ten days, and the current trend is exceeding internal expectations. Workover expenses did not materially increase, and our East Texas asset remains one of our highest margin and best cash flowing areas.
Non-Operated Eagle Ford:
| • | | Production: 88.5 MBoe; 1.0 Mboepd |
| • | | Commodity Mix: 84% oil, 9% NGLs, 7% natural gas |
| • | | LOE: $1.1 million or $12.30 per Boe |
Capital spend in the first quarter was focused on the completion of DUCs and associated production facilities, and a portion of the expenditures will carry over into the second quarter. As of May 5, 2021, 23 DUCs have been turned online. The wells are all located in the core of the Eagle Ford in Karnes County. Amplify is pleased with the well results, which have exceeded our type curves to date with an average gross oil IP rate of 1,200 Bopd. The Company will continue opportunistically participating in attractive non-operated Eagle Ford projects as they arise.
Costs and Expenses
Lease operating expenses in the first quarter of 2021 were approximately $28.9 million, or $13.01 per Boe, an increase of $0.4 million compared to $28.5 million, or $11.77 per Boe, in the fourth quarter of 2020. This increase was mainly attributed to the storm-related impacts to our Oklahoma, East Texas and Eagle Ford assets. The resulting impact was a $0.7 MM increase in workover expense in the month of February, or $0.30 per Boe for the quarter, and the aforementioned 700 MBoe production deferral. The quick and prudent decision-making of our operations teams minimized the financial impact and mitigated the overall cost of the storm to the Company. Amplify remains committed to the disciplined management of operating expenses, and the asset teams continue to explore additional methods of reducing costs moving forward.
Severance and Ad Valorem taxes in the first quarter of 2021 consisted of $4.6 million, an increase of $1.6 million compared to $3.0 million in the fourth quarter of 2020. On a percentage basis, Amplify paid approximately 6.4% of total oil, NGLs, and natural gas sales revenue in taxes this quarter compared to 5.4% in the previous quarter. The quarter-over-quarter increase was generally attributable to a positive tax adjustment made in the fourth quarter of 2020 and higher commodity pricing, partially offset by naturally declining production in the same period.
Amplify incurred $4.6 million, or $2.06 per Boe, of gathering, processing, and transportation expenses in the first quarter of 2021, compared to $5.5 million, or $2.29 per Boe, in the previous quarter. The decrease was generally attributable to natural production decline, along with production impacts from Winter Storm Uri in February of 2021.
Due to year-end processes, first quarter cash G&A expenses are typically the highest of the year, and the $6.6 million incurred this quarter was an expected increase of $0.8 million from the fourth quarter of 2020. However, first quarter cash G&A decreased year-over-year by $2.1 million as Amplify undertook a transformative corporate expense reduction plan in 2020. The Company’s current projected full year 2021 cash G&A estimate remains approximately $23 million.
Depreciation, depletion, and amortization expense for the first quarter of 2021 totaled $7.3 million, or $3.31 per Boe, compared to $9.1 million, or $3.77 per Boe, in the previous quarter. The decrease of $1.8 million can be attributed to non-cash impairments in 2020. Amplify does not expect further impairments in the foreseeable future.
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