Significant Collaboration Transactions - Additional Information (Details) | Jun. 01, 2019USD ($)$ / sharesshares | Oct. 01, 2018USD ($)shares | Sep. 30, 2017USD ($) | Dec. 31, 2017USD ($) | Dec. 31, 2020USD ($)shares | Dec. 31, 2021USD ($)shares | Dec. 31, 2020USD ($)Obligationsshares | Dec. 31, 2019USD ($)shares | Dec. 31, 2018USD ($)shares | Dec. 31, 2017USD ($) | Jan. 31, 2021USD ($)shares | Jul. 31, 2020USD ($) | Dec. 01, 2019USD ($) | Jul. 31, 2019USD ($) | Dec. 01, 2018USD ($) | Sep. 30, 2018USD ($) |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Number of additional common units received | shares | | | | | | | 5,400,000 | | | | | | | | | |
Income (loss) on equity method investments | | | | | | | $ (3,059,000) | $ (46,936,000) | | | | | | | | |
Deferred revenue | | | | | $ 120,078,000 | | 120,078,000 | 147,900,000 | $ 127,200,000 | | | | | | | |
Revenue remaining performance obligation amount | | | | | 20,700,000 | | 20,700,000 | 35,300,000 | | | | | | | | |
Deferred revenue, net of current portion | | | | | | | | 24,500 | | | | | | | | |
Revenue recognized | | | | | | | 25,500,000 | 16,800,000 | | | | | | | | |
Gain (Loss) on Extinguishment of Debt | | | | | | | | 71,000 | | | | | | | | |
Amyris, Inc. [Member] | Other Income (Expense) [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Gain (Loss) on Extinguishment of Debt | | | | | | | 8,300,000 | 1,600,000 | | | | | | | | |
Amyris, Inc. [Member] | Promissory Note [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Debt instrument, face amount | | | | | 12,000,000 | | $ 12,000,000 | | | | | | | | | |
Debt Instrument, maturity date | | | | | | | Oct. 19, 2022 | | | | | | | | | |
Repayments of debt | | | | | | | $ 9,800,000 | | | | | | | | | |
Synlogic, Inc. [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Deferred revenue | | | | | 100,000 | | 100,000 | 100,000 | | | | | | | | |
Deferred revenue recognized | | | | | | | 100,000 | 100,000 | | | | | | | | |
Revenue remaining performance obligation amount | | | | | | | | | | | | | | $ 30,000,000 | | |
Equity method investments | | | | | 13,700,000 | | 13,700,000 | 16,400,000 | | | | | | | | |
Equity method investments fair value | $ 35,800,000 | | | | | | | | | | | | | | | |
Equity method investment difference between aggregated cost and underlying equity | | | | | | | | | | | | | | 29,800,000 | | |
Equity method investment, aggregate cost | | | | | | | | | | | | | | 80,000,000 | | |
Synlogic, Inc. [Member] | Common Stock [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Equity metohd investment ownership percentage | 19.90% | | | | | | | | | | | | | | | |
Stock issued during period shares | shares | 6,340,771 | | | | | | | | | | | | | | | |
Shares issued price per share | $ / shares | $ 9 | | | | | | | | | | | | | | | |
Stock issued during period, value, new issues | $ 57,100,000 | | | | | | | | | | | | | | | |
Synlogic, Inc. [Member] | Warrant [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Equity method investments | | | | | | | | | | | | | | $ 14,400,000 | | |
Equity method investments fair value | | | | | $ 5,500,000 | | 5,500,000 | 6,600,000 | | | | | | | | |
Change in fair value of warrant liabilities | | | | | | | 1,100,000 | 7,800,000 | | | | | | | | |
Synlogic, Inc. [Member] | Loss on Equity Method Investments [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Income (loss) on equity method investments | | | | | | | $ 2,700,000 | 19,400,000 | | | | | | | | |
Amyris Collaboration Agreement [Member] | Amyris, Inc. [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Due from related parties | | | | $ 800,000 | | | | | | $ 800,000 | | | | | | |
Amyris Collaboration Agreement [Member] | Amyris, Inc. [Member] | Promissory Note [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Due to related parties | | | | 12,000,000 | | | | | | 12,000,000 | | | | | | |
Warrant Agreement [Member] | Synlogic, Inc. [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Number of ordinary share called by each warrant | shares | 2,548,117 | | | | | | | | | | | | | | | |
Exercise price of warrants or rights | $ / shares | $ 9 | | | | | | | | | | | | | | | |
Joyn Bio, LLC [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Equity metohd investment ownership percentage | | | | | 50.00% | | 50.00% | | | | | | | | | |
Equity method investments | | | | 97,900,000 | | | | | | $ 97,900,000 | | | | | | |
Revenue recognized | | | | | | | $ 9,900,000 | 17,100,000 | | | | | | | | |
Joyn Bio, LLC [Member] | Under Termination Scenarios [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Revenue recognized | | | | | | | 7,300,000 | 9,300,000 | | | | | | | | |
Contract with customers liability balance refundable | | | | | $ 9,900,000 | | 9,900,000 | | | | | | | | | |
Cooksonia [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Variable interest entity ownership percentage | | | 70.00% | | | | | | | | | | | | | |
Variable interest entity percentage of control of board of directors | | | 100.00% | | | | | | | | | | | | | |
Equity method investment, underlying equity in net assets | | | $ 8,100,000 | | | | | | | | | | | | | |
Cooksonia [Member] | Parent [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Equity method investments | | | 13,100,000 | | | | | | | | | | | | | |
Payment to acquire equity method investments | | | 5,000,000 | | | | | | | | | | | | | |
Cooksonia [Member] | Joyn Bio, LLC [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Income (loss) on equity method investments | | | | | | | $ 400,000 | 1,700,000 | | | | | | | | |
Equity metohd investment ownership percentage | | | | | 50.00% | | 50.00% | | | | | | | | | |
Equity method investments | | | | | $ 28,900,000 | | $ 28,900,000 | 29,300,000 | | | | | | | | |
Foundry Services Agreement [Member] | Synlogic, Inc. [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Exercise price of warrants or rights | $ / shares | $ 8.99 | | | | | | | | | | | | | | | |
Payments for repurchase of warrants | $ 22,900,000 | | | | | | | | | | | | | | | |
Cash | $ 30,000,000 | | | | | | | | | | | | | | | |
Foundry Services Agreement [Member] | Joyn Bio, LLC [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Upfront payment received | | | | $ 20,000,000 | | | | 15,000,000 | | | | | | | | |
Capital Unit, Class B [Member] | Cooksonia [Member] | Related Party Investor [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Equity metohd investment ownership percentage | | | | 20.00% | | | | | | 20.00% | | | | | | |
Payment to acquire equity method investments | | | | $ 20,000,000 | | | | | | | | | | | | |
Capital Units Class C [Member] | Joyn Bio, LLC [Member] | Bayer [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Equity metohd investment ownership percentage | | | | 50.00% | | | | | | 50.00% | | | | | | |
Payment to acquire equity method investments | | | | $ 20,000,000 | | | | | | | | | | | | |
Commitment to contribute additional capital | | | | $ 60,000,000 | | | | | | $ 60,000,000 | | | | | | |
Capital Units Class C [Member] | Cooksonia [Member] | Joyn Bio, LLC [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Equity metohd investment ownership percentage | | | | 50.00% | | | | | | 50.00% | | | | | | |
Glycosyn Prepaid Services [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Deferred revenue recognized | | | | | | | | 700,000 | | | | | | | | |
Collaborative arrangements value of service applied | | $ 1,000,000 | | | | | | | | | | | | | | |
Collaborative arrangements percentage of every invoice transferrable to service committed | | 25.00% | | | | | | | | | | | | | | |
Collaborative arrangements percentage of every invoice payable | | 75.00% | | | | | | | | | | | | | | |
Percentage of outstanding amount payable upon termination of agreement | | 50.00% | | | | | | | | | | | | | | |
Loans receivable | | | | | 5,400,000 | | 5,400,000 | 5,700,000 | | | | | $ 3,100,000 | | | |
Deferred revenue | | | | | | | | | $ 9,000,000 | | | | | | | |
Elimination of outstanding balance | | | | | | | | 8,400,000 | | | | | | | | |
Unrealized gain losses on loans receivable | | | | | | | | 1,500,000 | | | | | | | | |
Glycosyn Prepaid Services [Member] | Series C Redeemable Preferred Stock [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Equity metohd investment ownership percentage | | | | | | | | | 18.00% | | | | | | | |
Equity method investments | | | | | | | | | $ 10,000,000 | | | | | | | |
Impairment losses on equity method investments | | | | | | | | 1,300,000 | | | | | | | | |
Equity method investments fair value | | | | | | | | 8,500,000 | | | | | | | | |
Letter Contract [Member] | National Institutes of Health [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Revenue remaining performance obligation amount | | | | | | | | | | | | $ 40,500,000 | | | | |
Letter Contract [Member] | Other Income [Member] | National Institutes of Health [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Income recognized on achievement of milestone | | | | | 6,600,000 | | | | | | | | | | | |
Development and Collaboration Agreement [Member] | Octant Inc [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Payments to acquire in process research and development | | | | | | | 5,000,000 | | | | | | | | | |
Payments of profit sharing payments | | | | | | | $ 0 | | | | | | | | | |
Allonnia [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Number of common units received | shares | | | | | | | 3,600,000 | | | | | | | | | |
Carrying value of the equity method investment | | | | | | | $ 24,500,000 | | | | | | | | | |
carrying value outstanding | | | | | | | | 0 | | | | | | | | |
Income (loss) on equity method investments | | | | | | | | 24,500,000 | | | | | | | | |
Allocated upfront non-cash consideration | | | | | | | 2,500,000 | | | | | | | | | |
Deferred revenue | | | | | 26,100,000 | | 26,100,000 | 24,500,000 | | | | | | | | |
Deferred revenue recognized | | | | | | | $ 5,000,000 | | | | | | | | | |
Additional common units | shares | | | | | | 1,867,411 | | | | | 1,867,411 | | | | | |
Total consideration of common unit | | | | | | $ 12,700,000 | | | | | $ 12,700,000 | | | | | |
Number of allocated performance obligations | Obligations | | | | | | | 10 | | | | | | | | | |
Fixed non-cash consideration | | | | | | | $ 2,500,000 | | | | | | | | | |
Variable Interest Entity, Not Primary Beneficiary [Member] | Technical Development Agreement [Member] | Motiff Foodworks Inc [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Investment owned, balance, shares | shares | | | | | | | | | 9,000,900 | | | | | | | |
Equity method investments | | | | | | | | | | | | | | | $ 65,100,000 | |
Impairment losses on equity method investments | | | | | | | | | $ 65,100,000,000 | | | | | | | |
Equity method investments fair value | | | | | 0 | | 0 | 0 | | | | | | | | |
Contract with customers liability | | | | | 54,000,000 | | 54,000,000 | 62,500,000 | | | | | | | | |
Revenue recognized | | | | | | | 20,800,000 | 19,000,000 | | | | | | | | |
Variable Interest Entity, Not Primary Beneficiary [Member] | Technical Development Agreement [Member] | Motiff Foodworks Inc [Member] | Material Rights Tranche One [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Revenue remaining performance obligation amount | | | | | | | | | 6,500,000 | | | | | | | |
Variable Interest Entity, Not Primary Beneficiary [Member] | Technical Development Agreement [Member] | Motiff Foodworks Inc [Member] | Material Rights Tranche Two [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Revenue remaining performance obligation amount | | | | | | | | | 6,500,000 | | | | | | | |
Variable Interest Entity, Not Primary Beneficiary [Member] | Technical Development Agreement [Member] | Motiff Foodworks Inc [Member] | Material Rights Tranche Three [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Revenue remaining performance obligation amount | | | | | | | | | 6,500,000 | | | | | | | |
Variable Interest Entity, Not Primary Beneficiary [Member] | Technical Development Agreement [Member] | Motiff Foodworks Inc [Member] | Material Rights Tranche Four [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Revenue remaining performance obligation amount | | | | | | | | | 6,500,000 | | | | | | | |
Variable Interest Entity, Not Primary Beneficiary [Member] | Technical Development Agreement [Member] | Motiff Foodworks Inc [Member] | Material Rights Tranche Five [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Revenue remaining performance obligation amount | | | | | | | | | 6,500,000 | | | | | | | |
Variable Interest Entity, Not Primary Beneficiary [Member] | Technical Development Agreement [Member] | Motiff Foodworks Inc [Member] | Material Rights Tranche Six [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Revenue remaining performance obligation amount | | | | | | | | | 6,500,000 | | | | | | | |
Variable Interest Entity, Not Primary Beneficiary [Member] | Technical Development Agreement [Member] | Motiff Foodworks Inc [Member] | Material Rights Tranche Seven [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Revenue remaining performance obligation amount | | | | | | | | | 6,500,000 | | | | | | | |
Variable Interest Entity, Not Primary Beneficiary [Member] | Technical Development Agreement [Member] | Motiff Foodworks Inc [Member] | Material Rights Tranche Eight [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Revenue remaining performance obligation amount | | | | | | | | | 6,500,000 | | | | | | | |
Variable Interest Entity, Not Primary Beneficiary [Member] | Technical Development Agreement [Member] | Motiff Foodworks Inc [Member] | Material Rights Tranche Nine [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Revenue remaining performance obligation amount | | | | | | | | | 6,500,000 | | | | | | | |
Variable Interest Entity, Not Primary Beneficiary [Member] | Technical Development Agreement [Member] | Motiff Foodworks Inc [Member] | Material Rights Tranche Ten [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Revenue remaining performance obligation amount | | | | | | | | | 6,500,000 | | | | | | | |
Variable Interest Entity, Not Primary Beneficiary [Member] | Two Thousand And Eighteen Technical Development Agreement [Member] | Genomatica [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Revenue remaining performance obligation amount | | | | | | | | | 40,000,000 | | | | | | | |
Investment owned, at cost | | | | | | | | | $ 40,000,000 | | | | | | | |
Variable Interest Entity, Not Primary Beneficiary [Member] | Foundry Terms Of Service Agreement [Member] | Genomatica [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Contract with customers liability | | | | | 30,100,000 | | 30,100,000 | 38,100,000 | | | | | | | | |
Revenue recognized | | | | | | | 9,400,000 | 6,200,000 | | | | | | | | |
Potential consideration receivable | | | | | | | | | | | | | | | | $ 19,100,000 |
Potential amount payable | | | | | | | | | | | | | | | | $ 4,600,000 |
Upfront amount received | | | | | 6,900,000 | | 6,900,000 | 6,900,000 | | | | | | | | |
Variable Interest Entity, Not Primary Beneficiary [Member] | Foundry Terms Of Service Agreement [Member] | Genomatica [Member] | Preferred Stock [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Equity securities without readily determinable fair value | | | | | $ 55,000,000 | | 55,000,000 | 55,000,000 | | | | | | | | |
Variable Interest Entity, Primary Beneficiary [Member] | Capital Unit, Class A [Member] | Cooksonia [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Payment to acquire equity method investments | | | | $ 5,000,000 | | | | | | | | | | | | |
Variable interest entity ownership percentage | | | | | | | | | | 70.00% | | | | | | |
Noncontrolling Interest [Member] | Cooksonia [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Equity method investments | | | 29,700,000 | | | | | | | | | | | | | |
Equity method investment, underlying equity in net assets | | | $ 8,100,000 | | | | | | | | | | | | | |
Noncontrolling Interest [Member] | Cooksonia [Member] | Joyn Bio, LLC [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Income (loss) on equity method investments | | | | | | | 100,000 | 500,000 | | | | | | | | |
Maximum [Member] | Amyris, Inc. [Member] | Promissory Note [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Debt instrument, periodic payment | | | | | | | 300,000 | | | | | | | | | |
Maximum [Member] | Synlogic, Inc. [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Deferred revenue | | | | | | | | 200,000 | | | | | | | | |
Minimum [Member] | Amyris, Inc. [Member] | Promissory Note [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Debt instrument, periodic payment | | | | | | | $ 200,000 | | | | | | | | | |
Series A Preferred Stock | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Consideration | | | | | | | | $ 33,000,000 | | | | | | | | |
Share Sold | shares | | | | | | | | 2,970,000 | | | | | | | | |
Additional share | shares | | | | | | | | 630,000 | | | | | | | | |
Additional preferred units issued | shares | | | | | 5,400,000 | | 5,400,000 | | | | | | | | | |
Additional common units | shares | | | | | | 1,867,411 | | | | | | | | | | |
Series A Preferred Stock | Allonnia [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Consideration | | | | | | $ 200,000 | | | | | | | | | | |
Share Sold | shares | | | | | | | 1,664,911 | | | | | | | | | |
Number of additional common units received | shares | | | | | | | 5,400,000 | | | | | | | | | |
Additional preferred units issued | shares | | | | | 1,844,911 | | 1,844,911 | | | | | | | | | |
Share Issued In Exchange For The Rights To Certain Intellectual Property | shares | | | | | | | 180,000 | | | | | | | | | |
Series A Preferred Stock | Allonnia [Member] | Arcaea [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Business Combination, Issue | shares | | | | | | 22,500 | | | | | | | | | | |
Class C Units [Member] | Glycosyn Purchase Agreement [Member] | | | | | | | | | | | | | | | | |
Business Acquisition [Line Items] | | | | | | | | | | | | | | | | |
Investment owned, balance, shares | shares | | 80,142 | | | | | | | | | | | | | | |
Investment owned, balance, contracts | | 124.78 | | | | | | | | | | | | | | |
Investment owned, at cost | | $ 10,000,000 | | | | | | | | | | | | | | |
Cash paid upon collaboration agreement | | 1,000,000 | | | | | | | | | | | | | | |
Rights to utilize upon collaboration agreement | | $ 9,000,000 | | | | | | | | | | | | | | |