U.S. Bancorp (USB) 13F-HR Quarterly holdings report by institutional manager

The Securities and Exchange Commission has not necessarily reviewed the information in this filing and has not determined if it is accurate and complete.
The reader should not assume that the information is accurate and complete.

UNITED STATES SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549
FORM 13F

FORM 13F INFORMATION TABLE

OMB APPROVAL

OMB Number:		3235-0006
Expires:	Oct 31, 2018
Estimated average burden
hours per response:		23.8

COLUMN 1	COLUMN 2	COLUMN 3	COLUMN 4	COLUMN 5			COLUMN 6	COLUMN 7	COLUMN 8
			VALUE	SHRS OR	SH/	PUT/	INVESTMENT	OTHER	VOTING AUTHORITY
NAME OF ISSUER	TITLE OF CLASS	CUSIP	(x$1000)	PRN AMT	PRN	CALL	DISCRETION	MANAGER	SOLE	SHARED	NONE
DEUTSCHE BANK AG	COMMON STOCK	D18190898	131	12,054	SH		DFND	1	5,570	0	6,484
DEUTSCHE BANK AG	COMMON STOCK	D18190898	13	1,227	SH		OTR	1	50	888	289
ADIENT PLC	COMMON STOCK	G0084W101	80	2,295	SH		DFND	1	2,271	0	24
ADIENT PLC	COMMON STOCK	G0084W101	10	295	SH		OTR	1	295	0	0
AFYA LTD CLASS A	COMMON STOCK	G01125106	32	1,252	SH		DFND	1	1,252	0	0
ALKERMES PLC	COMMON STOCK	G01767105	36	1,819	SH		DFND	1	1,105	0	714
ALKERMES PLC	COMMON STOCK	G01767105	36	1,801	SH		OTR	1	0	1,743	58
ALLEGION PLC	COMMON STOCK	G0176J109	828	7,116	SH		DFND	1	6,517	0	599
ALLEGION PLC	COMMON STOCK	G0176J109	564	4,847	SH		OTR	1	4,672	0	175
ALLEGION PUBLIC LIMITED COMPNY	COMMON STOCK	G0176J109	11	92	SH		DFND	5	92	0	0
AMCOR PLC	COMMON STOCK	G0250X107	8,240	700,089	SH		DFND	1	700,018	0	71
AMCOR PLC	COMMON STOCK	G0250X107	3,067	260,569	SH		OTR	1	253,347	0	7,222
AMDOCS LTD	COMMON STOCK	G02602103	1,666	23,485	SH		DFND	1	21,863	0	1,622
AMDOCS LTD	COMMON STOCK	G02602103	381	5,376	SH		OTR	1	4,386	958	32
AMDOCS LTD	COMMON STOCK	G02602103	3	43	SH		DFND	5	11	0	32
AMBARELLA INC	COMMON STOCK	G037AX101	233	2,543	SH		DFND	1	2,453	0	90
AMBARELLA INC	COMMON STOCK	G037AX101	61	660	SH		OTR	1	660	0	0
AMBARELLA INC	COMMON STOCK	G037AX101	9	102	SH		DFND	5	102	0	0
AON PLC	COMMON STOCK	G0403H108	1,835	8,685	SH		DFND	1	6,833	0	1,852
AON PLC	COMMON STOCK	G0403H108	3,651	17,280	SH		OTR	1	5,499	619	11,162
AON PLC	COMMON STOCK	G0403H108	198	936	SH		DFND	5	936	0	0
ARCH CAP GROUP LTD	COMMON STOCK	G0450A105	1,375	38,109	SH		DFND	1	30,128	0	7,981
ARCH CAP GROUP LTD	COMMON STOCK	G0450A105	274	7,596	SH		OTR	1	7,504	0	92
ARCH CAPITAL GROUP LTD	COMMON STOCK	G0450A105	2	43	SH		DFND	5	43	0	0
ARCOS DORADOS HOLDINGS INC A	COMMON STOCK	G0457F107	6	1,183	SH		DFND	1	0	0	1,183
ARCOS DORADOS HOLDINGS INC A	COMMON STOCK	G0457F107	5	1,013	SH		OTR	1	1,013	0	0
ARGO GROUP INTL HLDGS LTD	COMMON STOCK	G0464B107	963	22,026	SH		DFND	1	21,426	0	600
ARGO GROUP INTL HLDGS LTD	COMMON STOCK	G0464B107	13	301	SH		OTR	1	301	0	0
ASSURED GUARANTY LTD	COMMON STOCK	G0585R106	70	2,229	SH		DFND	1	715	0	1,514
ATLASSIAN CORP PLC CLASS A	COMMON STOCK	G06242104	2,975	12,720	SH		DFND	1	12,336	0	384
ATLASSIAN CORP PLC CLASS A	COMMON STOCK	G06242104	387	1,655	SH		OTR	1	1,393	262	0
ATLASSIAN CORP PLC	COMMON STOCK	G06242104	137	586	SH		DFND	5	581	0	5
ATHENE HLDG LTD	COMMON STOCK	G0684D107	158	3,661	SH		DFND	1	2,936	0	725
ATHENE HLDG LTD	COMMON STOCK	G0684D107	8	188	SH		OTR	1	0	64	124
AXIS CAPITAL HOLDINGS LTD	COMMON STOCK	G0692U109	4	71	SH		DFND	1	1	0	70
AXALTA COATING SYSTEMS LTD	COMMON STOCK	G0750C108	73	2,557	SH		DFND	1	1,034	0	1,523
AXALTA COATING SYSTEMS LTD	COMMON STOCK	G0750C108	2	69	SH		OTR	1	0	0	69
ATLANTICA SUSTAINABLE INFRASTR	COMMON STOCK	G0751N103	1,743	45,910	SH		DFND	1	45,910	0	0
ATLANTICA SUSTAINABLE INFRASTR	COMMON STOCK	G0751N103	421	11,100	SH		OTR	1	11,000	0	100
BANK OF NT BUTTERFIELD SON	COMMON STOCK	G0772R208	164	5,265	SH		DFND	1	0	0	5,265
BIOHAVEN PHARMACEUTICAL HOLD	COMMON STOCK	G11196105	23	263	SH		DFND	1	250	0	13
BIOHAVEN PHARMACEUTICAL HOLD	COMMON STOCK	G11196105	2	21	SH		OTR	1	0	10	11
ACCENTURE PLC CL A	COMMON STOCK	G1151C101	245,716	940,685	SH		DFND	1	905,086	0	35,599
ACCENTURE PLC CL A	COMMON STOCK	G1151C101	69,049	264,341	SH		OTR	1	236,690	15,986	11,665
ACCENTURE PLC	COMMON STOCK	G1151C101	879	3,363	SH		DFND	5	3,363	0	0
MIMECAST LTD	COMMON STOCK	G14838109	623	10,966	SH		DFND	1	10,966	0	0
MIMECAST LTD	COMMON STOCK	G14838109	255	4,483	SH		OTR	1	4,483	0	0
BROOKFIELD BUSINESS PARTNERS L	COMMON STOCK	G16234109	3	72	SH		DFND	1	72	0	0
BROOKFIELD PROPERTY PARTNERS L	COMMON STOCK	G16249107	57	3,920	SH		DFND	1	3,920	0	0
BROOKFIELD PROPERTY PARTNERS L	COMMON STOCK	G16249107	174	12,000	SH		OTR	1	12,000	0	0
BROOKFIELD INFRASTRUCTURE PART	COMMON STOCK	G16252101	33	662	SH		DFND	1	662	0	0
BROOKFIELD INFRASTRUCTURE PART	COMMON STOCK	G16252101	229	4,642	SH		OTR	1	3,180	0	1,462
BROOKFIELD RENEWABLE PARTNERS	COMMON STOCK	G16258108	273	6,330	SH		DFND	1	4,830	0	1,500
BROOKFIELD RENEWABLE PARTNERS	COMMON STOCK	G16258108	210	4,878	SH		OTR	1	4,878	0	0
BUNGE LIMITED	COMMON STOCK	G16962105	253	3,859	SH		DFND	1	2,735	0	1,124
BUNGE LIMITED	COMMON STOCK	G16962105	2	26	SH		OTR	1	26	0	0
CAPRI HOLDINGS LTD	COMMON STOCK	G1890L107	98	2,339	SH		DFND	1	961	0	1,378
CAPRI HOLDINGS LTD	COMMON STOCK	G1890L107	11	257	SH		OTR	1	0	257	0
CAPRI HOLDINGS LTD	COMMON STOCK	G1890L107	1	18	SH		DFND	5	18	0	0
CHINA YUCHAI INTL LTD	COMMON STOCK	G21082105	117	7,181	SH		DFND	1	759	0	6,422
CIMPRESS PLC	COMMON STOCK	G2143T103	2	25	SH		DFND	1	25	0	0
CIMPRESS PLC	COMMON STOCK	G2143T103	1	13	SH		OTR	1	0	0	13
CHINA BIOLOGIC PRODUCTS HOLD	COMMON STOCK	G21515104	2	13	SH		OTR	1	13	0	0
CLARIVATE PLC	COMMON STOCK	G21810109	282	9,485	SH		OTR	1	9,485	0	0
CLARIVATE ANALYTICS PLC	COMMON STOCK	G21810109	0	7	SH		DFND	5	7	0	0
CREDICORP LTD	COMMON STOCK	G2519Y108	118	721	SH		DFND	1	325	0	396
CREDICORP LTD	COMMON STOCK	G2519Y108	5	29	SH		OTR	1	25	0	4
COSAN LTD CL A	COMMON STOCK	G25343107	39	2,120	SH		DFND	1	2,120	0	0
COCA COLA EUROREAN PARTNERS	COMMON STOCK	G25839104	1,158	23,225	SH		DFND	1	20,998	0	2,227
COCA COLA EUROREAN PARTNERS	COMMON STOCK	G25839104	128	2,569	SH		OTR	1	2,519	50	0
COCA-COLA EUROPEAN PARTNERS	COMMON STOCK	G25839104	29	592	SH		DFND	5	592	0	0
CUSHMAN WAKEFIELD PLC	COMMON STOCK	G2717B108	5	334	SH		DFND	1	187	0	147
CUSHMAN WAKEFIELD PLC	COMMON STOCK	G2717B108	2	140	SH		OTR	1	0	66	74
EATON CORP PLC	COMMON STOCK	G29183103	19,924	165,842	SH		DFND	1	157,278	0	8,564
EATON CORP PLC	COMMON STOCK	G29183103	6,624	55,137	SH		OTR	1	49,392	5,105	640
EATON CORPORATION PLC	COMMON STOCK	G29183103	220	1,829	SH		DFND	5	1,829	0	0
ENDO INTERNATIONAL PLC	COMMON STOCK	G30401106	6	794	SH		DFND	1	297	0	497
ENDO INTERNATIONAL PLC	COMMON STOCK	G30401106	11	1,527	SH		OTR	1	0	1,527	0
ENSTAR GROUP LTD	COMMON STOCK	G3075P101	15	74	SH		DFND	1	37	0	37
ENSTAR GROUP LTD	COMMON STOCK	G3075P101	20	98	SH		OTR	1	98	0	0
ASA GOLD AND PRECIOUS METALS L	OTHER	G3156P103	0	21	SH		DFND	1	21	0	0
ESSENT GROUP LTD	COMMON STOCK	G3198U102	1,739	40,255	SH		DFND	1	40,255	0	0
ESSENT GROUP LTD	COMMON STOCK	G3198U102	62	1,443	SH		OTR	1	1,443	0	0
EVEREST RE GROUP LTD	COMMON STOCK	G3223R108	2,029	8,670	SH		DFND	1	8,509	0	161
EVEREST RE GROUP LTD	COMMON STOCK	G3223R108	102	438	SH		OTR	1	438	0	0
EVEREST RE GROUP LTD	COMMON STOCK	G3223R108	27	115	SH		DFND	5	115	0	0
FABRINET	COMMON STOCK	G3323L100	486	6,261	SH		DFND	1	5,715	0	546
FABRINET	COMMON STOCK	G3323L100	49	634	SH		OTR	1	634	0	0
FRESH DEL MONTE PRODUCE INC	COMMON STOCK	G36738105	296	12,300	SH		DFND	1	12,300	0	0
FRESH DEL MONTE PRODUCE INC	COMMON STOCK	G36738105	63	2,626	SH		OTR	1	2,626	0	0
FRONTLINE LTD	COMMON STOCK	G3682E192	1	85	SH		DFND	1	85	0	0
EROS STX GLOBAL CORPORATION	COMMON STOCK	G3788M114	0	130	SH		DFND	1	130	0	0
GEOPARK LTD	COMMON STOCK	G38327105	3,162	243,396	SH		DFND	1	243,396	0	0
GEOPARK LTD	COMMON STOCK	G38327105	148	11,365	SH		OTR	1	11,365	0	0
GATES INDUSTRIAL CORP PLC	COMMON STOCK	G39108108	20	1,544	SH		DFND	1	0	0	1,544
GATES INDUSTRIAL CORP PLC	COMMON STOCK	G39108108	2	165	SH		OTR	1	0	0	165
GENPACT LIMITED	COMMON STOCK	G3922B107	3,671	88,767	SH		DFND	1	84,208	0	4,559
GENPACT LIMITED	COMMON STOCK	G3922B107	346	8,367	SH		OTR	1	7,100	353	914
GENPACT LTD	COMMON STOCK	G3922B107	92	2,225	SH		DFND	5	2,069	0	156
GOLDEN OCEAN GROUP LTD	COMMON STOCK	G39637205	14	3,078	SH		DFND	1	3,078	0	0
HELEN OF TROY CORP LTD	COMMON STOCK	G4388N106	2,307	10,384	SH		DFND	1	10,357	0	27
HELEN OF TROY CORP LTD	COMMON STOCK	G4388N106	78	351	SH		OTR	1	164	187	0
HERBALIFE NUTRITION LTD	COMMON STOCK	G4412G101	86	1,784	SH		DFND	1	1,387	0	397
HERBALIFE NUTRITION LTD	COMMON STOCK	G4412G101	13	278	SH		OTR	1	0	0	278
JANUS HENDERSON GROUP PLC	COMMON STOCK	G4474Y214	290	8,913	SH		DFND	1	4,137	0	4,776
JANUS HENDERSON GROUP PLC	COMMON STOCK	G4474Y214	2	54	SH		OTR	1	0	0	54
HOLLYSYS AUTOMATION TECHNOLOGI	COMMON STOCK	G45667105	24	1,662	SH		DFND	1	1,662	0	0
HOLLYSYS AUTOMATION TECHNOLOGI	COMMON STOCK	G45667105	6	402	SH		OTR	1	402	0	0
HORIZON THERAPEUTICS PLC	COMMON STOCK	G46188101	2,616	35,752	SH		DFND	1	34,419	0	1,333
HORIZON THERAPEUTICS PLC	COMMON STOCK	G46188101	505	6,908	SH		OTR	1	6,479	396	33
HORIZON THERAPEUTICS PLC	COMMON STOCK	G46188101	73	993	SH		DFND	5	933	0	60
IBEX LTD	COMMON STOCK	G4690M101	9	500	SH		OTR	1	500	0	0
ICON PLC	COMMON STOCK	G4705A100	2,309	11,843	SH		DFND	1	11,438	0	405
ICON PLC	COMMON STOCK	G4705A100	367	1,880	SH		OTR	1	1,596	284	0
ICON PLC	COMMON STOCK	G4705A100	182	934	SH		DFND	5	925	0	9
ICHOR HOLDINGS INC	COMMON STOCK	G4740B105	1	40	SH		DFND	1	40	0	0
ICHOR HOLDINGS INC	COMMON STOCK	G4740B105	1	33	SH		OTR	1	0	33	0
IHS MARKIT LTD	COMMON STOCK	G47567105	3,461	38,524	SH		DFND	1	36,912	0	1,612
IHS MARKIT LTD	COMMON STOCK	G47567105	659	7,341	SH		OTR	1	6,726	306	309
IHS MARKIT LTD	COMMON STOCK	G47567105	442	4,914	SH		DFND	5	4,914	0	0
INTERNATIONAL GAME TECHNOLOG	COMMON STOCK	G4863A108	3	155	SH		DFND	1	0	0	155
INTERNATIONAL GAME TECHNOLOGY	COMMON STOCK	G4863A108	0	3	SH		DFND	5	3	0	0
INVESCO LTD	COMMON STOCK	G491BT108	168	9,650	SH		DFND	1	4,157	0	5,493
INVESCO LTD	COMMON STOCK	G491BT108	31	1,781	SH		OTR	1	914	115	752
JAMES RIVER GROUP HOLDINGS L	COMMON STOCK	G5005R107	8	161	SH		DFND	1	161	0	0
JAZZ PHARMACEUTICALS PLC	COMMON STOCK	G50871105	412	2,495	SH		DFND	1	2,348	0	147
JAZZ PHARMACEUTICALS PLC	COMMON STOCK	G50871105	160	966	SH		OTR	1	839	43	84
JAZZ PHARMACEUTICALS PLC	COMMON STOCK	G50871105	66	397	SH		DFND	5	397	0	0
JOHNSON CTLS INTL PLC	COMMON STOCK	G51502105	1,855	39,802	SH		DFND	1	34,246	0	5,556
JOHNSON CTLS INTL PLC	COMMON STOCK	G51502105	410	8,789	SH		OTR	1	7,980	541	268
JOHNSON CONTROLS INTERNATIONAL	COMMON STOCK	G51502105	51	1,090	SH		DFND	5	638	0	452
LAZARD LTD CL A	COMMON STOCK	G54050102	34	797	SH		DFND	1	497	0	300
LIBERTY GLOBAL PLC A	COMMON STOCK	G5480U104	10	429	SH		DFND	1	429	0	0
LIBERTY GLOBAL PLC SERIES C	COMMON STOCK	G5480U120	110	4,663	SH		DFND	1	4,663	0	0
LINDE PLC	COMMON STOCK	G5494J103	38,121	144,664	SH		DFND	1	139,919	0	4,745
LINDE PLC	COMMON STOCK	G5494J103	15,479	58,740	SH		OTR	1	50,006	2,179	6,555
LINDE PLC	COMMON STOCK	G5494J103	41	156	SH		DFND	5	147	0	9
LIVANOVA PLC	COMMON STOCK	G5509L101	38	579	SH		DFND	1	414	0	165
LIVANOVA PLC	COMMON STOCK	G5509L101	53	807	SH		OTR	1	15	792	0
LUXFER HOLDINGS PLC	COMMON STOCK	G5698W116	2	150	SH		DFND	1	82	0	68
MANCHESTER UTD PLCCL A	COMMON STOCK	G5784H106	22	1,311	SH		OTR	1	0	1,311	0
MARVELL TECHNOLOGY GROUP LTD	COMMON STOCK	G5876H105	9,739	204,868	SH		DFND	1	201,991	0	2,877
MARVELL TECHNOLOGY GROUP LTD	COMMON STOCK	G5876H105	1,067	22,443	SH		OTR	1	21,665	251	527
MARVELL TECHNOLOGY GROUP LTD	COMMON STOCK	G5876H105	5	112	SH		DFND	5	112	0	0
MEDTRONIC PLC	COMMON STOCK	G5960L103	101,578	867,147	SH		DFND	1	840,641	0	26,506
MEDTRONIC PLC	COMMON STOCK	G5960L103	42,602	363,684	SH		OTR	1	337,861	7,212	18,611
MEDTRONIC PLC	COMMON STOCK	G5960L103	550	4,693	SH		DFND	5	4,693	0	0
APTIV PLC	COMMON STOCK	G6095L109	27,064	207,722	SH		DFND	1	203,283	0	4,439
APTIV PLC	COMMON STOCK	G6095L109	3,370	25,865	SH		OTR	1	25,116	502	247
APTIV PLC	COMMON STOCK	G6095L109	273	2,093	SH		DFND	5	2,093	0	0
NABORS INDUSTRIES LTD SHS	COMMON STOCK	G6359F137	1	18	SH		DFND	1	0	0	18
NABRIVA THERAPEUTICS PLC SHS N	COMMON STOCK	G63637113	0	50	SH		DFND	1	50	0	0
NATIONAL ENERGY SERVICES REU	COMMON STOCK	G6375R107	29	2,871	SH		DFND	1	792	0	2,079
MYOVANT SCIENCES LTD	COMMON STOCK	G637AM102	11	393	SH		DFND	1	69	0	324
NIELSEN HOLDINGS PLC	COMMON STOCK	G6518L108	72	3,462	SH		DFND	1	1,003	0	2,459
NOMAD FOODS LTD	COMMON STOCK	G6564A105	4,053	159,447	SH		DFND	1	159,447	0	0
NOMAD FOODS LTD	COMMON STOCK	G6564A105	906	35,641	SH		OTR	1	35,641	0	0
NORDIC AMERICAN TANKERS LTD	COMMON STOCK	G65773106	14	4,883	SH		DFND	1	4,162	0	721
NORDIC AMERICAN TANKERS LTD	COMMON STOCK	G65773106	8	2,674	SH		OTR	1	0	2,674	0
NORWEGIAN CRUISE LINE HOLDINGS	COMMON STOCK	G66721104	428	16,835	SH		DFND	1	15,651	0	1,184
NORWEGIAN CRUISE LINE HOLDINGS	COMMON STOCK	G66721104	18	712	SH		OTR	1	0	150	562
NOVOCURE LTD	COMMON STOCK	G6674U108	101	585	SH		DFND	1	562	0	23
NOVOCURE LTD	COMMON STOCK	G6674U108	375	2,171	SH		OTR	1	337	564	1,270
NVENT ELECTRIC PLC	COMMON STOCK	G6700G107	5,548	238,200	SH		DFND	1	236,279	0	1,921
NVENT ELECTRIC PLC	COMMON STOCK	G6700G107	8,677	372,564	SH		OTR	1	361,563	934	10,067
OXFORD IMMUNOTEC GLOBAL PLC	COMMON STOCK	G6855A103	17	965	SH		DFND	1	965	0	0
PAGSEGURO DIGITAL LTD CL A	COMMON STOCK	G68707101	201	3,536	SH		DFND	1	3,536	0	0
PAGSEGURO DIGITAL LTD CL A	COMMON STOCK	G68707101	58	1,021	SH		OTR	1	1,021	0	0
PROTHENA CORP PLC	COMMON STOCK	G72800108	14	1,160	SH		DFND	1	1,160	0	0
RENAISSANCE RE HOLDINGS LTD	COMMON STOCK	G7496G103	1,271	7,668	SH		DFND	1	7,512	0	156
RENAISSANCE RE HOLDINGS LTD	COMMON STOCK	G7496G103	326	1,963	SH		OTR	1	1,856	0	107
RENAISSANCERE HOLDINGS LTD	COMMON STOCK	G7496G103	34	203	SH		DFND	5	203	0	0
ROYALTY PHARMA PLC SHS CL A	COMMON STOCK	G7709Q104	45	905	SH		DFND	1	905	0	0
SFL CORP LTD	COMMON STOCK	G7738W106	13	2,000	SH		DFND	1	2,000	0	0
SEAGATE TECHNOLOGY	COMMON STOCK	G7945M107	646	10,402	SH		DFND	1	9,026	0	1,376
SEAGATE TECHNOLOGY	COMMON STOCK	G7945M107	222	3,568	SH		OTR	1	3,118	0	450
SEAGATE TECHNOLOGY	COMMON STOCK	G7945M107	1	0	SH		DFND	5	0	0	0
PENTAIR PLC	COMMON STOCK	G7S00T104	13,647	257,053	SH		DFND	1	255,640	0	1,413
PENTAIR PLC	COMMON STOCK	G7S00T104	11,953	225,142	SH		OTR	1	209,499	5,858	9,785
PENTAIR PLC	COMMON STOCK	G7S00T104	9	171	SH		DFND	5	171	0	0
SENSATA TECHNOLOGIES HOLDING	COMMON STOCK	G8060N102	1,723	32,658	SH		DFND	1	31,232	0	1,426
SENSATA TECHNOLOGIES HOLDING	COMMON STOCK	G8060N102	469	8,887	SH		OTR	1	8,734	100	53
SENSATA TECHNOLOGIES HLDG PLC	COMMON STOCK	G8060N102	57	1,078	SH		DFND	5	1,078	0	0
SIGNET JEWELERS LTD	COMMON STOCK	G81276100	432	15,836	SH		DFND	1	13,992	0	1,844
SINA CORP	COMMON STOCK	G81477104	159	3,741	SH		DFND	1	1,874	0	1,867
STERIS PLC	COMMON STOCK	G8473T100	4,112	21,693	SH		DFND	1	19,080	0	2,613
STERIS PLC	COMMON STOCK	G8473T100	860	4,538	SH		OTR	1	3,592	946	0
STERIS PLC	COMMON STOCK	G8473T100	344	1,811	SH		DFND	5	1,798	0	13
STONECO LTD A	COMMON STOCK	G85158106	1,312	15,632	SH		DFND	1	15,611	0	21
STONECO LTD A	COMMON STOCK	G85158106	1,020	12,155	SH		OTR	1	12,155	0	0
STONECO LTD	COMMON STOCK	G85158106	15	180	SH		DFND	5	156	0	24
TECHNIPFMC PLC	COMMON STOCK	G87110105	136	14,526	SH		DFND	1	9,918	0	4,608
TECHNIPFMC PLC	COMMON STOCK	G87110105	10	1,068	SH		OTR	1	923	0	145
TEXTAINER GROUP HOLDINGS LTD	COMMON STOCK	G8766E109	797	41,557	SH		DFND	1	41,557	0	0
THERAVANCE BIOPHARMA INC	COMMON STOCK	G8807B106	2	116	SH		DFND	1	116	0	0
TRANE TECHNOLOGIES PLC	COMMON STOCK	G8994E103	11,005	75,816	SH		DFND	1	72,914	0	2,902
TRANE TECHNOLOGIES PLC	COMMON STOCK	G8994E103	3,007	20,709	SH		OTR	1	16,512	720	3,477
TRANE TECHNOLOGIES PLC	COMMON STOCK	G8994E103	89	615	SH		DFND	5	615	0	0
LIBERTY LATIN AMERICA LTD	COMMON STOCK	G9001E102	6	526	SH		DFND	1	485	0	41
LIBERTY LATIN AMERICA LTD	COMMON STOCK	G9001E128	1	47	SH		DFND	1	47	0	0
TRITON INTERNATIONAL LTD BER	COMMON STOCK	G9078F107	342	7,036	SH		DFND	1	6,690	0	346
TRITON INTERNATIONAL LTD BER	COMMON STOCK	G9078F107	3	68	SH		OTR	1	0	68	0
TRONOX HOLDINGS PLC CLASS A	COMMON STOCK	G9087Q102	14	974	SH		DFND	1	686	0	288
TSAKOS ENERGY NAVIGATION LTD S	COMMON STOCK	G9108L173	96	12,000	SH		DFND	1	12,000	0	0
GOLAR LNG LTD	COMMON STOCK	G9456A100	109	11,345	SH		DFND	1	11,345	0	0
WATFORD HOLDINGS LTD	COMMON STOCK	G94787101	1	31	SH		DFND	1	0	0	31
WHITE MTNS INS GROUP LTD	COMMON STOCK	G9618E107	16	16	SH		DFND	1	8	0	8
WHITE MTNS INS GROUP LTD	COMMON STOCK	G9618E107	48	48	SH		OTR	1	0	15	33
WHITE MOUNTAINS INSURANCE GP	COMMON STOCK	G9618E107	30	30	SH		DFND	5	30	0	0
WILLIS TOWERS WATSON PLC	COMMON STOCK	G96629103	3,095	14,693	SH		DFND	1	13,830	0	863
WILLIS TOWERS WATSON PLC	COMMON STOCK	G96629103	208	990	SH		OTR	1	961	10	19
PERRIGO CO PLC	COMMON STOCK	G97822103	602	13,457	SH		DFND	1	13,051	0	406
PERRIGO CO PLC	COMMON STOCK	G97822103	6	139	SH		OTR	1	111	0	28
ALCON INC	COMMON STOCK	H01301128	7,382	111,895	SH		DFND	1	108,147	0	3,748
ALCON INC	COMMON STOCK	H01301128	1,930	29,255	SH		OTR	1	23,736	1,028	4,491
ALCON INC	COMMON STOCK	H01301128	814	12,332	SH		DFND	5	12,313	0	19
CHUBB LTD	COMMON STOCK	H1467J104	121,276	787,915	SH		DFND	1	779,409	0	8,506
CHUBB LTD	COMMON STOCK	H1467J104	26,042	169,184	SH		OTR	1	153,408	7,379	8,397
CHUBB LTD	COMMON STOCK	H1467J104	225	1,464	SH		DFND	5	1,464	0	0
CRISPR THERAPEUTICS AG	COMMON STOCK	H17182108	178	1,162	SH		DFND	1	1,162	0	0
CRISPR THERAPEUTICS AG	COMMON STOCK	H17182108	15	95	SH		OTR	1	95	0	0
GARMIN LTD	COMMON STOCK	H2906T109	1,936	16,181	SH		DFND	1	15,215	0	966
GARMIN LTD	COMMON STOCK	H2906T109	106	890	SH		OTR	1	841	44	5
UBS GROUP AG	COMMON STOCK	H42097107	632	44,731	SH		DFND	1	24,390	0	20,341
UBS GROUP AG	COMMON STOCK	H42097107	56	3,992	SH		OTR	1	512	1,045	2,435
UBS GROUP AG	COMMON STOCK	H42097107	6	425	SH		DFND	5	425	0	0
LOGITECH INTL SA	COMMON STOCK	H50430232	396	4,069	SH		DFND	1	2,255	0	1,814
LOGITECH INTL SA	COMMON STOCK	H50430232	129	1,330	SH		OTR	1	192	11	1,127
TE CONNECTIVITY LTD	COMMON STOCK	H84989104	5,032	41,565	SH		DFND	1	25,690	0	15,875
TE CONNECTIVITY LTD	COMMON STOCK	H84989104	1,443	11,920	SH		OTR	1	11,868	46	6
TE CONNECTIVITY LTD	COMMON STOCK	H84989104	68	560	SH		DFND	5	560	0	0
TRANSOCEAN LTD	COMMON STOCK	H8817H100	2	1,010	SH		DFND	1	1,010	0	0
TRANSOCEAN LTD	COMMON STOCK	H8817H100	1	438	SH		OTR	1	438	0	0
ADECOAGRO S A	COMMON STOCK	L00849106	1	210	SH		DFND	1	0	0	210
ARDAGH GROUP S A	COMMON STOCK	L0223L101	3	170	SH		DFND	1	170	0	0
ARDAGH FINANCE HOLDINGS SA	COMMON STOCK	L0223L101	1	35	SH		DFND	5	35	0	0
GLOBANT SA	COMMON STOCK	L44385109	2,411	11,078	SH		DFND	1	10,988	0	90
GLOBANT SA	COMMON STOCK	L44385109	303	1,392	SH		OTR	1	1,392	0	0
GLOBANT S A	COMMON STOCK	L44385109	441	2,027	SH		DFND	5	2,027	0	0
MILLICOM INTL CELLULAR SA	COMMON STOCK	L6388F110	16	400	SH		DFND	1	400	0	0
ORION ENGINEERED CARBONS SA	COMMON STOCK	L72967109	23	1,369	SH		DFND	1	290	0	1,079
ORION ENGINEERED CARBONS SA	COMMON STOCK	L72967109	9	540	SH		OTR	1	0	540	0
SPOTIFY TECHNOLOGY SA	COMMON STOCK	L8681T102	5,363	17,045	SH		DFND	1	16,717	0	328
SPOTIFY TECHNOLOGY SA	COMMON STOCK	L8681T102	974	3,094	SH		OTR	1	1,066	2,028	0
SPOTIFY TECHNOLOGY SA	COMMON STOCK	L8681T102	19	61	SH		DFND	5	48	0	13
TRINSEO S A	COMMON STOCK	L9340P101	48	935	SH		DFND	1	352	0	583
TRINSEO S A	COMMON STOCK	L9340P101	8	157	SH		OTR	1	102	55	0
ALLOT LTD	COMMON STOCK	M0854Q105	36	3,358	SH		DFND	1	3,358	0	0
ALLOT LTD	COMMON STOCK	M0854Q105	4	333	SH		OTR	1	0	333	0
AUDIOCODES LTD	COMMON STOCK	M15342104	90	3,280	SH		DFND	1	3,280	0	0
AUDIOCODES LTD	COMMON STOCK	M15342104	44	1,587	SH		OTR	1	1,587	0	0
AUDIOCODES LTD	COMMON STOCK	M15342104	9	311	SH		DFND	5	311	0	0
CAESAR STONE SDOT YAM LTD	COMMON STOCK	M20598104	0	31	SH		DFND	5	31	0	0
CERAGON NETWORKS LTD	COMMON STOCK	M22013102	56	20,000	SH		DFND	1	20,000	0	0
CHECK POINT SOFTWARE TECH LTD	COMMON STOCK	M22465104	11,130	83,747	SH		DFND	1	82,910	0	837
CHECK POINT SOFTWARE TECH LTD	COMMON STOCK	M22465104	3,388	25,491	SH		OTR	1	24,752	334	405
CHECK POINT SOFTWARE TECH	COMMON STOCK	M22465104	33	252	SH		DFND	5	252	0	0
CYBERARK SOFTWARE LTD ISRAEL	COMMON STOCK	M2682V108	1,132	7,008	SH		DFND	1	6,728	0	280
CYBERARK SOFTWARE LTD ISRAEL	COMMON STOCK	M2682V108	371	2,292	SH		OTR	1	2,177	106	9
ELBIT SYSTEMS LTD	COMMON STOCK	M3760D101	499	3,813	SH		DFND	1	3,372	0	441
ELBIT SYSTEMS LTD	COMMON STOCK	M3760D101	140	1,071	SH		OTR	1	621	0	450
GAMIDA CELL LTD	COMMON STOCK	M47364100	10	1,204	SH		DFND	1	1,204	0	0
FIVERR INTERNATIONAL LTD	COMMON STOCK	M4R82T106	10	50	SH		DFND	1	50	0	0
FIVERR INTERNATIONAL LTD	COMMON STOCK	M4R82T106	15	75	SH		OTR	1	75	0	0
GILAT SATELLITE NETWORKS LTD	COMMON STOCK	M51474118	7	1,000	SH		DFND	1	1,000	0	0
GILAT SATELLITE NETWORKS LTD	COMMON STOCK	M51474118	4	539	SH		OTR	1	0	539	0
ICL GROUP LTD	COMMON STOCK	M53213100	93	18,555	SH		DFND	1	8,343	0	10,212
ICL GROUP LTD	COMMON STOCK	M53213100	10	2,058	SH		OTR	1	0	2,058	0
INMODE LTD	COMMON STOCK	M5425M103	38	797	SH		DFND	1	797	0	0
INMODE LTD	COMMON STOCK	M5425M103	6	118	SH		OTR	1	118	0	0
JFROG LTD	COMMON STOCK	M6191J100	124	1,971	SH		DFND	1	1,971	0	0
KAMADA LTD	COMMON STOCK	M6240T109	32	4,875	SH		OTR	1	0	4,875	0
KORNIT DIGITAL LTD	COMMON STOCK	M6372Q113	1,081	12,129	SH		DFND	1	12,129	0	0
KORNIT DIGITAL LTD	COMMON STOCK	M6372Q113	110	1,229	SH		OTR	1	1,086	143	0
KORNIT DIGITAL LTD	COMMON STOCK	M6372Q113	290	3,251	SH		DFND	5	3,251	0	0
MAGAL SECURITY SYS LTD	COMMON STOCK	M6786D104	386	99,527	SH		DFND	1	99,527	0	0
NANO X IMAGING LTD	COMMON STOCK	M70700105	11	250	SH		DFND	1	250	0	0
NOVA MEASURING INSTRUMENTS LTD	COMMON STOCK	M7516K103	10	145	SH		DFND	1	0	0	145
NOVA MEASURING INSTRUMENTS LTD	COMMON STOCK	M7516K103	85	1,199	SH		OTR	1	0	1,199	0
NOVA MEASURING INSTRUMENTS LTD	COMMON STOCK	M7516K103	2	29	SH		DFND	5	29	0	0
RADWARE LTD	COMMON STOCK	M81873107	124	4,453	SH		DFND	1	3,959	0	494
RADWARE LTD	COMMON STOCK	M81873107	62	2,247	SH		OTR	1	2,129	118	0
REWALK ROBOTICS LTD	COMMON STOCK	M8216Q200	1	500	SH		DFND	1	500	0	0
STRATASYS LTD	COMMON STOCK	M85548101	4	200	SH		DFND	1	200	0	0
STRATASYS LTD	COMMON STOCK	M85548101	2	100	SH		OTR	1	100	0	0
TARO PHARMACEUTICAL INDUS	COMMON STOCK	M8737E108	646	8,804	SH		DFND	1	8,289	0	515
TARO PHARMACEUTICAL INDUS	COMMON STOCK	M8737E108	193	2,634	SH		OTR	1	2,522	112	0
TOWER SEMICONDUCTOR LTD	COMMON STOCK	M87915274	382	14,810	SH		DFND	1	14,757	0	53
TOWER SEMICONDUCTOR LTD	COMMON STOCK	M87915274	60	2,330	SH		OTR	1	2,330	0	0
TOWER SEMICONDUCTOR LTD	COMMON STOCK	M87915274	1	31	SH		DFND	5	31	0	0
UROGEN PHARMA LTD	COMMON STOCK	M96088105	28	1,538	SH		DFND	1	138	0	1,400
UROGEN PHARMA LTD	COMMON STOCK	M96088105	7	372	SH		OTR	1	0	372	0
WIX COM LTD	COMMON STOCK	M98068105	457	1,827	SH		DFND	1	1,514	0	313
WIX COM LTD	COMMON STOCK	M98068105	38	154	SH		OTR	1	92	49	13
AERCAP HOLDINGS NV	COMMON STOCK	N00985106	1,161	25,479	SH		DFND	1	24,052	0	1,427
AERCAP HOLDINGS NV	COMMON STOCK	N00985106	292	6,401	SH		OTR	1	6,065	288	48
AERCAP HOLDINGS NV	COMMON STOCK	N00985106	9	190	SH		DFND	5	190	0	0
AFFIMED NV	COMMON STOCK	N01045108	140	24,000	SH		OTR	1	24,000	0	0
ASML HOLDING NV NY REG SHS	COMMON STOCK	N07059210	5,881	12,059	SH		DFND	1	9,931	0	2,128
ASML HOLDING NV NY REG SHS	COMMON STOCK	N07059210	1,458	2,988	SH		OTR	1	2,287	674	27
ASML HOLDING N V	COMMON STOCK	N07059210	752	1,541	SH		DFND	5	1,536	0	5
ELASTIC NV	COMMON STOCK	N14506104	793	5,430	SH		DFND	1	5,324	0	106
ELASTIC NV	COMMON STOCK	N14506104	180	1,232	SH		OTR	1	1,147	66	19
ELASTIC NV	COMMON STOCK	N14506104	14	98	SH		DFND	5	82	0	16
CNH INDUSTRIAL NV	COMMON STOCK	N20944109	174	13,522	SH		DFND	1	10,311	0	3,211
CNH INDUSTRIAL NV	COMMON STOCK	N20944109	38	2,932	SH		OTR	1	2,932	0	0
CNH INDUSTRIAL NV	COMMON STOCK	N20944109	33	2,571	SH		DFND	5	2,571	0	0
CORE LABORATORIES N V	COMMON STOCK	N22717107	141	5,319	SH		DFND	1	4,246	0	1,073
CORE LABORATORIES N V	COMMON STOCK	N22717107	15	557	SH		OTR	1	170	96	291
FERRARI NV	COMMON STOCK	N3167Y103	2,463	10,730	SH		DFND	1	9,741	0	989
FERRARI NV	COMMON STOCK	N3167Y103	389	1,698	SH		OTR	1	1,670	0	28
FERRARI NV	COMMON STOCK	N3167Y103	666	2,902	SH		DFND	5	2,902	0	0
FIAT CHRYSLER AUTOMOBILES NV	COMMON STOCK	N31738102	323	17,850	SH		DFND	1	12,045	0	5,805
FIAT CHRYSLER AUTOMOBILES NV	COMMON STOCK	N31738102	35	1,926	SH		OTR	1	0	1,666	260
FRANK S INTERNATINAL NV	COMMON STOCK	N33462107	3	1,030	SH		DFND	1	1,030	0	0
LYONDELLBASELL INDU CL A	COMMON STOCK	N53745100	36,055	393,354	SH		DFND	1	386,030	0	7,324
LYONDELLBASELL INDU CL A	COMMON STOCK	N53745100	4,899	53,449	SH		OTR	1	49,209	470	3,770
LYONDELLBASELL INDUSTRIES NV	COMMON STOCK	N53745100	2	25	SH		DFND	5	1	0	24
NXP SEMICONDUCTORS NV	COMMON STOCK	N6596X109	5,499	34,582	SH		DFND	1	33,771	0	811
NXP SEMICONDUCTORS NV	COMMON STOCK	N6596X109	534	3,360	SH		OTR	1	2,889	471	0
NXP SEMICONDUTORS NV	COMMON STOCK	N6596X109	463	2,910	SH		DFND	5	2,836	0	74
PLAYA HOTELS RESORTS NV	COMMON STOCK	N70544106	3	533	SH		DFND	1	182	0	351
QIAGEN N V	COMMON STOCK	N72482123	2,148	40,648	SH		DFND	1	37,041	0	3,607
QIAGEN N V	COMMON STOCK	N72482123	998	18,898	SH		OTR	1	18,427	395	76
QIAGEN N.V.	COMMON STOCK	N72482123	2	40	SH		DFND	5	40	0	0
UNIQURE BV	COMMON STOCK	N90064101	6	155	SH		DFND	1	0	0	155
YANDEX NV	COMMON STOCK	N97284108	4,351	62,521	SH		DFND	1	62,429	0	92
YANDEX NV	COMMON STOCK	N97284108	747	10,729	SH		OTR	1	10,493	236	0
YANDEX NV	COMMON STOCK	N97284108	14	195	SH		DFND	5	195	0	0
COPA HOLDINGS SA CL A	COMMON STOCK	P31076105	68	885	SH		DFND	1	859	0	26
ONESPAWORLD HOLDINGS LTD	COMMON STOCK	P73684113	1	122	SH		DFND	1	0	0	122
ONESPAWORLD HOLDINGS LTD	COMMON STOCK	P73684113	31	3,092	SH		OTR	1	0	3,092	0
MAKEMYTRIP LTD	COMMON STOCK	V5633W109	121	4,106	SH		DFND	1	4,106	0	0
ROYAL CARIBBEAN CRUISES LTD	COMMON STOCK	V7780T103	1,132	15,162	SH		DFND	1	12,288	0	2,874
ROYAL CARIBBEAN CRUISES LTD	COMMON STOCK	V7780T103	285	3,813	SH		OTR	1	3,244	417	152
ROYAL CARIBBEAN CRUISES LTD	COMMON STOCK	V7780T103	2	23	SH		DFND	5	23	0	0
ATLAS CORP	COMMON STOCK	Y0436Q109	6	568	SH		DFND	1	476	0	92
ATLAS CORP	COMMON STOCK	Y0436Q109	4	330	SH		DFND	5	330	0	0
COSTAMARE INC	COMMON STOCK	Y1771G102	35	4,215	SH		DFND	1	0	0	4,215
DHT HOLDINGS INC	COMMON STOCK	Y2065G121	8	1,436	SH		DFND	1	1,436	0	0
DIANA SHIPPING INC	COMMON STOCK	Y2066G104	5	2,704	SH		DFND	1	2,704	0	0
FLEX LTD	COMMON STOCK	Y2573F102	380	21,151	SH		DFND	1	19,910	0	1,241
FLEX LTD	COMMON STOCK	Y2573F102	13	696	SH		OTR	1	696	0	0
FLEXTRONICS INTL LTD	COMMON STOCK	Y2573F102	5	282	SH		DFND	5	282	0	0
GASLOG PARTNERS LP	COMMON STOCK	Y2687W108	3	1,000	SH		OTR	1	1,000	0	0
INTERNATIONAL SEAWAYS INC WI	COMMON STOCK	Y41053102	2	135	SH		DFND	1	0	0	135
MAXEON SOLAR TECHNOLOGIES LTD	COMMON STOCK	Y58473102	1	52	SH		DFND	1	0	0	52
MAXEON SOLAR TECHNOLOGIES LTD	COMMON STOCK	Y58473102	7	241	SH		OTR	1	0	241	0
SCORPIO TANKERS INC	COMMON STOCK	Y7542C130	11	997	SH		DFND	1	270	0	727
SCORPIO BULKERS INC	COMMON STOCK	Y7546A130	5	270	SH		DFND	1	270	0	0
AAON INC	COMMON STOCK	000360206	3,046	45,717	SH		DFND	1	43,174	0	2,543
AAON INC	COMMON STOCK	000360206	536	8,042	SH		OTR	1	5,132	0	2,910
A A R CORP	COMMON STOCK	000361105	301	8,330	SH		DFND	1	8,287	0	43
A A R CORP	COMMON STOCK	000361105	7	187	SH		OTR	1	187	0	0
ABB LTD A D R	COMMON STOCK	000375204	2,780	99,452	SH		DFND	1	94,137	0	5,315
ABB LTD A D R	COMMON STOCK	000375204	672	24,013	SH		OTR	1	23,464	484	65
ABB LTD	COMMON STOCK	000375204	4	128	SH		DFND	5	128	0	0
ABCAM PLC	COMMON STOCK	000380204	4	183	SH		DFND	5	183	0	0
ABM INDUSTRIES INC	COMMON STOCK	000957100	8	213	SH		DFND	5	213	0	0
AFLAC INC	COMMON STOCK	001055102	8,561	192,513	SH		DFND	1	184,534	0	7,979
AFLAC INC	COMMON STOCK	001055102	2,901	65,251	SH		OTR	1	64,115	823	313
AFLAC INC	COMMON STOCK	001055102	26	586	SH		DFND	5	586	0	0
AGCO CORP	COMMON STOCK	001084102	160	1,550	SH		DFND	1	1,209	0	341
AG MORTGAGE INVESTMENT TRUST	COMMON STOCK	001228105	2	600	SH		OTR	1	600	0	0
AMN HEALTHCARE SVCS INC	COMMON STOCK	001744101	1,789	26,213	SH		DFND	1	25,755	0	458
AMN HEALTHCARE SVCS INC	COMMON STOCK	001744101	229	3,348	SH		OTR	1	2,725	623	0
AMN HEALTHCARE SERVICES INC	COMMON STOCK	001744101	220	3,217	SH		DFND	5	3,217	0	0
A10 NETWORKS INC	COMMON STOCK	002121101	165	16,749	SH		DFND	1	16,749	0	0
A10 NETWORKS INC	COMMON STOCK	002121101	36	3,628	SH		OTR	1	3,628	0	0
AZZ INC	COMMON STOCK	002474104	20	429	SH		DFND	1	320	0	109
ABBOTT LABORATORIES	COMMON STOCK	002824100	237,925	2,173,031	SH		DFND	1	2,111,458	0	61,573
ABBOTT LABORATORIES	COMMON STOCK	002824100	94,990	867,571	SH		OTR	1	816,957	31,362	19,252
ABBOTT LABORATORIES	COMMON STOCK	002824100	269	2,457	SH		DFND	5	2,457	0	0
ABBOTT LABORATORIES	COMMON STOCK	002824100	0	5	SH	Put	DFND	1	5	0	0
ABBOTT LABORATORIES	COMMON STOCK	002824100	3	14	SH	Put	OTR	1	14	0	0
ABERDEEN ASIA PACIFIC PRIME IN	OTHER	003009107	689	154,900	SH		DFND	1	10,000	0	144,900
ABERDEEN ASIA PACIFIC PRIME IN	OTHER	003009107	134	30,000	SH		OTR	1	0	30,000	0
ABERDEEN AUSTRALIA EQUITY FUND	OTHER	003011103	28	5,356	SH		DFND	1	5,356	0	0
ABERDEEN AUSTRALIA EQUITY FUND	OTHER	003011103	149	28,238	SH		OTR	1	28,238	0	0
ABERDEEN STANDARD PHYSICAL P E	OTHER	003262102	107	468	SH		OTR	1	0	0	468
ABIOMED INC	COMMON STOCK	003654100	872	2,689	SH		DFND	1	2,303	0	386
ABIOMED INC	COMMON STOCK	003654100	507	1,564	SH		OTR	1	1,516	38	10
ABIOMED INC	COMMON STOCK	003654100	3	10	SH		DFND	5	10	0	0
ACADIA PHARMACEUTICALS INC	COMMON STOCK	004225108	194	3,637	SH		DFND	1	3,293	0	344
ACADIA PHARMACEUTICALS INC	COMMON STOCK	004225108	8	166	SH		OTR	1	121	45	0
ACADIA RLTY TR	COMMON STOCK	004239109	34	2,428	SH		DFND	1	1,323	0	1,105
ACCURAY INC	COMMON STOCK	004397105	6	1,500	SH		DFND	1	1,500	0	0
ACI WORLDWIDE INC	COMMON STOCK	004498101	9	243	SH		DFND	1	243	0	0
ACI WORLDWIDE INC	COMMON STOCK	004498101	335	8,724	SH		OTR	1	260	0	8,464
ACUSHNET HOLDINGS CORP	COMMON STOCK	005098108	13	329	SH		DFND	1	152	0	177
ADAMS DIVERSIFIED EQUITY FUND	OTHER	006212104	35	2,033	SH		DFND	1	2,033	0	0
ADDUS HOMECARE CORP	COMMON STOCK	006739106	3	26	SH		DFND	1	20	0	6
ADDUS HOMECARE CORP	COMMON STOCK	006739106	86	735	SH		OTR	1	0	735	0
ADDVANTAGE TECHNOLOGIES GRP	COMMON STOCK	006743306	1	250	SH		DFND	1	250	0	0
AEROJET ROCKETDYNE HOLDINGS	COMMON STOCK	007800105	35	666	SH		DFND	1	666	0	0
AEROJET ROCKETDYNE HOLDINGS	COMMON STOCK	007800105	43	812	SH		OTR	1	0	812	0
ADVANCED MICRO DEVICES INC	COMMON STOCK	007903107	4,292	46,798	SH		DFND	1	37,325	0	9,473
ADVANCED MICRO DEVICES INC	COMMON STOCK	007903107	10,626	115,863	SH		OTR	1	2,632	2,814	110,417
ADVANCED MICRO DEVICES INC	COMMON STOCK	007903107	19	209	SH		DFND	5	147	0	62
AEGON N V A D R	COMMON STOCK	007924103	1,479	374,356	SH		DFND	1	344,525	0	29,831
AEGON N V A D R	COMMON STOCK	007924103	107	26,918	SH		OTR	1	19,655	3,898	3,365
ADVANCED ENERGY INDUSTRIES	COMMON STOCK	007973100	52	534	SH		DFND	1	490	0	44
ADVANCED ENERGY INDUSTRIES	COMMON STOCK	007973100	191	1,973	SH		OTR	1	0	823	1,150
AEROVIRONMENT INC	COMMON STOCK	008073108	50	578	SH		DFND	1	578	0	0
AEROVIRONMENT INC	COMMON STOCK	008073108	12	141	SH		OTR	1	141	0	0
AFFILIATED MANAGERS GROUP INC	COMMON STOCK	008252108	483	4,755	SH		DFND	1	4,438	0	317
AFFILIATED MANAGERS GROUP INC	COMMON STOCK	008252108	21	209	SH		OTR	1	168	19	22
AGNICO EAGLE MINES LIMITED	COMMON STOCK	008474108	119	1,682	SH		DFND	1	1,280	0	402
AGNICO EAGLE MINES LIMITED	COMMON STOCK	008474108	83	1,173	SH		OTR	1	315	831	27
AGREE REALTY CORP	COMMON STOCK	008492100	784	11,774	SH		DFND	1	11,727	0	47
AGREE REALTY CORP	COMMON STOCK	008492100	337	5,068	SH		OTR	1	4,332	736	0
AGREE REALTY CORP	COMMON STOCK	008492100	9	130	SH		DFND	5	130	0	0
AIRBNB INC CL A	COMMON STOCK	009066101	109	740	SH		DFND	1	740	0	0
AIRBNB INC CL A	COMMON STOCK	009066101	21	140	SH		OTR	1	140	0	0
AIR PRODUCTS CHEMICALS INC	COMMON STOCK	009158106	15,347	56,170	SH		DFND	1	47,707	0	8,463
AIR PRODUCTS CHEMICALS INC	COMMON STOCK	009158106	10,872	39,791	SH		OTR	1	36,612	2,864	315
AIR PRODUCTS & CHEMICALS INC	COMMON STOCK	009158106	7	27	SH		DFND	5	27	0	0
ALAMO GROUP INC	COMMON STOCK	011311107	1,684	12,209	SH		DFND	1	12,209	0	0
ALAMOS GOLD INC NEW CLASS A	COMMON STOCK	011532108	12	1,385	SH		DFND	1	1,385	0	0
ALARM COM HOLDINGS INC	COMMON STOCK	011642105	2,115	20,440	SH		DFND	1	17,392	0	3,048
ALARM COM HOLDINGS INC	COMMON STOCK	011642105	222	2,139	SH		OTR	1	1,708	431	0
ALARM.COM INC	COMMON STOCK	011642105	370	3,575	SH		DFND	5	3,575	0	0
ALASKA AIR GROUP INC	COMMON STOCK	011659109	791	15,229	SH		DFND	1	12,754	0	2,475
ALASKA AIR GROUP INC	COMMON STOCK	011659109	113	2,178	SH		OTR	1	1,235	747	196
ALBANY INTL CORP CL A	COMMON STOCK	012348108	1,108	15,095	SH		DFND	1	15,095	0	0
ALBANY INTL CORP CL A	COMMON STOCK	012348108	109	1,483	SH		OTR	1	298	1,185	0
ALBEMARLE CORP	COMMON STOCK	012653101	104,798	710,395	SH		DFND	1	705,083	0	5,312
ALBEMARLE CORP	COMMON STOCK	012653101	12,333	83,603	SH		OTR	1	79,927	2,239	1,437
ALBEMARLE CORP	COMMON STOCK	012653101	38	257	SH		DFND	5	257	0	0
ALBERTSONS COS INC COMMON STOC	COMMON STOCK	013091103	35	2,000	SH		DFND	1	2,000	0	0
ALBERTSONS COS INC COMMON STOC	COMMON STOCK	013091103	412	23,426	SH		OTR	1	500	0	22,926
ALCOA CORPORATION	COMMON STOCK	013872106	138	5,971	SH		DFND	1	4,613	0	1,358
ALCOA CORPORATION	COMMON STOCK	013872106	23	1,014	SH		OTR	1	615	399	0
ALECTOR INC	COMMON STOCK	014442107	18	1,168	SH		DFND	1	992	0	176
ALEXANDER BALDWIN INC NEW	COMMON STOCK	014491104	4	239	SH		DFND	1	239	0	0
ALEXANDER BALDWIN INC NEW	COMMON STOCK	014491104	1	59	SH		OTR	1	0	59	0
ALEXANDERS INC	COMMON STOCK	014752109	1	5	SH		DFND	1	0	0	5
ALEXANDRIA REAL ESTATE EQUITIE	COMMON STOCK	015271109	1,792	10,057	SH		DFND	1	9,221	0	836
ALEXANDRIA REAL ESTATE EQUITIE	COMMON STOCK	015271109	416	2,333	SH		OTR	1	1,291	967	75
ALEXANDRIA REAL ESTATE EQUITY	COMMON STOCK	015271109	18	102	SH		DFND	5	102	0	0
ALEXION PHARMACEUTICALS INC	COMMON STOCK	015351109	3,936	25,187	SH		DFND	1	23,828	0	1,359
ALEXION PHARMACEUTICALS INC	COMMON STOCK	015351109	434	2,776	SH		OTR	1	2,403	215	158
ALEXION PHARMACEUTICALS INC	COMMON STOCK	015351109	406	2,597	SH		DFND	5	2,597	0	0
ALGONQUIN PWR & UTILITIES CORP	COMMON STOCK	015857105	14	850	SH		DFND	5	850	0	0
ALIGN TECHNOLOGY INC	COMMON STOCK	016255101	46,461	86,945	SH		DFND	1	86,581	0	364
ALIGN TECHNOLOGY INC	COMMON STOCK	016255101	17,146	32,086	SH		OTR	1	30,200	1,075	811
ALIGN TECHNOLOGY INC	COMMON STOCK	016255101	285	534	SH		DFND	5	531	0	3
ALLEGHANY CORP	COMMON STOCK	017175100	35	58	SH		DFND	1	48	0	10
ALLEGHANY CORP	COMMON STOCK	017175100	54	89	SH		OTR	1	31	58	0
ALLETE INC	COMMON STOCK	018522300	1,523	24,580	SH		DFND	1	24,030	0	550
ALLETE INC	COMMON STOCK	018522300	450	7,273	SH		OTR	1	7,073	0	200
ALLIANCE DATA SYSTEMS CORP	COMMON STOCK	018581108	950	12,825	SH		DFND	1	12,085	0	740
ALLIANCE DATA SYSTEMS CORP	COMMON STOCK	018581108	20	263	SH		OTR	1	53	187	23
ALLIANT ENERGY CORP	COMMON STOCK	018802108	5,163	100,188	SH		DFND	1	98,439	0	1,749
ALLIANT ENERGY CORP	COMMON STOCK	018802108	2,648	51,388	SH		OTR	1	46,726	4,662	0
ALLIANZGI CONV INVOME II	OTHER	018825109	8	1,556	SH		DFND	1	1,556	0	0
ALLIANZGI CONVERTIBLE INCO	OTHER	018828103	15	2,590	SH		DFND	1	2,590	0	0
ALLIED MOTION TECHNOLOGIES INC	COMMON STOCK	019330109	16	312	SH		OTR	1	0	312	0
ALLOGENE THERAPEUTICS INC	COMMON STOCK	019770106	6	247	SH		DFND	1	0	0	247
ALLOGENE THERAPEUTICS INC	COMMON STOCK	019770106	1	50	SH		OTR	1	50	0	0
ALLSTATE CORP	COMMON STOCK	020002101	14,107	128,330	SH		DFND	1	123,529	0	4,801
ALLSTATE CORP	COMMON STOCK	020002101	6,098	55,477	SH		OTR	1	52,286	3,131	60
ALLSTATE CORP	COMMON STOCK	020002101	114	1,038	SH		DFND	5	950	0	88
ALTABANCORP	COMMON STOCK	021347109	66	2,361	SH		DFND	1	2,307	0	54
ALTAIR ENGINEERING INC A C	COMMON STOCK	021369103	39	678	SH		DFND	1	678	0	0
ALTAIR ENGINEERING INC A C	COMMON STOCK	021369103	249	4,279	SH		OTR	1	190	634	3,455
AMALGAMATED BK OF NEW YORK	COMMON STOCK	022663108	112	8,153	SH		DFND	1	0	0	8,153
AMARIN CORPORATION PLC A D R	COMMON STOCK	023111206	97	19,895	SH		DFND	1	19,895	0	0
AMARIN CORPORATION PLC A D R	COMMON STOCK	023111206	120	24,613	SH		OTR	1	24,613	0	0
AMAZON COM INC	COMMON STOCK	023135106	751,573	230,761	SH		DFND	1	224,954	0	5,807
AMAZON COM INC	COMMON STOCK	023135106	162,651	49,940	SH		OTR	1	41,904	2,065	5,971
AMAZON.COM INC	COMMON STOCK	023135106	3,338	1,025	SH		DFND	5	1,001	0	24
AMBAC FINANCIAL GROUP INC	COMMON STOCK	023139884	3	198	SH		DFND	1	198	0	0
AMEDISYS INC	COMMON STOCK	023436108	3,230	11,014	SH		DFND	1	10,879	0	135
AMEDISYS INC	COMMON STOCK	023436108	1,515	5,164	SH		OTR	1	5,157	0	7
AMEDISYS INC	COMMON STOCK	023436108	310	1,057	SH		DFND	5	1,057	0	0
AMERCO	COMMON STOCK	023586100	93	204	SH		DFND	1	94	0	110
AMERCO	COMMON STOCK	023586100	46	101	SH		DFND	5	101	0	0
AMEREN CORP	COMMON STOCK	023608102	4,477	57,354	SH		DFND	1	44,849	0	12,505
AMEREN CORP	COMMON STOCK	023608102	2,072	26,539	SH		OTR	1	20,019	2,997	3,523
AMEREN CORP	COMMON STOCK	023608102	39	504	SH		DFND	5	504	0	0
AMERICAN ASSETS TRUST INC	COMMON STOCK	024013104	26	906	SH		DFND	1	228	0	678
AMERICAN AXLE & MFG HLDGS INC	COMMON STOCK	024061103	6	672	SH		DFND	1	672	0	0
AMERICAN CAMPUS COMMUNITIES	COMMON STOCK	024835100	7,859	183,745	SH		DFND	1	178,845	0	4,900
AMERICAN CAMPUS COMMUNITIES	COMMON STOCK	024835100	1,534	35,867	SH		OTR	1	35,767	0	100
AMERICAN CAMPUS COMMUNITIES	COMMON STOCK	024835100	64	1,485	SH		DFND	5	1,485	0	0
AMERICAN ELECTRIC POWER	COMMON STOCK	025537101	13,392	160,834	SH		DFND	1	156,231	0	4,603
AMERICAN ELECTRIC POWER	COMMON STOCK	025537101	8,082	97,064	SH		OTR	1	92,484	1,624	2,956
AMERICAN ELECTRIC POWER CO INC	COMMON STOCK	025537101	213	2,552	SH		DFND	5	2,552	0	0
AMERICAN EQUITY INVT LIFE HL	COMMON STOCK	025676206	682	24,649	SH		DFND	1	22,675	0	1,974
AMERICAN EQUITY INVT LIFE HL	COMMON STOCK	025676206	92	3,344	SH		OTR	1	3,193	112	39
AMERICAN EXPRESS CO	COMMON STOCK	025816109	29,905	247,332	SH		DFND	1	242,322	0	5,010
AMERICAN EXPRESS CO	COMMON STOCK	025816109	18,432	152,439	SH		OTR	1	140,927	5,713	5,799
AMERICAN EXPRESS COMPANY	COMMON STOCK	025816109	43	352	SH		DFND	5	352	0	0
AMERICAN FINL GROUP INC OHIO	COMMON STOCK	025932104	586	6,689	SH		DFND	1	4,475	0	2,214
AMERICAN FINL GROUP INC OHIO	COMMON STOCK	025932104	49	560	SH		OTR	1	500	11	49
AMERICAN FINANCIAL GROUP INC	COMMON STOCK	025932104	39	446	SH		DFND	5	446	0	0
AMERICAN INTL GROUP INC WT 1	OTHER	026874156	0	153	SH		DFND	1	153	0	0
AMERICAN INTL GROUP INC WT 1	OTHER	026874156	0	5	SH		OTR	1	5	0	0
AMERICAN INTERNATIONAL GROUP	COMMON STOCK	026874784	1,143	30,187	SH		DFND	1	24,152	0	6,035
AMERICAN INTERNATIONAL GROUP	COMMON STOCK	026874784	156	4,104	SH		OTR	1	2,756	1,297	51
AMERICAN INTERNATIONAL GROUP	COMMON STOCK	026874784	50	1,312	SH		DFND	5	1,119	0	193
AMERICAN NATL BNKSHS/DANVILLE	COMMON STOCK	027745108	35	1,350	SH		OTR	1	0	1,350	0
AMERICAN RENAL ASSOCS HLDGS IN	COMMON STOCK	029227105	2	202	SH		DFND	1	0	0	202
AMERICAN SOFTWARE INC CL A	COMMON STOCK	029683109	21	1,229	SH		DFND	1	1,229	0	0
AMERICAN STATES WATER CO	COMMON STOCK	029899101	264	3,326	SH		DFND	1	3,093	0	233
AMERICAN STATES WATER CO	COMMON STOCK	029899101	119	1,500	SH		OTR	1	500	0	1,000
AMERICAN STATES WATER CO	COMMON STOCK	029899101	9	112	SH		DFND	5	112	0	0
AMERICAN VANGUARD CORP	COMMON STOCK	030371108	13	814	SH		DFND	1	814	0	0
AMERICAN WATER WORKS CO INC	COMMON STOCK	030420103	4,108	26,765	SH		DFND	1	22,036	0	4,729
AMERICAN WATER WORKS CO INC	COMMON STOCK	030420103	2,944	19,184	SH		OTR	1	10,600	1,687	6,897
AMERICAN WATER WORKS CO INC	COMMON STOCK	030420103	2	15	SH		DFND	5	15	0	0
AMERICAN WOODMARK CORP	COMMON STOCK	030506109	961	10,244	SH		DFND	1	10,174	0	70
AMERICAN WOODMARK CORP	COMMON STOCK	030506109	70	746	SH		OTR	1	746	0	0
AMERICAN WOODMARK CORP	COMMON STOCK	030506109	9	98	SH		DFND	5	98	0	0
AMETEK INC	COMMON STOCK	031100100	5,758	47,603	SH		DFND	1	46,071	0	1,532
AMETEK INC	COMMON STOCK	031100100	1,375	11,372	SH		OTR	1	10,989	258	125
AMETEK INC	COMMON STOCK	031100100	8	67	SH		DFND	5	48	0	19
AMGEN INC	COMMON STOCK	031162100	120,390	523,617	SH		DFND	1	497,515	0	26,102
AMGEN INC	COMMON STOCK	031162100	43,032	187,160	SH		OTR	1	169,232	2,998	14,930
AMGEN INC	COMMON STOCK	031162100	695	3,014	SH		DFND	5	3,014	0	0
AMKOR TECHNOLOGY INC	COMMON STOCK	031652100	23	1,500	SH		DFND	1	1,500	0	0
AMPCO-PITTSBURG CORP	COMMON STOCK	032037103	1	106	SH		DFND	1	106	0	0
AMPHENOL CORP CL A	COMMON STOCK	032095101	21,541	164,724	SH		DFND	1	163,253	0	1,471
AMPHENOL CORP CL A	COMMON STOCK	032095101	6,382	48,796	SH		OTR	1	36,712	4,844	7,240
AMPHENOL CORP	COMMON STOCK	032095101	644	4,923	SH		DFND	5	4,923	0	0
AMPLIFY TRANSFORMATIONAL DATA	OTHER	032108607	96	2,757	SH		OTR	1	2,757	0	0
YIELDSHARES HIGH INCOME ETF	OTHER	032108847	6	355	SH		DFND	1	355	0	0
ANALOG DEVICES INC	COMMON STOCK	032654105	8,732	59,106	SH		DFND	1	51,866	0	7,240
ANALOG DEVICES INC	COMMON STOCK	032654105	3,622	24,520	SH		OTR	1	14,599	2,382	7,539
ANALOG DEVICES INC	COMMON STOCK	032654105	92	622	SH		DFND	5	605	0	17
ANAPTYSBIO INC	COMMON STOCK	032724106	18	814	SH		DFND	1	506	0	308
ANAPTYSBIO INC	COMMON STOCK	032724106	4	172	SH		OTR	1	0	172	0
ANGLOGOLD ASHANTI LTD SPON A D	COMMON STOCK	035128206	108	4,780	SH		DFND	1	3,376	0	1,404
ANIKA THERAPEUTICS INC	COMMON STOCK	035255108	9	200	SH		DFND	1	200	0	0
ANNALY CAPITAL MANAGEMENT INC	COMMON STOCK	035710409	1,112	131,506	SH		DFND	1	116,536	0	14,970
ANNALY CAPITAL MANAGEMENT INC	COMMON STOCK	035710409	195	23,087	SH		OTR	1	23,087	0	0
ANNALY CAPITAL MANAGEMENT	COMMON STOCK	035710409	7	851	SH		DFND	5	0	0	851
ANTARES PHARMA INC	COMMON STOCK	036642106	8	1,990	SH		DFND	1	1,990	0	0
ANTHEM INC	COMMON STOCK	036752103	9,579	29,833	SH		DFND	1	27,477	0	2,356
ANTHEM INC	COMMON STOCK	036752103	4,194	13,060	SH		OTR	1	7,668	104	5,288
ANTHEM INC	COMMON STOCK	036752103	255	795	SH		DFND	5	788	0	7
ANWORTH MORTGAGE ASSET CORP	COMMON STOCK	037347101	10	3,800	SH		OTR	1	0	0	3,800
APACHE CORP	COMMON STOCK	037411105	509	35,888	SH		DFND	1	32,342	0	3,546
APACHE CORP	COMMON STOCK	037411105	133	9,358	SH		OTR	1	8,850	0	508
APOGEE ENTERPRISES INC	COMMON STOCK	037598109	247	7,799	SH		DFND	1	7,709	0	90
APPLE INC	COMMON STOCK	037833100	1,248,069	9,405,906	SH		DFND	1	9,087,636	0	318,270
APPLE INC	COMMON STOCK	037833100	443,694	3,343,844	SH		OTR	1	2,928,299	107,102	308,443
APPLE INC	COMMON STOCK	037833100	2,540	19,133	SH		DFND	5	17,713	0	1,420
APPLE INC	COMMON STOCK	037833100	10	52	SH	Put	DFND	1	52	0	0
APPLE INC	COMMON STOCK	037833100	20	31	SH	Put	OTR	1	11	20	0
APPLIED MATERIALS INC	COMMON STOCK	038222105	25,660	297,334	SH		DFND	1	284,346	0	12,988
APPLIED MATERIALS INC	COMMON STOCK	038222105	7,836	90,800	SH		OTR	1	87,303	805	2,692
APPLIED MATERIALS INC	COMMON STOCK	038222105	111	1,281	SH		DFND	5	1,249	0	32
APTAR GROUP INC	COMMON STOCK	038336103	2,154	15,730	SH		DFND	1	15,301	0	429
APTAR GROUP INC	COMMON STOCK	038336103	378	2,757	SH		OTR	1	1,216	41	1,500
APTAR GROUP INC	COMMON STOCK	038336103	49	354	SH		DFND	5	354	0	0
ARBOR REALTY TRUST INC	COMMON STOCK	038923108	27	1,905	SH		DFND	1	1,767	0	138
ARBOR REALTY TRUST INC	COMMON STOCK	038923108	1	78	SH		OTR	1	0	0	78
ARCHER DANIELS MIDLAND CO	COMMON STOCK	039483102	7,918	157,072	SH		DFND	1	150,434	0	6,638
ARCHER DANIELS MIDLAND CO	COMMON STOCK	039483102	1,389	27,548	SH		OTR	1	26,768	775	5
ARCHER DANIELS MIDLAND CO	COMMON STOCK	039483102	40	790	SH		DFND	5	659	0	131
ARCIMOTO INC COM	COMMON STOCK	039587100	3	200	SH		OTR	1	200	0	0
ARCOSA INC	COMMON STOCK	039653100	336	6,115	SH		DFND	1	5,851	0	264
ARCOSA INC	COMMON STOCK	039653100	73	1,339	SH		OTR	1	116	1,223	0
ARENA PHARMACEUTICALS INC	COMMON STOCK	040047607	19	253	SH		DFND	1	226	0	27
ARISTA NETWORKS INC	COMMON STOCK	040413106	1,849	6,365	SH		DFND	1	5,062	0	1,303
ARISTA NETWORKS INC	COMMON STOCK	040413106	539	1,856	SH		OTR	1	1,265	0	591
ARISTA NETWORKS INC	COMMON STOCK	040413106	9	30	SH		DFND	5	30	0	0
ARMOUR RESIDENTIAL REIT INC	COMMON STOCK	042315507	4	364	SH		DFND	1	289	0	75
ARROW ELECTRS INC	COMMON STOCK	042735100	1,195	12,287	SH		DFND	1	11,043	0	1,244
ARROW ELECTRS INC	COMMON STOCK	042735100	611	6,282	SH		OTR	1	5,117	1,140	25
ARROW ELECTRONICS INC	COMMON STOCK	042735100	20	201	SH		DFND	5	201	0	0
ARTESIAN RESOURCES CORP CL A C	COMMON STOCK	043113208	10	274	SH		OTR	1	274	0	0
ASBURY AUTOMOTIVE GROUP	COMMON STOCK	043436104	486	3,330	SH		DFND	1	3,330	0	0
ASBURY AUTOMOTIVE GROUP	COMMON STOCK	043436104	10	70	SH		OTR	1	70	0	0
ASHLAND GLOBAL HOLDINGS INC	COMMON STOCK	044186104	181	2,289	SH		DFND	1	1,897	0	392
ASHLAND GLOBAL HOLDINGS INC	COMMON STOCK	044186104	116	1,466	SH		OTR	1	1,309	58	99
ASHLAND GLOBAL HOLDINGS INC	COMMON STOCK	044186104	1	13	SH		DFND	5	13	0	0
ASPEN TECHNOLOGY INC	COMMON STOCK	045327103	4,273	32,810	SH		DFND	1	32,399	0	411
ASPEN TECHNOLOGY INC	COMMON STOCK	045327103	251	1,929	SH		OTR	1	1,884	34	11
ASPEN TECHNOLOGY INC	COMMON STOCK	045327103	168	1,288	SH		DFND	5	1,288	0	0
ASSEMBLY BIOSCIENCES INC	COMMON STOCK	045396108	1	157	SH		OTR	1	0	157	0
ASSOCIATED BANC CORP	COMMON STOCK	045487105	1,226	71,908	SH		DFND	1	71,125	0	783
ASSOCIATED BANC CORP	COMMON STOCK	045487105	289	16,963	SH		OTR	1	15,822	0	1,141
ASSOCIATED CAPITAL GROUP W I	COMMON STOCK	045528106	246	7,000	SH		DFND	1	7,000	0	0
ASTEC INDUSTRIES INC	COMMON STOCK	046224101	76	1,312	SH		DFND	1	1,199	0	113
ASTEC INDUSTRIES INC	COMMON STOCK	046224101	54	939	SH		OTR	1	0	939	0
ASTRAZENECA P L C SPSD A D R	COMMON STOCK	046353108	2,946	58,936	SH		DFND	1	43,604	0	15,332
ASTRAZENECA P L C SPSD A D R	COMMON STOCK	046353108	890	17,797	SH		OTR	1	10,418	3,829	3,550
ASTRAZENECA PLC	COMMON STOCK	046353108	4	71	SH		DFND	5	71	0	0
ATARA BIOTHERAPEUTICS INC	COMMON STOCK	046513107	2	91	SH		DFND	1	0	0	91
ATARA BIOTHERAPEUTICS INC	COMMON STOCK	046513107	2	118	SH		OTR	1	0	118	0
ATKORE INTERNATIONAL GROUP I	COMMON STOCK	047649108	19	458	SH		DFND	1	53	0	405
ATKORE INTERNATIONAL GROUP I	COMMON STOCK	047649108	62	1,513	SH		OTR	1	0	1,513	0
ATLAS AIR WORLDWIDE HOLDINGS	COMMON STOCK	049164205	726	13,312	SH		DFND	1	12,736	0	576
ATLAS AIR WORLDWIDE HOLDINGS	COMMON STOCK	049164205	10	192	SH		OTR	1	117	75	0
ATMOS ENERGY CORP	COMMON STOCK	049560105	668	7,001	SH		DFND	1	6,439	0	562
ATMOS ENERGY CORP	COMMON STOCK	049560105	71	741	SH		OTR	1	693	7	41
ATMOS ENERGY CORP	COMMON STOCK	049560105	3	32	SH		DFND	5	32	0	0
ATRION CORPORATION	COMMON STOCK	049904105	77	120	SH		DFND	1	104	0	16
ATRION CORPORATION	COMMON STOCK	049904105	48	74	SH		OTR	1	4	70	0
AUBURN NATIONAL BANCORPORATION	COMMON STOCK	050473107	2	39	SH		DFND	1	39	0	0
AUTODESK INC	COMMON STOCK	052769106	14,801	48,472	SH		DFND	1	44,017	0	4,455
AUTODESK INC	COMMON STOCK	052769106	1,407	4,610	SH		OTR	1	4,016	396	198
AUTODESK INC	COMMON STOCK	052769106	295	965	SH		DFND	5	901	0	64
AUTOLIV INC	COMMON STOCK	052800109	324	3,520	SH		DFND	1	3,479	0	41
AUTOLIV INC	COMMON STOCK	052800109	121	1,311	SH		OTR	1	582	729	0
AUTOMATIC DATA PROCESSING	COMMON STOCK	053015103	74,561	423,161	SH		DFND	1	403,298	0	19,863
AUTOMATIC DATA PROCESSING	COMMON STOCK	053015103	40,616	230,508	SH		OTR	1	211,287	10,400	8,821
AUTOMATIC DATA PROCESSING INC	COMMON STOCK	053015103	256	1,454	SH		DFND	5	1,454	0	0
AUTOMATIC DATA PROCESSING	COMMON STOCK	053015103	13	36	SH	Put	DFND	1	36	0	0
AUTOMATIC DATA PROCESSING	COMMON STOCK	053015103	1	23	SH	Put	OTR	1	8	15	0
AUTOZONE INC	COMMON STOCK	053332102	2,920	2,463	SH		DFND	1	2,179	0	284
AUTOZONE INC	COMMON STOCK	053332102	1,023	863	SH		OTR	1	677	54	132
AUTOZONE INC	COMMON STOCK	053332102	83	70	SH		DFND	5	70	0	0
AVALONBAY COMMUNITIES INC	COMMON STOCK	053484101	1,885	11,749	SH		DFND	1	11,086	0	663
AVALONBAY COMMUNITIES INC	COMMON STOCK	053484101	964	6,010	SH		OTR	1	2,778	13	3,219
AVALONBAY COMMUNITIES INC	COMMON STOCK	053484101	15	93	SH		DFND	5	93	0	0
AVERY DENNISON CORP	COMMON STOCK	053611109	2,730	17,602	SH		DFND	1	15,737	0	1,865
AVERY DENNISON CORP	COMMON STOCK	053611109	796	5,130	SH		OTR	1	4,129	983	18
AVERY DENNISON CORP	COMMON STOCK	053611109	61	389	SH		DFND	5	379	0	10
AVIS BUDGET GROUP INC	COMMON STOCK	053774105	15	414	SH		DFND	1	357	0	57
AVIS BUDGET GROUP INC	COMMON STOCK	053774105	9	227	SH		OTR	1	0	178	49
AVNET INC	COMMON STOCK	053807103	29	832	SH		DFND	1	105	0	727
AVNET INC	COMMON STOCK	053807103	2	57	SH		DFND	5	57	0	0
BP PLC SPON A D R	COMMON STOCK	055622104	9,829	478,997	SH		DFND	1	462,182	0	16,815
BP PLC SPON A D R	COMMON STOCK	055622104	7,271	354,313	SH		OTR	1	343,449	4,079	6,785
BP PLC	COMMON STOCK	055622104	4	184	SH		DFND	5	184	0	0
BP PRUDHOE BAY ROYALTY TRUST	COMMON STOCK	055630107	6	2,400	SH		DFND	1	2,400	0	0
BADGER METER INC	COMMON STOCK	056525108	1,106	11,758	SH		DFND	1	11,758	0	0
BADGER METER INC	COMMON STOCK	056525108	2,321	24,679	SH		OTR	1	23,251	0	1,428
BADGER METER INC	COMMON STOCK	056525108	15	162	SH		DFND	5	162	0	0
BAIDU INC SPON A D R	COMMON STOCK	056752108	7,894	36,507	SH		DFND	1	35,162	0	1,345
BAIDU INC SPON A D R	COMMON STOCK	056752108	1,523	7,042	SH		OTR	1	7,038	0	4
BAIDU INC	COMMON STOCK	056752108	2	11	SH		DFND	5	11	0	0
BALCHEM CORP	COMMON STOCK	057665200	1,886	16,365	SH		DFND	1	14,570	0	1,795
BALCHEM CORP	COMMON STOCK	057665200	322	2,797	SH		OTR	1	1,377	0	1,420
BALCHEM CORP	COMMON STOCK	057665200	12	107	SH		DFND	5	107	0	0
BALL CORP	COMMON STOCK	058498106	6,291	67,520	SH		DFND	1	65,354	0	2,166
BALL CORP	COMMON STOCK	058498106	213	2,282	SH		OTR	1	1,839	351	92
BALL CORP	COMMON STOCK	058498106	3	27	SH		DFND	5	27	0	0
BALLARD POWER SYSTEMS INC	COMMON STOCK	058586108	16	700	SH		DFND	1	700	0	0
BALLARD POWER SYSTEMS INC	COMMON STOCK	058586108	274	11,700	SH		OTR	1	11,700	0	0
BANCO BRADESCO A D R	COMMON STOCK	059460303	274	51,984	SH		DFND	1	29,487	0	22,497
BANCO BRADESCO A D R	COMMON STOCK	059460303	10	1,829	SH		OTR	1	1,621	0	208
BANCO DE CHILE A D R	COMMON STOCK	059520106	17	830	SH		DFND	1	620	0	210
BANK OF AMERICA CORP	COMMON STOCK	060505104	206,691	6,819,235	SH		DFND	1	6,590,919	0	228,316
BANK OF AMERICA CORP	COMMON STOCK	060505104	29,636	977,727	SH		OTR	1	926,262	16,654	34,811
BANK OF AMERICA CORP	COMMON STOCK	060505104	261	8,612	SH		DFND	5	8,612	0	0
BANK OF HAWAII CORP	COMMON STOCK	062540109	128	1,665	SH		DFND	1	937	0	728
BANK OF HAWAII CORP	COMMON STOCK	062540109	75	983	SH		OTR	1	954	0	29
BANK OF MARIN BANCORP CA	COMMON STOCK	063425102	11	312	SH		DFND	1	246	0	66
BANK MONTREAL QUE	COMMON STOCK	063671101	2,518	33,120	SH		DFND	1	18,902	0	14,218
BANK MONTREAL QUE	COMMON STOCK	063671101	96	1,262	SH		OTR	1	1,229	0	33
BANK OF MONTREAL	COMMON STOCK	063671101	0	1	SH		DFND	5	1	0	0
BANK OF NEW YORK MELLON CORP	COMMON STOCK	064058100	4,541	107,011	SH		DFND	1	99,295	0	7,716
BANK OF NEW YORK MELLON CORP	COMMON STOCK	064058100	1,931	45,488	SH		OTR	1	45,047	400	41
BANK OF NEW YORK MELLON CORP	COMMON STOCK	064058100	62	1,453	SH		DFND	5	1,291	0	162
BANK OF NOVA SCOTIA	COMMON STOCK	064149107	2,918	53,991	SH		DFND	1	51,435	0	2,556
BANK OF NOVA SCOTIA	COMMON STOCK	064149107	415	7,673	SH		OTR	1	7,513	0	160
BAR HBR BANKSHARES	COMMON STOCK	066849100	3	127	SH		DFND	1	127	0	0
BARNES GROUP INC	COMMON STOCK	067806109	515	10,160	SH		DFND	1	7,757	0	2,403
BARNES GROUP INC	COMMON STOCK	067806109	151	2,981	SH		OTR	1	1,246	1,735	0
BARNES GROUP INC (THE)	COMMON STOCK	067806109	6	125	SH		DFND	5	125	0	0
BARRICK GOLD CORP	COMMON STOCK	067901108	1,889	82,914	SH		DFND	1	77,246	0	5,668
BARRICK GOLD CORP	COMMON STOCK	067901108	592	26,006	SH		OTR	1	14,548	0	11,458
BAUSCH HEALTH COS INC	COMMON STOCK	071734107	309	14,876	SH		DFND	1	1,538	0	13,338
BAUSCH HEALTH COS INC	COMMON STOCK	071734107	193	9,299	SH		OTR	1	1,183	8,047	69
BAXTER INTERNATIONAL INC	COMMON STOCK	071813109	13,281	165,512	SH		DFND	1	158,733	0	6,779
BAXTER INTERNATIONAL INC	COMMON STOCK	071813109	6,318	78,733	SH		OTR	1	74,632	195	3,906
BAXTER INTERNATIONAL INC	COMMON STOCK	071813109	33	413	SH		DFND	5	413	0	0
BEACON ROOFING SUPPLY INC	COMMON STOCK	073685109	514	12,786	SH		DFND	1	12,759	0	27
BEACON ROOFING SUPPLY INC	COMMON STOCK	073685109	12	300	SH		OTR	1	272	0	28
BEASLEY BROADCAST GRP INC A	COMMON STOCK	074014101	0	50	SH		DFND	1	50	0	0
BECTON DICKINSON AND CO	COMMON STOCK	075887109	13,917	55,623	SH		DFND	1	44,445	0	11,178
BECTON DICKINSON AND CO	COMMON STOCK	075887109	5,737	22,928	SH		OTR	1	18,153	327	4,448
BECTON DICKINSON & CO	COMMON STOCK	075887109	17	68	SH		DFND	5	68	0	0
BECTON DICKINSON AND CO	COMMON STOCK	075887109	2	4	SH	Put	DFND	1	4	0	0
BED BATH & BEYOND INC	COMMON STOCK	075896100	91	5,143	SH		DFND	1	4,539	0	604
BELDEN INC	COMMON STOCK	077454106	58	1,389	SH		DFND	1	329	0	1,060
BELDEN INC	COMMON STOCK	077454106	34	808	SH		OTR	1	808	0	0
BELLEROPHON THERAPEUTICS INC	COMMON STOCK	078771300	8	1,221	SH		OTR	1	1,221	0	0
BELLRING BRANDS INC CLASS A	COMMON STOCK	079823100	59	2,440	SH		OTR	1	45	2,395	0
WR BERKLEY CORP	COMMON STOCK	084423102	115	1,726	SH		DFND	1	1,075	0	651
WR BERKLEY CORP	COMMON STOCK	084423102	156	2,351	SH		OTR	1	487	0	1,864
BERKSHIRE HATHAWAY INC CL A	COMMON STOCK	084670108	155,474	447	SH		DFND	1	433	0	14
BERKSHIRE HATHAWAY INC CL A	COMMON STOCK	084670108	253,209	728	SH		OTR	1	719	1	8
BERKSHIRE HATHAWAY INC CL B	COMMON STOCK	084670702	109,864	473,818	SH		DFND	1	437,085	0	36,733
BERKSHIRE HATHAWAY INC CL B	COMMON STOCK	084670702	46,097	198,807	SH		OTR	1	176,461	8,907	13,439
BERKSHIRE HATHAWAY INC	COMMON STOCK	084670702	324	1,396	SH		DFND	5	1,396	0	0
BERKSHIRE HATHAWAY INC CL B	COMMON STOCK	084670702	2	23	SH	Put	OTR	1	23	0	0
BERKSHIRE HILLS BANCORP INC	COMMON STOCK	084680107	93	5,454	SH		DFND	1	3,426	0	2,028
BEST BUY CO INC	COMMON STOCK	086516101	6,394	64,068	SH		DFND	1	60,474	0	3,594
BEST BUY CO INC	COMMON STOCK	086516101	2,752	27,581	SH		OTR	1	26,904	676	1
BEST BUY CO INC	COMMON STOCK	086516101	84	844	SH		DFND	5	844	0	0
B H P BILLITON LIMITED A D R	COMMON STOCK	088606108	1,952	29,877	SH		DFND	1	17,712	0	12,165
B H P BILLITON LIMITED A D R	COMMON STOCK	088606108	418	6,405	SH		OTR	1	4,716	1,427	262
BICYCLE THERAPEUTICS LTD A D R	COMMON STOCK	088786108	232	12,911	SH		DFND	1	12,911	0	0
BICYCLE THERAPEUTICS LTD A D R	COMMON STOCK	088786108	3	162	SH		OTR	1	162	0	0
BIG LOTS INC	COMMON STOCK	089302103	89	2,078	SH		DFND	1	1,814	0	264
BIG LOTS INC	COMMON STOCK	089302103	11	252	SH		OTR	1	75	166	11
BILIBILI INC A D R	COMMON STOCK	090040106	1,792	20,902	SH		DFND	1	19,709	0	1,193
BILIBILI INC A D R	COMMON STOCK	090040106	279	3,259	SH		OTR	1	3,259	0	0
BILIBILI INC	COMMON STOCK	090040106	1	13	SH		DFND	5	13	0	0
BILL COM HOLDINGS INC	COMMON STOCK	090043100	3,038	22,260	SH		DFND	1	22,228	0	32
BILL COM HOLDINGS INC	COMMON STOCK	090043100	744	5,455	SH		OTR	1	5,408	35	12
BIO RAD LABS INC CL A	COMMON STOCK	090572207	2,367	4,061	SH		DFND	1	3,790	0	271
BIO RAD LABS INC CL A	COMMON STOCK	090572207	575	988	SH		OTR	1	970	6	12
BIO-RAD LABORATORIES	COMMON STOCK	090572207	114	195	SH		DFND	5	195	0	0
BIOTELEMETRY INC	COMMON STOCK	090672106	24	332	SH		DFND	1	115	0	217
BLACK HILLS CORP	COMMON STOCK	092113109	1,365	22,205	SH		DFND	1	20,833	0	1,372
BLACK HILLS CORP	COMMON STOCK	092113109	151	2,455	SH		OTR	1	2,455	0	0
BLACKROCK MUNICIPAL INCOME QUA	OTHER	092479104	121	8,002	SH		DFND	1	8,002	0	0
BLACKROCK ENHD GLB DIV TRT	OTHER	092501105	32	2,931	SH		DFND	1	2,931	0	0
BLACKROCK CREDIT ALLOCATION IN	OTHER	092508100	22	1,500	SH		DFND	1	1,500	0	0
BLACKROCK CAPITAL INVESTMENT	COMMON STOCK	092533108	21	7,841	SH		OTR	1	0	0	7,841
BLOCK H R INC	COMMON STOCK	093671105	324	20,421	SH		DFND	1	18,772	0	1,649
BLOCK H R INC	COMMON STOCK	093671105	34	2,150	SH		OTR	1	2,150	0	0
BLOOM ENERGY CORP A	COMMON STOCK	093712107	25	874	SH		DFND	1	132	0	742
BLOOM ENERGY CORP A	COMMON STOCK	093712107	43	1,500	SH		OTR	1	1,500	0	0
BLOOMIN BRANDS INC	COMMON STOCK	094235108	27	1,406	SH		DFND	1	1,334	0	72
BLUCORA INC	COMMON STOCK	095229100	29	1,833	SH		DFND	1	1,833	0	0
BLUEGREEN VACATIONS HLDG COR	COMMON STOCK	096308101	1	80	SH		OTR	1	80	0	0
BOEING CO THE	COMMON STOCK	097023105	72,140	337,005	SH		DFND	1	330,472	0	6,533
BOEING CO THE	COMMON STOCK	097023105	26,782	125,114	SH		OTR	1	115,508	1,414	8,192
BOEING COMPANY	COMMON STOCK	097023105	54	251	SH		DFND	5	218	0	33
BONANZA CREEK ENERGY INC	COMMON STOCK	097793400	18	910	SH		DFND	1	0	0	910
BOOT BARN HOLDINGS INC	COMMON STOCK	099406100	238	5,494	SH		DFND	1	5,494	0	0
BOOT BARN HOLDINGS INC	COMMON STOCK	099406100	70	1,618	SH		OTR	1	1,618	0	0
BOOT BARN HOLDINGS INC	COMMON STOCK	099406100	14	314	SH		DFND	5	314	0	0
BOOZ ALLEN HAMILTON HOLDING	COMMON STOCK	099502106	2,240	25,687	SH		DFND	1	23,489	0	2,198
BOOZ ALLEN HAMILTON HOLDING	COMMON STOCK	099502106	722	8,288	SH		OTR	1	6,442	311	1,535
BOOZ ALLEN HAMILTON HOLDING	COMMON STOCK	099502106	126	1,446	SH		DFND	5	1,424	0	22
BORG WARNER INC	COMMON STOCK	099724106	585	15,127	SH		DFND	1	13,133	0	1,994
BORG WARNER INC	COMMON STOCK	099724106	747	19,347	SH		OTR	1	16,290	1,367	1,690
BORG WARNER INC	COMMON STOCK	099724106	22	568	SH		DFND	5	568	0	0
BOSTON BEER INC A	COMMON STOCK	100557107	441	444	SH		DFND	1	320	0	124
BOSTON BEER INC A	COMMON STOCK	100557107	22	22	SH		OTR	1	0	15	7
BOSTON OMAHA CORP	COMMON STOCK	101044105	2	61	SH		DFND	1	61	0	0
BOSTON PRIVATE FINL HLDNGS INC	COMMON STOCK	101119105	16	1,846	SH		DFND	1	1,150	0	696
BOSTON PPTYS INC	COMMON STOCK	101121101	1,384	14,636	SH		DFND	1	13,323	0	1,313
BOSTON PPTYS INC	COMMON STOCK	101121101	841	8,893	SH		OTR	1	8,872	10	11
BOSTON PROPERTIES INC	COMMON STOCK	101121101	12	130	SH		DFND	5	130	0	0
BOSTON SCIENTIFIC CORP	COMMON STOCK	101137107	6,128	170,456	SH		DFND	1	158,339	0	12,117
BOSTON SCIENTIFIC CORP	COMMON STOCK	101137107	3,074	85,514	SH		OTR	1	79,995	580	4,939
BOSTON SCIENTIFIC CORP	COMMON STOCK	101137107	3	82	SH		DFND	5	18	0	64
BOTTOMLINE TECHNOLOGIES DEL IN	COMMON STOCK	101388106	153	2,900	SH		DFND	1	198	0	2,702
BOTTOMLINE TECHNOLOGIES DEL IN	COMMON STOCK	101388106	169	3,210	SH		OTR	1	160	0	3,050
BOULDER GROWTH AND INCOME FUND	OTHER	101507101	1	100	SH		DFND	1	100	0	0
BOYD GAMING CORPORATION	COMMON STOCK	103304101	595	13,849	SH		DFND	1	12,617	0	1,232
BOYD GAMING CORPORATION	COMMON STOCK	103304101	39	929	SH		OTR	1	216	0	713
BOYD GAMING CORP	COMMON STOCK	103304101	1	23	SH		DFND	5	23	0	0
BRADY CORPORATION CL A	COMMON STOCK	104674106	30	577	SH		DFND	1	577	0	0
BRADY CORPORATION CL A	COMMON STOCK	104674106	1	15	SH		OTR	1	0	15	0
BRADY CORP	COMMON STOCK	104674106	3	57	SH		DFND	5	57	0	0
BRANDYWINE REALTY TRUST	COMMON STOCK	105368203	56	4,686	SH		DFND	1	4,251	0	435
BRANDYWINE REALTY TRUST	COMMON STOCK	105368203	4	330	SH		OTR	1	330	0	0
BRIDGE BANCORP INC	COMMON STOCK	108035106	2	74	SH		DFND	1	0	0	74
BRIGHT HORIZONS FAMILY SOLUT	COMMON STOCK	109194100	1,261	7,287	SH		DFND	1	5,624	0	1,663
BRIGHT HORIZONS FAMILY SOLUT	COMMON STOCK	109194100	18	101	SH		OTR	1	90	0	11
BRIGHT HORIZONS FAMILY SOLTNS	COMMON STOCK	109194100	40	233	SH		DFND	5	233	0	0
BRINKER INTL INC	COMMON STOCK	109641100	106	1,871	SH		DFND	1	1,119	0	752
BRINKER INTL INC	COMMON STOCK	109641100	21	377	SH		OTR	1	0	199	178
THE BRINKS CO	COMMON STOCK	109696104	28	395	SH		DFND	1	373	0	22
THE BRINKS CO	COMMON STOCK	109696104	65	898	SH		OTR	1	24	855	19
BRISTOL MYERS SQUIBB CO	COMMON STOCK	110122108	108,559	1,750,107	SH		DFND	1	1,670,573	0	79,534
BRISTOL MYERS SQUIBB CO	COMMON STOCK	110122108	51,955	837,580	SH		OTR	1	742,241	82,588	12,751
BRISTOL MYERS SQUIBB CO	COMMON STOCK	110122108	263	4,233	SH		DFND	5	4,199	0	34
BRISTOL MYERS SQUIBB RT 3	OTHER	110122157	220	319,027	SH		DFND	1	313,385	0	5,642
BRISTOL MYERS SQUIBB RT 3	OTHER	110122157	80	117,394	SH		OTR	1	112,095	4,291	1,008
BRITISH AMERN TOB PLC SPON A D	COMMON STOCK	110448107	1,077	28,740	SH		DFND	1	18,345	0	10,395
BRITISH AMERN TOB PLC SPON A D	COMMON STOCK	110448107	174	4,646	SH		OTR	1	3,643	940	63
BRITISH AMERICAN TOBACCO	COMMON STOCK	110448107	317	8,468	SH		DFND	5	8,468	0	0
BROADWAY FINANCIAL CORP DE	COMMON STOCK	111444105	284	153,512	SH		DFND	1	153,512	0	0
BROOKDALE SR LIVING INC	COMMON STOCK	112463104	54	12,168	SH		DFND	1	10,818	0	1,350
BROOKDALE SR LIVING INC	COMMON STOCK	112463104	2	551	SH		OTR	1	551	0	0
BROOKFIELD ASSET MANAGE CL A	COMMON STOCK	112585104	530	12,831	SH		DFND	1	12,831	0	0
BROOKFIELD ASSET MANAGE CL A	COMMON STOCK	112585104	1,006	24,373	SH		OTR	1	18,003	558	5,812
BROOKFIELD ASSET MGMT INC	COMMON STOCK	112585104	23	560	SH		DFND	5	560	0	0
BROOKS AUTOMATION INC	COMMON STOCK	114340102	1,343	19,793	SH		DFND	1	18,851	0	942
BROOKS AUTOMATION INC	COMMON STOCK	114340102	64	947	SH		OTR	1	947	0	0
BROWN & BROWN INC	COMMON STOCK	115236101	1,106	23,335	SH		DFND	1	22,145	0	1,190
BROWN & BROWN INC	COMMON STOCK	115236101	320	6,752	SH		OTR	1	6,710	0	42
BROWN FORMAN CORP	COMMON STOCK	115637100	2,799	38,093	SH		DFND	1	27,653	0	10,440
BROWN FORMAN CORP	COMMON STOCK	115637100	403	5,484	SH		OTR	1	5,484	0	0
BROWN FORMAN CORP CLASS B	COMMON STOCK	115637209	9,692	122,012	SH		DFND	1	119,155	0	2,857
BROWN FORMAN CORP CLASS B	COMMON STOCK	115637209	1,166	14,676	SH		OTR	1	13,859	610	207
BROWN FORMAN CORP	COMMON STOCK	115637209	145	1,825	SH		DFND	5	1,698	0	127
BRUKER CORPORATION	COMMON STOCK	116794108	187	3,463	SH		DFND	1	1,698	0	1,765
BRUKER CORPORATION	COMMON STOCK	116794108	78	1,449	SH		OTR	1	0	1,373	76
BRUNSWICK CORP	COMMON STOCK	117043109	681	8,926	SH		DFND	1	8,869	0	57
BRUNSWICK CORP	COMMON STOCK	117043109	121	1,592	SH		OTR	1	0	1,592	0
BRUNSWICK CORP	COMMON STOCK	117043109	13	173	SH		DFND	5	173	0	0
BUCKLE INC	COMMON STOCK	118440106	2	61	SH		DFND	1	61	0	0
BURLINGTON STORES INC	COMMON STOCK	122017106	4,127	15,780	SH		DFND	1	14,129	0	1,651
BURLINGTON STORES INC	COMMON STOCK	122017106	1,233	4,714	SH		OTR	1	3,070	19	1,625
BURLINGTON STORES INC	COMMON STOCK	122017106	142	544	SH		DFND	5	535	0	9
BYLINE BANCORP INC	COMMON STOCK	124411109	330	21,373	SH		DFND	1	5,655	0	15,718
C A E INC	COMMON STOCK	124765108	72	2,588	SH		DFND	1	1,833	0	755
C A E INC	COMMON STOCK	124765108	4	145	SH		OTR	1	0	40	105
CBIZ INC	COMMON STOCK	124805102	7	263	SH		DFND	1	90	0	173
CF INDUSTRIES HOLDINGS INC	COMMON STOCK	125269100	361	9,342	SH		DFND	1	8,323	0	1,019
CF INDUSTRIES HOLDINGS INC	COMMON STOCK	125269100	426	11,004	SH		OTR	1	10,517	487	0
CF INDUSTRIES HOLDINGS INC	COMMON STOCK	125269100	2	49	SH		DFND	5	49	0	0
CIGNA CORP	COMMON STOCK	125523100	16,238	77,997	SH		DFND	1	74,757	0	3,240
CIGNA CORP	COMMON STOCK	125523100	12,000	57,642	SH		OTR	1	54,908	2,141	593
CIGNA CORP	COMMON STOCK	125523100	90	434	SH		DFND	5	400	0	34
CIT GROUP INC	COMMON STOCK	125581801	194	5,397	SH		DFND	1	3,892	0	1,505
CIT GROUP INC	COMMON STOCK	125581801	32	900	SH		OTR	1	67	180	653
C M S ENERGY CORP	COMMON STOCK	125896100	7,293	119,543	SH		DFND	1	117,931	0	1,612
C M S ENERGY CORP	COMMON STOCK	125896100	4,471	73,285	SH		OTR	1	71,611	133	1,541
CMS ENERGY CORPORATION	COMMON STOCK	125896100	33	537	SH		DFND	5	537	0	0
CNA FINANCIAL CORP	COMMON STOCK	126117100	72	1,842	SH		DFND	1	332	0	1,510
CNA FINANCIAL CORP	COMMON STOCK	126117100	4	107	SH		OTR	1	0	0	107
CNA FINANCIAL CORPORATION	COMMON STOCK	126117100	6	142	SH		DFND	5	142	0	0
CNOOC LTD A D R	COMMON STOCK	126132109	191	2,080	SH		DFND	1	2,080	0	0
CNOOC LTD A D R	COMMON STOCK	126132109	63	687	SH		OTR	1	687	0	0
CSG SYS INTL INC	COMMON STOCK	126349109	25	554	SH		DFND	1	554	0	0
CSG SYS INTL INC	COMMON STOCK	126349109	58	1,285	SH		OTR	1	1,285	0	0
CSW INDUSTRIALS INC	COMMON STOCK	126402106	137	1,222	SH		DFND	1	1,195	0	27
CSW INDUSTRIALS INC	COMMON STOCK	126402106	52	462	SH		OTR	1	462	0	0
CSW INDUSTRIES	COMMON STOCK	126402106	8	74	SH		DFND	5	74	0	0
CSX CORP	COMMON STOCK	126408103	21,634	238,386	SH		DFND	1	230,174	0	8,212
CSX CORP	COMMON STOCK	126408103	15,716	173,171	SH		OTR	1	169,036	1,352	2,783
CSX CORP	COMMON STOCK	126408103	47	516	SH		DFND	5	468	0	48
CTS CORP	COMMON STOCK	126501105	163	4,754	SH		DFND	1	4,754	0	0
CTS CORP	COMMON STOCK	126501105	53	1,535	SH		OTR	1	1,535	0	0
CTS CORP	COMMON STOCK	126501105	5	146	SH		DFND	5	146	0	0
CVB FINL CORP	COMMON STOCK	126600105	411	21,069	SH		DFND	1	21,069	0	0
CVB FINL CORP	COMMON STOCK	126600105	59	3,015	SH		OTR	1	3,015	0	0
CVS HEALTH CORP	COMMON STOCK	126650100	27,681	405,287	SH		DFND	1	386,940	0	18,347
CVS HEALTH CORP	COMMON STOCK	126650100	9,301	136,183	SH		OTR	1	119,660	4,739	11,784
CVS HEALTH CORP	COMMON STOCK	126650100	97	1,419	SH		DFND	5	1,312	0	107
CABOT CORP	COMMON STOCK	127055101	1,061	23,646	SH		DFND	1	21,351	0	2,295
CABOT CORP	COMMON STOCK	127055101	7	163	SH		OTR	1	163	0	0
CABOT OIL GAS CORP CL A	COMMON STOCK	127097103	32,304	1,984,274	SH		DFND	1	1,972,674	0	11,600
CABOT OIL GAS CORP CL A	COMMON STOCK	127097103	2,904	178,401	SH		OTR	1	161,873	4,225	12,303
CABOT OIL & GAS CORPORATION	COMMON STOCK	127097103	1	77	SH		DFND	5	77	0	0
CACI INTERNATIONAL INC CL A	COMMON STOCK	127190304	1,558	6,251	SH		DFND	1	5,638	0	613
CACI INTERNATIONAL INC CL A	COMMON STOCK	127190304	166	669	SH		OTR	1	511	158	0
CACI INTERNATIONAL INC	COMMON STOCK	127190304	9	38	SH		DFND	5	38	0	0
CACTUS INC A	COMMON STOCK	127203107	374	14,328	SH		DFND	1	14,122	0	206
CACTUS INC A	COMMON STOCK	127203107	116	4,446	SH		OTR	1	4,446	0	0
CADENCE DESIGN SYS INC	COMMON STOCK	127387108	2,819	20,658	SH		DFND	1	17,073	0	3,585
CADENCE DESIGN SYS INC	COMMON STOCK	127387108	414	3,029	SH		OTR	1	2,235	477	317
CADENCE DESIGN SYSTEMS INC	COMMON STOCK	127387108	38	277	SH		DFND	5	255	0	22
CAL MAINE FOODS INC	COMMON STOCK	128030202	117	3,104	SH		DFND	1	1,332	0	1,772
CAL MAINE FOODS INC	COMMON STOCK	128030202	2	65	SH		OTR	1	65	0	0
CALAMOS CONV OPPORTUNITIES & I	OTHER	128117108	48	3,552	SH		DFND	1	3,552	0	0
CALAMOS STRATEGIC TOTAL RETURN	OTHER	128125101	59	3,747	SH		DFND	1	3,747	0	0
CALAMP CORP	COMMON STOCK	128126109	22	2,248	SH		DFND	1	2,248	0	0
CALAVO GROWERS INC	COMMON STOCK	128246105	343	4,942	SH		DFND	1	4,662	0	280
CALAVO GROWERS INC	COMMON STOCK	128246105	39	565	SH		OTR	1	565	0	0
CALAVO GROWERS INC	COMMON STOCK	128246105	10	148	SH		DFND	5	148	0	0
CALERES INC	COMMON STOCK	129500104	10	637	SH		DFND	1	102	0	535
CALERES INC	COMMON STOCK	129500104	151	9,675	SH		OTR	1	9,675	0	0
CALIFORNIA WTR SVC GROUP	COMMON STOCK	130788102	183	3,380	SH		DFND	1	3,135	0	245
CALLAWAY GOLF CO	COMMON STOCK	131193104	2,263	94,270	SH		DFND	1	91,362	0	2,908
CALLAWAY GOLF CO	COMMON STOCK	131193104	4	155	SH		OTR	1	155	0	0
CAMBRIA SHAREHOLDER YIELD ETF	OTHER	132061201	13	300	SH		DFND	1	300	0	0
CAMBIA TRINITY	OTHER	132061839	107	4,345	SH		DFND	5	0	0	4,345
CAMBRIDGE BANCORP	COMMON STOCK	132152109	1,494	21,415	SH		DFND	1	21,415	0	0
CAMBRIDGE BANCORP	COMMON STOCK	132152109	224	3,205	SH		OTR	1	3,205	0	0
CAMDEN NATL CORP	COMMON STOCK	133034108	7	192	SH		DFND	1	192	0	0
CAMDEN PPTY TR SBI	COMMON STOCK	133131102	859	8,592	SH		DFND	1	8,044	0	548
CAMDEN PPTY TR SBI	COMMON STOCK	133131102	207	2,074	SH		OTR	1	1,969	5	100
CAMDEN PROPERTY TRUST	COMMON STOCK	133131102	30	296	SH		DFND	5	296	0	0
CAMPBELL SOUP CO	COMMON STOCK	134429109	961	19,873	SH		DFND	1	18,129	0	1,744
CAMPBELL SOUP CO	COMMON STOCK	134429109	886	18,322	SH		OTR	1	17,566	29	727
CAMPBELL SOUP COMPANY	COMMON STOCK	134429109	1	24	SH		DFND	5	24	0	0
CANADA GOOSE HOLDINGS INC	COMMON STOCK	135086106	2	67	SH		DFND	1	67	0	0
CANADA GOOSE HOLDINGS INC	COMMON STOCK	135086106	19	626	SH		OTR	1	0	626	0
CANADIAN IMPERIAL BK OF COMMER	COMMON STOCK	136069101	221	2,584	SH		DFND	1	1,259	0	1,325
CANADIAN IMPERIAL BK OF COMMER	COMMON STOCK	136069101	3	34	SH		OTR	1	0	0	34
CANADIAN IMPERIAL BK OF COMMRC	COMMON STOCK	136069101	350	4,097	SH		DFND	5	4,097	0	0
CANADIAN NATL RAILWAY CO	COMMON STOCK	136375102	10,439	95,033	SH		DFND	1	91,285	0	3,748
CANADIAN NATL RAILWAY CO	COMMON STOCK	136375102	2,749	25,028	SH		OTR	1	22,520	734	1,774
CANADIAN NATIONAL RAILWAY CO	COMMON STOCK	136375102	679	6,170	SH		DFND	5	6,170	0	0
CANADIAN NATURAL RESOURCES LTD	COMMON STOCK	136385101	1,736	72,173	SH		DFND	1	70,159	0	2,014
CANADIAN NATURAL RESOURCES LTD	COMMON STOCK	136385101	223	9,283	SH		OTR	1	9,081	45	157
CANADIAN NATURAL RESOURCES LTD	COMMON STOCK	136385101	19	772	SH		DFND	5	772	0	0
CANON INC SPONS A D R	COMMON STOCK	138006309	2,974	153,220	SH		DFND	1	143,984	0	9,236
CANON INC SPONS A D R	COMMON STOCK	138006309	572	29,475	SH		OTR	1	29,363	58	54
CANON INC	COMMON STOCK	138006309	7	335	SH		DFND	5	335	0	0
CANOPY GROWTH CORP	COMMON STOCK	138035100	131	5,298	SH		DFND	1	5,298	0	0
CANOPY GROWTH CORP	COMMON STOCK	138035100	49	2,000	SH		OTR	1	2,000	0	0
CANTEL MEDICAL CORP	COMMON STOCK	138098108	415	5,253	SH		DFND	1	3,986	0	1,267
CANTEL MEDICAL CORP	COMMON STOCK	138098108	126	1,604	SH		OTR	1	161	28	1,415
CAPITAL CITY BK GROUP INC	COMMON STOCK	139674105	3	107	SH		DFND	1	107	0	0
CAPITAL SOUTHWEST CORP	COMMON STOCK	140501107	243	13,700	SH		DFND	1	13,700	0	0
CARDIOVASCULAR SYSTEMS INC	COMMON STOCK	141619106	921	21,039	SH		DFND	1	21,039	0	0
CARDIOVASCULAR SYSTEMS INC	COMMON STOCK	141619106	603	13,783	SH		OTR	1	13,783	0	0
CARGURUS INC	COMMON STOCK	141788109	12	382	SH		DFND	1	290	0	92
CARLISLE COS INC	COMMON STOCK	142339100	4,000	25,613	SH		DFND	1	25,275	0	338
CARLISLE COS INC	COMMON STOCK	142339100	706	4,523	SH		OTR	1	4,480	28	15
CARLISLE COMPANIES INC	COMMON STOCK	142339100	27	172	SH		DFND	5	172	0	0
CARMAX INC	COMMON STOCK	143130102	2,435	25,770	SH		DFND	1	8,809	0	16,961
CARMAX INC	COMMON STOCK	143130102	378	3,997	SH		OTR	1	1,199	1,010	1,788
CARMAX INC	COMMON STOCK	143130102	145	1,531	SH		DFND	5	1,531	0	0
CARNIVAL CORP	COMMON STOCK	143658300	381	17,585	SH		DFND	1	14,017	0	3,568
CARNIVAL CORP	COMMON STOCK	143658300	83	3,795	SH		OTR	1	1,670	1,747	378
CARNIVAL CORPORATION	COMMON STOCK	143658300	0	8	SH		DFND	5	8	0	0
CARRIAGE SERVICES INC	COMMON STOCK	143905107	1	16	SH		DFND	5	16	0	0
CARPENTER TECHNOLOGY CORP	COMMON STOCK	144285103	133	4,573	SH		DFND	1	4,508	0	65
CARPENTER TECHNOLOGY CORP	COMMON STOCK	144285103	10	374	SH		OTR	1	324	0	50
CARTERS INC	COMMON STOCK	146229109	471	5,007	SH		DFND	1	4,223	0	784
CARTERS INC	COMMON STOCK	146229109	216	2,301	SH		OTR	1	1,107	71	1,123
CARTER INC	COMMON STOCK	146229109	33	347	SH		DFND	5	347	0	0
CARVANA CO	COMMON STOCK	146869102	71	298	SH		DFND	1	206	0	92
CARVANA CO	COMMON STOCK	146869102	33	139	SH		OTR	1	0	75	64
CARVANA CO	COMMON STOCK	146869102	1	4	SH		DFND	5	4	0	0
CASELLA WASTE SYSTEMS INC A	COMMON STOCK	147448104	309	4,989	SH		OTR	1	220	734	4,035
CASEYS GEN STORES INC	COMMON STOCK	147528103	637	3,564	SH		DFND	1	3,401	0	163
CASEYS GEN STORES INC	COMMON STOCK	147528103	10	53	SH		OTR	1	14	19	20
CASEYS GENERAL STORES INC	COMMON STOCK	147528103	12	67	SH		DFND	5	67	0	0
CATALENT INC	COMMON STOCK	148806102	490	4,707	SH		DFND	1	652	0	4,055
CATALENT INC	COMMON STOCK	148806102	134	1,289	SH		OTR	1	1,011	89	189
CATALENT INC	COMMON STOCK	148806102	135	1,300	SH		DFND	5	1,300	0	0
CATERPILLAR INC	COMMON STOCK	149123101	51,300	281,837	SH		DFND	1	277,440	0	4,397
CATERPILLAR INC	COMMON STOCK	149123101	24,106	132,433	SH		OTR	1	121,905	3,428	7,100
CATERPILLAR INC	COMMON STOCK	149123101	45	246	SH		DFND	5	202	0	44
CATHAY GENERAL BANCORP	COMMON STOCK	149150104	67	2,081	SH		DFND	1	524	0	1,557
CATHAY GENERAL BANCORP	COMMON STOCK	149150104	5	184	SH		OTR	1	46	0	138
CAVCO INDUSTRIES INC	COMMON STOCK	149568107	44	248	SH		DFND	5	248	0	0
CEDAR FAIR LP	COMMON STOCK	150185106	20	514	SH		DFND	1	514	0	0
CEL SCI CORPORATION	COMMON STOCK	150837607	11	942	SH		DFND	1	400	0	542
CELANESE CORP SER A	COMMON STOCK	150870103	1,502	11,557	SH		DFND	1	10,221	0	1,336
CELANESE CORP SER A	COMMON STOCK	150870103	600	4,624	SH		OTR	1	4,598	26	0
CEMEX SAB A D R	COMMON STOCK	151290889	1,317	254,815	SH		DFND	1	253,221	0	1,594
CEMEX SAB A D R	COMMON STOCK	151290889	326	63,127	SH		OTR	1	57,845	0	5,282
CENTRAL GARDEN & PET CO	COMMON STOCK	153527106	11	290	SH		DFND	1	33	0	257
CENTRAL PACIFIC FINANCIAL CO	COMMON STOCK	154760409	16	816	SH		DFND	1	285	0	531
CENTRAL VALLEY COMM BANCORP	COMMON STOCK	155685100	5	330	SH		DFND	1	330	0	0
CENTURY ALUMINUM COMPANY	COMMON STOCK	156431108	2	216	SH		DFND	1	216	0	0
CENTURY COMMUNITIES INC	COMMON STOCK	156504300	48	1,091	SH		DFND	1	259	0	832
CENTURY COMMUNITIES INC	COMMON STOCK	156504300	1	24	SH		OTR	1	0	0	24
CENTURYLINK INC	COMMON STOCK	156700106	380	38,976	SH		DFND	1	24,134	0	14,842
CENTURYLINK INC	COMMON STOCK	156700106	84	8,614	SH		OTR	1	2,976	0	5,638
CENTURYLINK INC	COMMON STOCK	156700106	8	773	SH		DFND	5	773	0	0
CERENCE INC	COMMON STOCK	156727109	1,961	19,517	SH		DFND	1	16,824	0	2,693
CERENCE INC	COMMON STOCK	156727109	82	822	SH		OTR	1	593	229	0
CERENCE INC	COMMON STOCK	156727109	74	740	SH		DFND	5	740	0	0
CERNER CORPORATION	COMMON STOCK	156782104	8,480	108,055	SH		DFND	1	105,562	0	2,493
CERNER CORPORATION	COMMON STOCK	156782104	5,239	66,752	SH		OTR	1	65,943	546	263
CERNER CORP	COMMON STOCK	156782104	17	216	SH		DFND	5	216	0	0
CERUS CORP	COMMON STOCK	157085101	2	228	SH		DFND	1	0	0	228
CEVA INC	COMMON STOCK	157210105	59	1,300	SH		DFND	1	1,300	0	0
CHANNELADVISOR CORP	COMMON STOCK	159179100	1	47	SH		DFND	1	47	0	0
CHARLES RIVER LABORATORIES	COMMON STOCK	159864107	4,800	19,209	SH		DFND	1	19,107	0	102
CHARLES RIVER LABORATORIES	COMMON STOCK	159864107	1,095	4,382	SH		OTR	1	4,217	43	122
CHARLES RIVER LABORATORIES	COMMON STOCK	159864107	45	182	SH		DFND	5	171	0	11
CHEESECAKE FACTORY INC	COMMON STOCK	163072101	590	15,928	SH		DFND	1	15,841	0	87
CHEESECAKE FACTORY INC	COMMON STOCK	163072101	43	1,134	SH		OTR	1	958	0	176
CHEFS WAREHOUSE HOLDINGS LLC	COMMON STOCK	163086101	719	27,997	SH		DFND	1	27,997	0	0
CHEFS WAREHOUSE HOLDINGS LLC	COMMON STOCK	163086101	79	3,060	SH		OTR	1	3,060	0	0
CHEGG INC	COMMON STOCK	163092109	780	8,633	SH		DFND	1	8,612	0	21
CHEGG INC	COMMON STOCK	163092109	7	82	SH		OTR	1	82	0	0
CHEGG INC	COMMON STOCK	163092109	22	249	SH		DFND	5	249	0	0
CHEMOURS COMPANY	COMMON STOCK	163851108	131	5,288	SH		DFND	1	3,545	0	1,743
CHEMOURS COMPANY	COMMON STOCK	163851108	85	3,443	SH		OTR	1	1,576	234	1,633
CHESAPEAKE UTILITIES CORP	COMMON STOCK	165303108	361	3,337	SH		DFND	1	3,268	0	69
CHESAPEAKE UTILITIES CORP	COMMON STOCK	165303108	58	533	SH		OTR	1	533	0	0
CHESAPEAKE UTILITIES CORP	COMMON STOCK	165303108	16	147	SH		DFND	5	147	0	0
CHEVRON CORPORATION	COMMON STOCK	166764100	99,435	1,177,436	SH		DFND	1	1,140,207	0	37,229
CHEVRON CORPORATION	COMMON STOCK	166764100	55,310	654,945	SH		OTR	1	610,438	22,606	21,901
CHEVRON CORPORATION	COMMON STOCK	166764100	539	6,384	SH		DFND	5	6,364	0	20
CHEVRON CORPORATION	COMMON STOCK	166764100	1	10	SH	Put	DFND	1	10	0	0
CHICOS FAS INC	COMMON STOCK	168615102	0	73	SH		OTR	1	73	0	0
THE CHILDRENS PLACE, INC	COMMON STOCK	168905107	14	272	SH		DFND	1	171	0	101
CHINA SOUTHN AIRLS LTD SPON A	COMMON STOCK	169409109	25	848	SH		DFND	1	675	0	173
CHINA TELECOM CORP LTD A D R	COMMON STOCK	169426103	285	10,347	SH		DFND	1	9,341	0	1,006
CHINA TELECOM CORP LTD A D R	COMMON STOCK	169426103	61	2,220	SH		OTR	1	1,352	0	868
CHIPOTLE MEXICAN GRILL INC	COMMON STOCK	169656105	2,154	1,553	SH		DFND	1	1,392	0	161
CHIPOTLE MEXICAN GRILL INC	COMMON STOCK	169656105	350	252	SH		OTR	1	200	22	30
CHIPOTLE MEXICAN GRILLE INC	COMMON STOCK	169656105	61	44	SH		DFND	5	44	0	0
CHOICE HOTELS INTL INC	COMMON STOCK	169905106	28	265	SH		DFND	1	99	0	166
CHOICE HOTELS INTL INC	COMMON STOCK	169905106	222	2,081	SH		OTR	1	2,000	81	0
CHURCH AND DWIGHT CO INC	COMMON STOCK	171340102	7,079	81,151	SH		DFND	1	77,330	0	3,821
CHURCH AND DWIGHT CO INC	COMMON STOCK	171340102	3,827	43,872	SH		OTR	1	42,867	334	671
CHURCH & DWIGHT COMPANY INC	COMMON STOCK	171340102	460	5,274	SH		DFND	5	5,235	0	39
CHURCHILL DOWNS INC	COMMON STOCK	171484108	471	2,416	SH		DFND	1	2,333	0	83
CHURCHILL DOWNS INC	COMMON STOCK	171484108	197	1,013	SH		OTR	1	0	364	649
CIENA CORP	COMMON STOCK	171779309	2,835	53,655	SH		DFND	1	51,530	0	2,125
CIENA CORP	COMMON STOCK	171779309	540	10,216	SH		OTR	1	9,979	41	196
CIENA CORP	COMMON STOCK	171779309	27	516	SH		DFND	5	477	0	39
CIMAREX ENERGY CO	COMMON STOCK	171798101	915	24,387	SH		DFND	1	20,490	0	3,897
CIMAREX ENERGY CO	COMMON STOCK	171798101	452	12,058	SH		OTR	1	8,876	2,799	383
CIMAREX ENERGY CO	COMMON STOCK	171798101	11	299	SH		DFND	5	299	0	0
CINCINNATI BELL INC	COMMON STOCK	171871502	12	754	SH		DFND	1	697	0	57
CINCINNATI BELL INC	COMMON STOCK	171871502	6	396	SH		OTR	1	396	0	0
CINCINNATI FINL CORP	COMMON STOCK	172062101	46,757	535,164	SH		DFND	1	516,850	0	18,314
CINCINNATI FINL CORP	COMMON STOCK	172062101	11,056	126,539	SH		OTR	1	123,327	0	3,212
CIRRUS LOGIC INC	COMMON STOCK	172755100	96	1,171	SH		DFND	1	934	0	237
CIRRUS LOGIC INC	COMMON STOCK	172755100	69	844	SH		OTR	1	36	711	97
CINTAS CORP	COMMON STOCK	172908105	22,715	64,265	SH		DFND	1	59,229	0	5,036
CINTAS CORP	COMMON STOCK	172908105	4,581	12,959	SH		OTR	1	11,107	1,438	414
CINTAS CORP	COMMON STOCK	172908105	200	565	SH		DFND	5	524	0	41
CITIGROUP INC	COMMON STOCK	172967424	20,686	335,504	SH		DFND	1	302,048	0	33,456
CITIGROUP INC	COMMON STOCK	172967424	6,168	100,038	SH		OTR	1	91,050	6,361	2,627
CITIGROUP INC	COMMON STOCK	172967424	90	1,451	SH		DFND	5	1,451	0	0
CITIZENS FINANCIAL GROUP	COMMON STOCK	174610105	960	26,852	SH		DFND	1	16,764	0	10,088
CITIZENS FINANCIAL GROUP	COMMON STOCK	174610105	287	8,025	SH		OTR	1	771	159	7,095
CITIZENS FINANCIAL GROUP	COMMON STOCK	174610105	43	1,202	SH		DFND	5	1,202	0	0
CITRIX SYS INC	COMMON STOCK	177376100	1,887	14,503	SH		DFND	1	12,694	0	1,809
CITRIX SYS INC	COMMON STOCK	177376100	671	5,155	SH		OTR	1	4,451	172	532
CITRIX SYSTEMS INC	COMMON STOCK	177376100	76	585	SH		DFND	5	567	0	18
CITY HLDG CO	COMMON STOCK	177835105	56	812	SH		DFND	1	454	0	358
CITY OFFICE REIT INC	COMMON STOCK	178587101	6	648	SH		DFND	1	0	0	648
CLEAN HBRS INC	COMMON STOCK	184496107	204	2,687	SH		DFND	1	217	0	2,470
CLEAN ENERGY FUELS	COMMON STOCK	184499101	39	4,915	SH		DFND	1	4,575	0	340
CLEARBRIDGE MLP AND MIDSTREAM	OTHER	184692200	207	12,359	SH		DFND	1	359	0	12,000
CLEARBRIDGE MLP AND MIDSTREAM	OTHER	184692200	45	2,700	SH		OTR	1	0	2,700	0
CLEVELAND CLIFFS INC	COMMON STOCK	185899101	242	16,639	SH		DFND	1	16,228	0	411
CLEVELAND CLIFFS INC	COMMON STOCK	185899101	12	820	SH		OTR	1	820	0	0
CLOROX CO	COMMON STOCK	189054109	15,744	77,970	SH		DFND	1	75,803	0	2,167
CLOROX CO	COMMON STOCK	189054109	13,983	69,248	SH		OTR	1	66,783	932	1,533
THE CLOROX COMPANY	COMMON STOCK	189054109	1	6	SH		DFND	5	6	0	0
CLOVIS ONCOLOGY INC	COMMON STOCK	189464100	93	19,383	SH		DFND	1	9,383	0	10,000
CLOVIS ONCOLOGY INC	COMMON STOCK	189464100	44	9,100	SH		OTR	1	9,100	0	0
COCA COLA BOTTLING CO CONS	COMMON STOCK	191098102	95	356	SH		DFND	1	300	0	56
COCA COLA BOTTLING CO CONS	COMMON STOCK	191098102	50	188	SH		OTR	1	188	0	0
COCA COLA CONSOLIDATED INC	COMMON STOCK	191098102	7	28	SH		DFND	5	28	0	0
COCA COLA COMPANY	COMMON STOCK	191216100	90,134	1,643,567	SH		DFND	1	1,587,702	0	55,865
COCA COLA COMPANY	COMMON STOCK	191216100	61,501	1,121,462	SH		OTR	1	1,012,645	63,737	45,080
COCA-COLA COMPANY	COMMON STOCK	191216100	144	2,621	SH		DFND	5	2,561	0	60
COCA COLA COMPANY	COMMON STOCK	191216100	1	30	SH	Put	DFND	1	30	0	0
COCA COLA FEMSA SAB SP A D R	COMMON STOCK	191241108	415	9,005	SH		DFND	1	7,586	0	1,419
COCA COLA FEMSA SAB SP A D R	COMMON STOCK	191241108	67	1,454	SH		OTR	1	903	0	551
CODEXIS INC	COMMON STOCK	192005106	10	465	SH		DFND	1	465	0	0
COEUR MINING INC	COMMON STOCK	192108504	5	494	SH		DFND	1	290	0	204
COEUR MINING INC	COMMON STOCK	192108504	17	1,640	SH		OTR	1	0	1,640	0
COGNEX CORP	COMMON STOCK	192422103	8,831	110,004	SH		DFND	1	100,846	0	9,158
COGNEX CORP	COMMON STOCK	192422103	927	11,550	SH		OTR	1	9,365	25	2,160
COGNEX CORP	COMMON STOCK	192422103	254	3,154	SH		DFND	5	3,154	0	0
COGNIZANT TECH SOLUTIONS CL A	COMMON STOCK	192446102	10,382	126,698	SH		DFND	1	122,495	0	4,203
COGNIZANT TECH SOLUTIONS CL A	COMMON STOCK	192446102	2,508	30,607	SH		OTR	1	27,385	2,809	413
COGNIZANT TECH SOLUTIONS CORP	COMMON STOCK	192446102	36	438	SH		DFND	5	347	0	91
COHERENT INC	COMMON STOCK	192479103	771	5,138	SH		DFND	1	1,003	0	4,135
COHERENT INC	COMMON STOCK	192479103	348	2,323	SH		OTR	1	0	2,000	323
COHU INC	COMMON STOCK	192576106	5	122	SH		DFND	1	71	0	51
COHU INC	COMMON STOCK	192576106	27	705	SH		OTR	1	705	0	0
COLFAX CORP	COMMON STOCK	194014106	92	2,398	SH		DFND	1	1,871	0	527
COLFAX CORP	COMMON STOCK	194014106	1	31	SH		OTR	1	0	0	31
COLGATE PALMOLIVE CO	COMMON STOCK	194162103	54,213	633,989	SH		DFND	1	273,639	0	360,350
COLGATE PALMOLIVE CO	COMMON STOCK	194162103	28,195	329,718	SH		OTR	1	309,931	8,593	11,194
COLGATE-PALMOLIVE CO	COMMON STOCK	194162103	6	74	SH		DFND	5	74	0	0
COLGATE PALMOLIVE CO	COMMON STOCK	194162103	0	10	SH	Put	DFND	1	10	0	0
COLLIERS INTERNATIONAL GROUP	COMMON STOCK	194693107	49	553	SH		DFND	1	436	0	117
COLUMBIA BKG SYS INC	COMMON STOCK	197236102	1,645	45,814	SH		DFND	1	45,749	0	65
COLUMBIA BKG SYS INC	COMMON STOCK	197236102	727	20,256	SH		OTR	1	19,266	990	0
COLUMBIA FINANCIAL INC	COMMON STOCK	197641103	17	1,117	SH		DFND	1	0	0	1,117
COLUMBIA PROPERTY TRUST INC	COMMON STOCK	198287203	114	7,955	SH		DFND	1	1,249	0	6,706
COLUMBIA SPORTSWEAR CO	COMMON STOCK	198516106	57	647	SH		DFND	1	425	0	222
COLUMBUS MCKINNON CORPORATION	COMMON STOCK	199333105	36	939	SH		DFND	1	939	0	0
COMFORT SYS USA INC	COMMON STOCK	199908104	752	14,279	SH		DFND	1	14,279	0	0
COMFORT SYS USA INC	COMMON STOCK	199908104	95	1,798	SH		OTR	1	1,039	759	0
COMFORT SYSTEMS USA INC	COMMON STOCK	199908104	7	126	SH		DFND	5	126	0	0
COMERICA INC	COMMON STOCK	200340107	813	14,560	SH		DFND	1	8,595	0	5,965
COMERICA INC	COMMON STOCK	200340107	261	4,672	SH		OTR	1	4,292	311	69
COMERICA INCORPORATED	COMMON STOCK	200340107	42	756	SH		DFND	5	756	0	0
COMMERCE BANCSHARES INC	COMMON STOCK	200525103	3,158	48,066	SH		DFND	1	46,912	0	1,154
COMMERCE BANCSHARES INC	COMMON STOCK	200525103	5,200	79,148	SH		OTR	1	76,539	0	2,609
COMMERCIAL METALS CO	COMMON STOCK	201723103	226	11,000	SH		DFND	1	10,410	0	590
COMMUNITY BK SYS INC	COMMON STOCK	203607106	91	1,468	SH		DFND	1	1,468	0	0
COMMUNITY HEALTH SYSTEMS INC	COMMON STOCK	203668108	5	616	SH		DFND	1	249	0	367
COMMUNITY TR BANCORP INC	COMMON STOCK	204149108	9	244	SH		DFND	1	244	0	0
COMMVAULT SYSTEMS INC	COMMON STOCK	204166102	984	17,765	SH		DFND	1	17,735	0	30
COMMVAULT SYSTEMS INC	COMMON STOCK	204166102	155	2,811	SH		OTR	1	2,715	0	96
CEMIG SA A D R	COMMON STOCK	204409601	417	144,917	SH		DFND	1	133,271	0	11,646
CEMIG SA A D R	COMMON STOCK	204409601	79	27,449	SH		OTR	1	27,012	437	0
COMPANHIA ENERGETICA DE MINAS	COMMON STOCK	204409601	7	2,406	SH		DFND	5	2,406	0	0
CIA CERVECERIAS UNIDAS A D R	COMMON STOCK	204429104	411	27,961	SH		DFND	1	26,891	0	1,070
CIA CERVECERIAS UNIDAS A D R	COMMON STOCK	204429104	49	3,340	SH		OTR	1	3,340	0	0
MINAS BUENAVENTURA A D R	COMMON STOCK	204448104	4	348	SH		DFND	1	175	0	173
COMPUTER PROGRAMS SYSTEMS	COMMON STOCK	205306103	2	80	SH		DFND	1	80	0	0
COMTECH TELECOMMUNICATIONS COR	COMMON STOCK	205826209	22	1,052	SH		DFND	1	1,000	0	52
CONAGRA BRANDS, INC	COMMON STOCK	205887102	8,860	244,330	SH		DFND	1	238,951	0	5,379
CONAGRA BRANDS, INC	COMMON STOCK	205887102	2,072	57,168	SH		OTR	1	55,491	1,254	423
CONAGRA FOODS INC.	COMMON STOCK	205887102	14	391	SH		DFND	5	217	0	174
CONCRETE PUMPING HOLDINGS IN	COMMON STOCK	206704108	85	22,225	SH		DFND	1	22,225	0	0
CONDUENT INC	COMMON STOCK	206787103	8	1,569	SH		DFND	1	1,569	0	0
CONDUENT INC	COMMON STOCK	206787103	2	400	SH		OTR	1	400	0	0
CONMED CORP	COMMON STOCK	207410101	1,309	11,690	SH		DFND	1	11,690	0	0
CONMED CORP	COMMON STOCK	207410101	38	341	SH		OTR	1	330	0	11
CONNS INC	COMMON STOCK	208242107	4	345	SH		DFND	1	0	0	345
CONSOLIDATED COMMUNICATIONS HL	COMMON STOCK	209034107	2	370	SH		OTR	1	0	0	370
CONSOLIDATED EDISON INC	COMMON STOCK	209115104	1,858	25,711	SH		DFND	1	22,828	0	2,883
CONSOLIDATED EDISON INC	COMMON STOCK	209115104	1,707	23,621	SH		OTR	1	21,614	39	1,968
CONSOLIDATED EDISON INC.	COMMON STOCK	209115104	7	97	SH		DFND	5	97	0	0
CONTAINER STORE GROUP INC	COMMON STOCK	210751103	1	140	SH		DFND	1	0	0	140
CONTINENTAL RESOURCES INC	COMMON STOCK	212015101	732	44,897	SH		DFND	1	39,458	0	5,439
CONTINENTAL RESOURCES INC	COMMON STOCK	212015101	224	13,735	SH		OTR	1	13,424	0	311
CONTINENTAL RESOURCES INC	COMMON STOCK	212015101	9	553	SH		DFND	5	553	0	0
COOPER COS INC	COMMON STOCK	216648402	5,681	15,638	SH		DFND	1	14,178	0	1,460
COOPER COS INC	COMMON STOCK	216648402	769	2,118	SH		OTR	1	2,046	36	36
COOPER COMPANIES INC (THE)	COMMON STOCK	216648402	68	187	SH		DFND	5	182	0	5
COOPER TIRE & RUBR CO	COMMON STOCK	216831107	211	5,210	SH		DFND	1	5,197	0	13
COOPER TIRE & RUBR CO	COMMON STOCK	216831107	2	38	SH		OTR	1	38	0	0
COPART INC	COMMON STOCK	217204106	487,688	3,832,521	SH		DFND	1	446,413	0	3,386,108
COPART INC	COMMON STOCK	217204106	3,653	28,705	SH		OTR	1	24,797	2,424	1,484
COPART INC	COMMON STOCK	217204106	153	1,203	SH		DFND	5	1,178	0	25
CORE MARK HLDG CO INC	COMMON STOCK	218681104	183	6,236	SH		DFND	1	6,236	0	0
CORE MARK HLDG CO INC	COMMON STOCK	218681104	8	257	SH		OTR	1	257	0	0
CORE MARK HOLDING CO INC	COMMON STOCK	218681104	9	316	SH		DFND	5	316	0	0
CORNING INC	COMMON STOCK	219350105	4,609	128,005	SH		DFND	1	120,531	0	7,474
CORNING INC	COMMON STOCK	219350105	3,607	100,192	SH		OTR	1	97,695	635	1,862
CORNING INC	COMMON STOCK	219350105	5	149	SH		DFND	5	149	0	0
COTY INC CL A	COMMON STOCK	222070203	23	3,230	SH		DFND	1	2,854	0	376
COUSINS PROPERTIES INC	COMMON STOCK	222795502	24	728	SH		DFND	1	326	0	402
CRANE CO	COMMON STOCK	224399105	759	9,784	SH		DFND	1	8,723	0	1,061
CRANE CO	COMMON STOCK	224399105	5	71	SH		OTR	1	53	0	18
CREDIT SUISSE ASSET MGMT INCCO	OTHER	224916106	3	1,000	SH		OTR	1	1,000	0	0
CREDIT ACCEPTANCE CORP	COMMON STOCK	225310101	44	126	SH		DFND	1	61	0	65
CREDIT ACCEPTANCE CORP	COMMON STOCK	225310101	7	20	SH		OTR	1	0	20	0
CREDIT SUISSE GROUP SPON A D R	COMMON STOCK	225401108	2,942	229,803	SH		DFND	1	215,301	0	14,502
CREDIT SUISSE GROUP SPON A D R	COMMON STOCK	225401108	732	57,217	SH		OTR	1	53,523	1,030	2,664
CREDIT SUISSE GROUP AG	COMMON STOCK	225401108	1	47	SH		DFND	5	47	0	0
CREE RESEARCH INC	COMMON STOCK	225447101	98	927	SH		DFND	1	702	0	225
CREE RESEARCH INC	COMMON STOCK	225447101	1	14	SH		DFND	5	14	0	0
CRESTWOOD EQUITY PARTNERS L P	COMMON STOCK	226344208	304	16,000	SH		DFND	1	16,000	0	0
CRESTWOOD EQUITY PARTNERS L P	COMMON STOCK	226344208	134	7,070	SH		OTR	1	0	0	7,070
CRITEO SA SPON A D R	COMMON STOCK	226718104	3,665	178,716	SH		DFND	1	177,425	0	1,291
CRITEO SA SPON A D R	COMMON STOCK	226718104	871	42,485	SH		OTR	1	42,123	362	0
CRITEO S A	COMMON STOCK	226718104	9	425	SH		DFND	5	425	0	0
CROCS INC	COMMON STOCK	227046109	578	9,226	SH		DFND	1	9,226	0	0
CROCS INC	COMMON STOCK	227046109	1,456	23,240	SH		OTR	1	22,452	770	18
CROWN HOLDINGS INC	COMMON STOCK	228368106	104	1,035	SH		DFND	1	781	0	254
CROWN HOLDINGS INC	COMMON STOCK	228368106	298	2,978	SH		OTR	1	2,532	193	253
CROWN HOLDINGS INC	COMMON STOCK	228368106	48	476	SH		DFND	5	450	0	26
CRYOPORT INC	COMMON STOCK	229050307	1,188	27,063	SH		DFND	1	27,063	0	0
CRYOPORT INC	COMMON STOCK	229050307	18	415	SH		OTR	1	415	0	0
CUBESMART	COMMON STOCK	229663109	648	19,274	SH		DFND	1	18,467	0	807
CUBESMART	COMMON STOCK	229663109	206	6,122	SH		OTR	1	6,087	0	35
CUBESMART	COMMON STOCK	229663109	1	28	SH		DFND	5	28	0	0
CUBIC CORP	COMMON STOCK	229669106	4	61	SH		DFND	1	49	0	12
CUBIC CORP	COMMON STOCK	229669106	46	742	SH		OTR	1	742	0	0
CULLEN FROST BANKERS INC	COMMON STOCK	229899109	135	1,557	SH		DFND	1	1,228	0	329
CULLEN FROST BANKERS INC	COMMON STOCK	229899109	136	1,554	SH		OTR	1	269	1,285	0
CUMMINS INC	COMMON STOCK	231021106	16,146	71,097	SH		DFND	1	69,190	0	1,907
CUMMINS INC	COMMON STOCK	231021106	4,771	21,008	SH		OTR	1	18,536	1,317	1,155
CUMMINS INC	COMMON STOCK	231021106	106	465	SH		DFND	5	465	0	0
CUMULUS MEDIA INC CL A	COMMON STOCK	231082801	1	100	SH		DFND	1	100	0	0
CURTISS WRIGHT CORP	COMMON STOCK	231561101	346	2,975	SH		DFND	1	2,567	0	408
CURTISS WRIGHT CORP	COMMON STOCK	231561101	204	1,755	SH		OTR	1	35	0	1,720
CUTERA INC	COMMON STOCK	232109108	26	1,092	SH		DFND	1	1,000	0	92
DEUTSCHE X-TRACKERS MSCI EM MK	OTHER	233051101	18	649	SH		DFND	1	649	0	0
DEUTSCHE XTRACKERS MSCI EAFE E	OTHER	233051200	6,181	183,889	SH		DFND	1	183,139	0	750
DEUTSCHE XTRACKERS MSCI EAFE E	OTHER	233051200	2,639	78,522	SH		OTR	1	75,022	0	3,500
XTRACKERS RUSSELL 1000 US ETF	OTHER	233051242	40	1,188	SH		DFND	1	1,188	0	0
DEUTSCHE XTRACKERS USD HIGH ET	OTHER	233051432	9	188	SH		DFND	1	188	0	0
DEUTSCHE X-TRACKERS HY CORP	OTHER	233051432	100	1,987	SH		DFND	5	1,987	0	0
DEUTSCHE XTRACKERS MSCI JAPAN	OTHER	233051507	35	764	SH		DFND	1	764	0	0
DEUTSCHE XTRACKERS MSCI JAPAN	OTHER	233051507	389	8,499	SH		OTR	1	8,499	0	0
XTRACKERS MUNICIPAL INFRAS REV	OTHER	233051705	5	175	SH		DFND	1	175	0	0
DEUTSCHE X TRACKERS MSCI EUROP	OTHER	233051853	167	5,574	SH		DFND	1	5,574	0	0
DEUTSCHE X TRACKERS MSCI EUROP	OTHER	233051853	830	27,728	SH		OTR	1	27,728	0	0
D T E ENERGY CO	COMMON STOCK	233331107	2,438	20,084	SH		DFND	1	17,801	0	2,283
D T E ENERGY CO	COMMON STOCK	233331107	1,001	8,239	SH		OTR	1	7,579	660	0
DTE ENERGY CORP	COMMON STOCK	233331107	60	490	SH		DFND	5	490	0	0
DAKTRONICS INC	COMMON STOCK	234264109	4	800	SH		DFND	1	800	0	0
DANA INCORPORATED	COMMON STOCK	235825205	6	326	SH		DFND	1	185	0	141
DANA INCORPORATED	COMMON STOCK	235825205	2	128	SH		OTR	1	0	128	0
DANAHER CORP	COMMON STOCK	235851102	35,930	161,746	SH		DFND	1	151,342	0	10,404
DANAHER CORP	COMMON STOCK	235851102	13,595	61,206	SH		OTR	1	45,480	1,666	14,060
DANAHER CORP	COMMON STOCK	235851102	171	771	SH		DFND	5	695	0	76
DARDEN RESTAURANTS INC	COMMON STOCK	237194105	3,240	27,198	SH		DFND	1	25,595	0	1,603
DARDEN RESTAURANTS INC	COMMON STOCK	237194105	3,040	25,524	SH		OTR	1	25,322	109	93
DARDEN RESTAURANTS INC	COMMON STOCK	237194105	59	499	SH		DFND	5	477	0	22
DARLING INGREDIENTS INC	COMMON STOCK	237266101	2,010	34,849	SH		DFND	1	17,015	0	17,834
DAVE BUSTERS ENTERTAINMEN	COMMON STOCK	238337109	35	1,180	SH		DFND	1	879	0	301
DAVE BUSTERS ENTERTAINMEN	COMMON STOCK	238337109	6	214	SH		OTR	1	0	214	0
DAWSON GEOPHYSICAL CO	COMMON STOCK	239360100	399	188,388	SH		DFND	1	188,388	0	0
DECKERS OUTDOOR CORP	COMMON STOCK	243537107	1,080	3,766	SH		DFND	1	3,473	0	293
DECKERS OUTDOOR CORP	COMMON STOCK	243537107	98	339	SH		OTR	1	138	201	0
DEERE CO	COMMON STOCK	244199105	49,202	182,874	SH		DFND	1	178,719	0	4,155
DEERE CO	COMMON STOCK	244199105	21,368	79,419	SH		OTR	1	73,728	801	4,890
DEERE & CO	COMMON STOCK	244199105	91	339	SH		DFND	5	339	0	0
DELTA AIR LINES INC	COMMON STOCK	247361702	16,436	408,750	SH		DFND	1	385,391	0	23,359
DELTA AIR LINES INC	COMMON STOCK	247361702	2,522	62,732	SH		OTR	1	54,146	7,542	1,044
DELUXE CORP	COMMON STOCK	248019101	730	24,989	SH		DFND	1	24,553	0	436
DELUXE CORP	COMMON STOCK	248019101	38	1,296	SH		OTR	1	800	0	496
THE DESCARTES SYSTEMS GROUP IN	COMMON STOCK	249906108	528	9,033	SH		DFND	1	5,610	0	3,423
THE DESCARTES SYSTEMS GROUP IN	COMMON STOCK	249906108	436	7,450	SH		OTR	1	1,188	1,722	4,540
DESIGNER BRANDS INC	COMMON STOCK	250565108	7	933	SH		DFND	1	0	0	933
DESIGNER BRANDS INC	COMMON STOCK	250565108	4	561	SH		OTR	1	0	561	0
DEXCOM INC	COMMON STOCK	252131107	6,713	18,157	SH		DFND	1	17,435	0	722
DEXCOM INC	COMMON STOCK	252131107	1,179	3,190	SH		OTR	1	1,160	1,985	45
DEXCOM INC	COMMON STOCK	252131107	182	491	SH		DFND	5	456	0	35
DIAMONDROCK HOSPITALITY CO	COMMON STOCK	252784301	6	681	SH		DFND	1	681	0	0
DICERNA PHARMACEUTICALS INC	COMMON STOCK	253031108	422	19,176	SH		DFND	1	19,005	0	171
DICERNA PHARMACEUTICALS INC	COMMON STOCK	253031108	6	272	SH		OTR	1	272	0	0
DICKS SPORTING GOODS INC	COMMON STOCK	253393102	188	3,336	SH		DFND	1	2,963	0	373
DICKS SPORTING GOODS INC	COMMON STOCK	253393102	185	3,286	SH		OTR	1	650	0	2,636
DIEBOLD INCORPORATED	COMMON STOCK	253651103	3	316	SH		DFND	1	316	0	0
DIGI INTERNATIONAL INC	COMMON STOCK	253798102	139	7,367	SH		DFND	1	6,567	0	800
DIGI INTERNATIONAL INC	COMMON STOCK	253798102	104	5,475	SH		OTR	1	5,075	0	400
DIGITAL REALTY TRUST INC	COMMON STOCK	253868103	41,012	293,974	SH		DFND	1	291,390	0	2,584
DIGITAL REALTY TRUST INC	COMMON STOCK	253868103	4,735	33,935	SH		OTR	1	32,996	91	848
DIGITAL REALTY TRUST INC	COMMON STOCK	253868103	55	397	SH		DFND	5	381	0	16
DIME CMNTY BANCORP INC	COMMON STOCK	253922108	5	310	SH		DFND	1	310	0	0
DILLARDS INC CL A	COMMON STOCK	254067101	4	65	SH		DFND	1	42	0	23
DINE BRANDS GLOBAL INC	COMMON STOCK	254423106	25	425	SH		DFND	1	362	0	63
DINE BRANDS GLOBAL INC	COMMON STOCK	254423106	1	17	SH		OTR	1	0	0	17
DIODES INC	COMMON STOCK	254543101	4,455	63,185	SH		DFND	1	61,477	0	1,708
DIODES INC	COMMON STOCK	254543101	901	12,780	SH		OTR	1	12,780	0	0
DIODES INC	COMMON STOCK	254543101	43	604	SH		DFND	5	563	0	41
WALT DISNEY CO THE	COMMON STOCK	254687106	172,422	951,659	SH		DFND	1	898,612	0	53,047
WALT DISNEY CO THE	COMMON STOCK	254687106	67,402	372,018	SH		OTR	1	336,885	14,049	21,084
DISNEY CO (WALT)	COMMON STOCK	254687106	593	3,266	SH		DFND	5	3,217	0	49
DISCOVER FINL SVCS	COMMON STOCK	254709108	3,423	37,805	SH		DFND	1	33,769	0	4,036
DISCOVER FINL SVCS	COMMON STOCK	254709108	1,149	12,681	SH		OTR	1	12,143	454	84
DISCOVER FINANCIAL SERVICES	COMMON STOCK	254709108	198	2,184	SH		DFND	5	2,184	0	0
DR REDDYS LABORATORIES LTD A D	COMMON STOCK	256135203	651	9,137	SH		DFND	1	4,825	0	4,312
DR REDDYS LABORATORIES LTD A D	COMMON STOCK	256135203	46	645	SH		OTR	1	0	36	609
DOCUSIGN INC	COMMON STOCK	256163106	4,753	21,382	SH		DFND	1	20,523	0	859
DOCUSIGN INC	COMMON STOCK	256163106	331	1,491	SH		OTR	1	769	650	72
DOCUSIGN INC	COMMON STOCK	256163106	1	4	SH		DFND	5	1	0	3
DOLLAR GENERAL CORP	COMMON STOCK	256677105	3,080	14,646	SH		DFND	1	12,399	0	2,247
DOLLAR GENERAL CORP	COMMON STOCK	256677105	2,139	10,171	SH		OTR	1	8,000	1,583	588
DOLLAR GENERAL CORP	COMMON STOCK	256677105	235	1,117	SH		DFND	5	1,117	0	0
DOLLAR TREE INC	COMMON STOCK	256746108	11,320	104,777	SH		DFND	1	102,081	0	2,696
DOLLAR TREE INC	COMMON STOCK	256746108	4,521	41,852	SH		OTR	1	40,376	195	1,281
DOLLAR TREE INC	COMMON STOCK	256746108	13	118	SH		DFND	5	54	0	64
DOMO INC	COMMON STOCK	257554105	3	51	SH		DFND	1	51	0	0
DOMO INC	COMMON STOCK	257554105	2	30	SH		OTR	1	0	0	30
DOMTAR CORP	COMMON STOCK	257559203	47	1,504	SH		DFND	1	243	0	1,261
DOMTAR CORP	COMMON STOCK	257559203	17	551	SH		OTR	1	551	0	0
DONALDSON CO INC	COMMON STOCK	257651109	21,836	390,765	SH		DFND	1	386,739	0	4,026
DONALDSON CO INC	COMMON STOCK	257651109	19,509	349,131	SH		OTR	1	182,414	9,644	157,073
DONALDSON CO INC	COMMON STOCK	257651109	317	144	SH	Call	DFND	1	144	0	0
RR DONNELLEY SONS CO	COMMON STOCK	257867200	1	385	SH		DFND	1	385	0	0
DORMAN PRODUCTS INC	COMMON STOCK	258278100	1,076	12,400	SH		DFND	1	10,982	0	1,418
DORMAN PRODUCTS INC	COMMON STOCK	258278100	264	3,036	SH		OTR	1	1,611	0	1,425
DORMAN PRODUCTS INC	COMMON STOCK	258278100	61	701	SH		DFND	5	701	0	0
DOVER CORP	COMMON STOCK	260003108	11,358	89,969	SH		DFND	1	86,283	0	3,686
DOVER CORP	COMMON STOCK	260003108	6,148	48,695	SH		OTR	1	46,252	2,243	200
DOVER CORP	COMMON STOCK	260003108	80	636	SH		DFND	5	636	0	0
DOW INC	COMMON STOCK	260557103	18,344	330,526	SH		DFND	1	312,447	0	18,079
DOW INC	COMMON STOCK	260557103	9,274	167,097	SH		OTR	1	154,338	3,981	8,778
DOW INC	COMMON STOCK	260557103	18	318	SH		DFND	5	318	0	0
DRIL-QUIP	COMMON STOCK	262037104	10	335	SH		DFND	5	335	0	0
DUCK CREEK TECHNOLOGIES INC	COMMON STOCK	264120106	1,716	39,636	SH		DFND	1	39,636	0	0
DUCK CREEK TECHNOLOGIES INC	COMMON STOCK	264120106	468	10,817	SH		OTR	1	10,817	0	0
DUCOMMUN INC	COMMON STOCK	264147109	59	1,100	SH		OTR	1	0	1,100	0
DUKE REALTY CORPORATION	COMMON STOCK	264411505	700	17,518	SH		DFND	1	15,160	0	2,358
DUKE REALTY CORPORATION	COMMON STOCK	264411505	59	1,482	SH		OTR	1	1,353	36	93
DUKE REALTY CORP	COMMON STOCK	264411505	13	327	SH		DFND	5	327	0	0
DYCOM INDS INC	COMMON STOCK	267475101	805	10,663	SH		DFND	1	10,287	0	376
DYCOM INDS INC	COMMON STOCK	267475101	218	2,892	SH		OTR	1	1,858	1,034	0
DYNATRACE INC	COMMON STOCK	268150109	160	3,683	SH		DFND	1	3,630	0	53
DYNATRACE INC	COMMON STOCK	268150109	143	3,319	SH		OTR	1	2,819	185	315
DYNATRACE HOLDINGS LLC	COMMON STOCK	268150109	36	825	SH		DFND	5	825	0	0
DYNAVAX TECHNOLOGIES CORP	COMMON STOCK	268158201	6	1,361	SH		DFND	1	245	0	1,116
DYNAVAX TECHNOLOGIES CORP	COMMON STOCK	268158201	3	697	SH		OTR	1	0	697	0
EL POLLO LOCO HOLDINGS INC	COMMON STOCK	268603107	2	98	SH		DFND	1	98	0	0
EAGLE BANCORP INC	COMMON STOCK	268948106	9	222	SH		DFND	1	202	0	20
EAGLE PHARMACEUTICALS INC	COMMON STOCK	269796108	313	6,730	SH		DFND	1	6,468	0	262
EAGLE PHARMACEUTICALS INC	COMMON STOCK	269796108	86	1,857	SH		OTR	1	1,857	0	0
EASTGROUP PROPERTIES INC	COMMON STOCK	277276101	1,006	7,287	SH		DFND	1	7,287	0	0
EASTGROUP PROPERTIES INC	COMMON STOCK	277276101	424	3,072	SH		OTR	1	2,672	0	400
EASTGROUP PROPERTIES INC	COMMON STOCK	277276101	2	14	SH		DFND	5	14	0	0
EASTMAN CHEM CO	COMMON STOCK	277432100	3,452	34,424	SH		DFND	1	31,510	0	2,914
EASTMAN CHEM CO	COMMON STOCK	277432100	816	8,135	SH		OTR	1	8,039	20	76
EASTMAN CHEMICAL COMPANY	COMMON STOCK	277432100	29	285	SH		DFND	5	285	0	0
EATON VANCE CORP	COMMON STOCK	278265103	1,211	17,827	SH		DFND	1	15,724	0	2,103
EATON VANCE CORP	COMMON STOCK	278265103	131	1,920	SH		OTR	1	1,717	0	203
EATON VANCE CORP	COMMON STOCK	278265103	31	460	SH		DFND	5	460	0	0
EATON VANCE	OTHER	278274105	0	0	SH		DFND	5	0	0	0
EATON VANCE ENH EQTY INC FD II	OTHER	278277108	31	1,452	SH		DFND	1	1,452	0	0
EATON VANCE ENH EQTY INC FD II	OTHER	278277108	202	9,326	SH		OTR	1	9,326	0	0
E BAY INC	COMMON STOCK	278642103	3,695	73,542	SH		DFND	1	68,038	0	5,504
E BAY INC	COMMON STOCK	278642103	2,453	48,824	SH		OTR	1	40,603	331	7,890
EBAY INC	COMMON STOCK	278642103	133	2,644	SH		DFND	5	2,644	0	0
EBIX INC	COMMON STOCK	278715206	24	629	SH		DFND	1	629	0	0
EBIX INC	COMMON STOCK	278715206	6	150	SH		OTR	1	14	136	0
ECHOSTAR CORP A	COMMON STOCK	278768106	3	165	SH		DFND	1	82	0	83
ECOLAB INC	COMMON STOCK	278865100	143,476	663,137	SH		DFND	1	574,454	0	88,683
ECOLAB INC	COMMON STOCK	278865100	79,188	366,002	SH		OTR	1	328,006	3,839	34,157
ECOLAB INC	COMMON STOCK	278865100	868	4,006	SH		DFND	5	4,006	0	0
ECOLAB INC	COMMON STOCK	278865100	0	5	SH	Put	DFND	1	5	0	0
ECOPETROL SA A D R	COMMON STOCK	279158109	121	9,329	SH		DFND	1	6,482	0	2,847
ECOPETROL SA A D R	COMMON STOCK	279158109	1	77	SH		OTR	1	0	0	77
EDISON INTERNATIONAL	COMMON STOCK	281020107	2,314	36,844	SH		DFND	1	34,265	0	2,579
EDISON INTERNATIONAL	COMMON STOCK	281020107	1,343	21,379	SH		OTR	1	20,637	722	20
EDISON INTERNATIONAL	COMMON STOCK	281020107	37	583	SH		DFND	5	473	0	110
8X8 INC	COMMON STOCK	282914100	5	132	SH		DFND	1	132	0	0
8X8 INC	COMMON STOCK	282914100	50	1,462	SH		OTR	1	0	1,462	0
ELDORADO GOLD CORP	COMMON STOCK	284902509	36	2,726	SH		DFND	1	25	0	2,701
ELECTRONIC ARTS INC	COMMON STOCK	285512109	49,726	346,278	SH		DFND	1	341,662	0	4,616
ELECTRONIC ARTS INC	COMMON STOCK	285512109	6,283	43,760	SH		OTR	1	40,744	981	2,035
ELECTRONIC ARTS INC	COMMON STOCK	285512109	247	1,717	SH		DFND	5	1,614	0	103
EMERSON ELECTRIC CO	COMMON STOCK	291011104	112,052	1,394,205	SH		DFND	1	1,361,527	0	32,678
EMERSON ELECTRIC CO	COMMON STOCK	291011104	88,004	1,094,981	SH		OTR	1	1,006,733	37,926	50,322
EMERSON ELECTRIC CO	COMMON STOCK	291011104	8	101	SH		DFND	5	70	0	31
EMERSON ELECTRIC CO	COMMON STOCK	291011104	21	116	SH	Put	DFND	1	116	0	0
EMERSON ELECTRIC CO	COMMON STOCK	291011104	29	160	SH	Put	OTR	1	0	160	0
EMERSON RADIO CORP	COMMON STOCK	291087203	2	2,000	SH		DFND	1	2,000	0	0
EMPIRE STATE REALTY TRUST IN	COMMON STOCK	292104106	64	6,913	SH		DFND	1	4,000	0	2,913
EMPIRE STATE REALTY TRUST IN	COMMON STOCK	292104106	2	183	SH		OTR	1	0	0	183
EMPLOYERS HOLDINGS INC	COMMON STOCK	292218104	7	207	SH		DFND	1	207	0	0
ENABLE MIDSTREAM PARTNERS LP	COMMON STOCK	292480100	1	130	SH		DFND	5	130	0	0
ENCORE CAPITAL GROUP INC	COMMON STOCK	292554102	5	130	SH		DFND	1	130	0	0
ENCORE CAPITAL GROUP INC	COMMON STOCK	292554102	39	990	SH		OTR	1	0	990	0
ENCORE WIRE CORP	COMMON STOCK	292562105	8	139	SH		DFND	1	108	0	31
ENERGY FUELS INC	COMMON STOCK	292671708	9	2,000	SH		DFND	1	2,000	0	0
ENERPAC TOOL GROUP CORP	COMMON STOCK	292765104	75	3,327	SH		DFND	1	202	0	3,125
ENERPLUS CORP	COMMON STOCK	292766102	218	69,844	SH		DFND	1	69,844	0	0
ENNIS INC	COMMON STOCK	293389102	27	1,510	SH		DFND	1	1,416	0	94
ENTERCOM COMMUNICATIONS CORP C	COMMON STOCK	293639100	6	2,516	SH		DFND	1	2,516	0	0
ENTERCOM COMMUNICATIONS CORP C	COMMON STOCK	293639100	0	141	SH		OTR	1	0	0	141
ENTERPRISE FINL SVCS	COMMON STOCK	293712105	96	2,734	SH		DFND	1	202	0	2,532
ENTERPRISE FINL SVCS	COMMON STOCK	293712105	62	1,783	SH		OTR	1	0	1,783	0
ENTERPRISE PRODUCTS PARTNERS	COMMON STOCK	293792107	2,773	141,536	SH		DFND	1	126,836	0	14,700
ENTERPRISE PRODUCTS PARTNERS	COMMON STOCK	293792107	2,312	117,996	SH		OTR	1	93,649	14,100	10,247
ENTERPRISE PRODUCTS PARTNERS	COMMON STOCK	293792107	30	1,543	SH		DFND	5	1,543	0	0
E PLUS INC	COMMON STOCK	294268107	138	1,564	SH		DFND	1	0	0	1,564
EQUIFAX INC	COMMON STOCK	294429105	2,174	11,275	SH		DFND	1	10,118	0	1,157
EQUIFAX INC	COMMON STOCK	294429105	314	1,633	SH		OTR	1	1,442	175	16
EQUITRANS MIDSTREAM CORP	COMMON STOCK	294600101	151	18,842	SH		DFND	1	8,452	0	10,390
EQUITRANS MIDSTREAM CORP	COMMON STOCK	294600101	1	123	SH		OTR	1	123	0	0
EQUITRANS MIDSTREAM CORP	COMMON STOCK	294600101	5	614	SH		DFND	5	614	0	0
EQUITY COMMONWEALTH	COMMON STOCK	294628102	245	8,998	SH		DFND	1	6,208	0	2,790
EQUITY COMMONWEALTH	COMMON STOCK	294628102	141	5,163	SH		OTR	1	39	170	4,954
ERICSSON LM TEL SP A D R	COMMON STOCK	294821608	4,093	342,585	SH		DFND	1	338,591	0	3,994
ERICSSON LM TEL SP A D R	COMMON STOCK	294821608	1,026	85,854	SH		OTR	1	85,472	0	382
ERICCSON (LM) TEL	COMMON STOCK	294821608	8	658	SH		DFND	5	658	0	0
E S C O TECHNOLOGIES INC	COMMON STOCK	296315104	2,425	23,493	SH		DFND	1	20,119	0	3,374
E S C O TECHNOLOGIES INC	COMMON STOCK	296315104	349	3,384	SH		OTR	1	1,258	486	1,640
ESPEY MFG & ELECTRONICS CORP	COMMON STOCK	296650104	43	2,283	SH		DFND	1	2,283	0	0
ESSEX PROPERTY TRUST INC	COMMON STOCK	297178105	505	2,128	SH		DFND	1	1,740	0	388
ESSEX PROPERTY TRUST INC	COMMON STOCK	297178105	126	534	SH		OTR	1	507	27	0
ESSEX PROPERTY TRUST INC	COMMON STOCK	297178105	8	35	SH		DFND	5	35	0	0
EURONET WORLDWIDE INC	COMMON STOCK	298736109	1,226	8,463	SH		DFND	1	8,024	0	439
EURONET WORLDWIDE INC	COMMON STOCK	298736109	466	3,214	SH		OTR	1	2,810	394	10
EURONET WORLDWIDE INC	COMMON STOCK	298736109	40	275	SH		DFND	5	275	0	0
ROBO GLOBAL ROBOTICS ETF	OTHER	301505707	63	1,035	SH		OTR	1	1,035	0	0
EMQQ EMERGING MARKETS ETF	OTHER	301505889	131	2,067	SH		DFND	1	2,067	0	0
EXLSERVICE HOLDINGS INC	COMMON STOCK	302081104	1,077	12,655	SH		DFND	1	9,031	0	3,624
EXLSERVICE HOLDINGS INC	COMMON STOCK	302081104	115	1,346	SH		OTR	1	1,346	0	0
EXLSERVICE HOLDINGS INC	COMMON STOCK	302081104	12	143	SH		DFND	5	143	0	0
EXONE CO	COMMON STOCK	302104104	5	500	SH		DFND	1	500	0	0
EXPEDITORS INTL WASH INC	COMMON STOCK	302130109	12,296	129,290	SH		DFND	1	107,208	0	22,082
EXPEDITORS INTL WASH INC	COMMON STOCK	302130109	8,855	93,092	SH		OTR	1	85,613	4,961	2,518
EXPEDITORS INTL WASH INC	COMMON STOCK	302130109	16	171	SH		DFND	5	171	0	0
FLIR SYSTEMS INC	COMMON STOCK	302445101	910	20,743	SH		DFND	1	18,069	0	2,674
FLIR SYSTEMS INC	COMMON STOCK	302445101	186	4,250	SH		OTR	1	4,193	57	0
F M C CORPORATION	COMMON STOCK	302491303	1,291	11,234	SH		DFND	1	10,005	0	1,229
F M C CORPORATION	COMMON STOCK	302491303	734	6,386	SH		OTR	1	3,895	1,422	1,069
FMC CORP	COMMON STOCK	302491303	54	470	SH		DFND	5	470	0	0
FNB CORP	COMMON STOCK	302520101	1,008	106,176	SH		DFND	1	88,388	0	17,788
FNB CORP	COMMON STOCK	302520101	44	4,612	SH		OTR	1	4,612	0	0
FS KKR CAPITAL CORP COM	COMMON STOCK	302635206	8	457	SH		DFND	1	457	0	0
FS KKR CAPITAL CORP COM	COMMON STOCK	302635206	33	1,985	SH		OTR	1	1,000	985	0
FTI CONSULTING INC	COMMON STOCK	302941109	21	188	SH		DFND	1	188	0	0
FTI CONSULTING INC	COMMON STOCK	302941109	22	194	SH		OTR	1	0	0	194
FACTSET RESEARCH SYSTEMS INC	COMMON STOCK	303075105	3,319	9,982	SH		DFND	1	9,793	0	189
FACTSET RESEARCH SYSTEMS INC	COMMON STOCK	303075105	1,824	5,485	SH		OTR	1	5,458	27	0
FACTSET RESEARCH SYSTEMS INC	COMMON STOCK	303075105	182	547	SH		DFND	5	547	0	0
FAIR ISAAC CORPORATION	COMMON STOCK	303250104	2,917	5,708	SH		DFND	1	3,614	0	2,094
FAIR ISAAC CORPORATION	COMMON STOCK	303250104	427	836	SH		OTR	1	259	567	10
FAIR ISAAC & CO INC	COMMON STOCK	303250104	437	856	SH		DFND	5	856	0	0
FARMER BROS CO	COMMON STOCK	307675108	110	23,603	SH		DFND	1	23,603	0	0
FARO TECHNOLOGIES INC	COMMON STOCK	311642102	22	307	SH		DFND	1	307	0	0
FARO TECHNOLOGIES INC	COMMON STOCK	311642102	46	656	SH		OTR	1	55	601	0
FASTENAL CO	COMMON STOCK	311900104	21,805	446,537	SH		DFND	1	386,658	0	59,879
FASTENAL CO	COMMON STOCK	311900104	9,593	196,466	SH		OTR	1	191,281	423	4,762
FASTENAL COMPANY	COMMON STOCK	311900104	396	8,101	SH		DFND	5	8,101	0	0
FAUQUIER BANKSHS INC	COMMON STOCK	312059108	480	27,600	SH		DFND	1	27,600	0	0
FEDERAL AGRIC MTG CORP CL A	COMMON STOCK	313148108	163	2,500	SH		DFND	1	2,500	0	0
FEDERAL AGRIC MTG CORP CL C	COMMON STOCK	313148306	15	200	SH		DFND	1	182	0	18
FEDERAL AGRIC MTG CORP CL C	COMMON STOCK	313148306	1	13	SH		OTR	1	0	13	0
FEDERAL AGRICULTURE MTG CORP	COMMON STOCK	313148306	9	125	SH		DFND	5	125	0	0
FEDERAL RLTY INVT TR SBI NEW	COMMON STOCK	313747206	273	3,208	SH		DFND	1	1,976	0	1,232
FEDERAL RLTY INVT TR SBI NEW	COMMON STOCK	313747206	56	660	SH		OTR	1	574	10	76
FEDERAL REALTY INVESTMENT TR	COMMON STOCK	313747206	1	16	SH		DFND	5	16	0	0
FEDERAL SIGNAL CORP	COMMON STOCK	313855108	91	2,745	SH		DFND	1	2,745	0	0
FEDERAL SIGNAL CORP	COMMON STOCK	313855108	71	2,133	SH		OTR	1	0	2,133	0
FEDERAL SIGNAL CORP	COMMON STOCK	313855108	40	1,219	SH		DFND	5	1,219	0	0
FEDERATED HERMES INC	COMMON STOCK	314211103	143	4,959	SH		DFND	1	4,263	0	696
FEDERATED HERMES INC	COMMON STOCK	314211103	13	427	SH		OTR	1	201	0	226
FERRO CORP	COMMON STOCK	315405100	973	66,551	SH		DFND	1	66,551	0	0
FERRO CORP	COMMON STOCK	315405100	99	6,756	SH		OTR	1	6,756	0	0
F5 NETWORKS INC	COMMON STOCK	315616102	1,093	6,210	SH		DFND	1	4,695	0	1,515
F5 NETWORKS INC	COMMON STOCK	315616102	187	1,063	SH		OTR	1	923	45	95
FIDELITY NASDAQ COMP INDX TS E	OTHER	315912808	136	272	SH		DFND	1	272	0	0
FIDELITY MSCI CONSUMER DISCRET	OTHER	316092204	671	9,384	SH		OTR	1	0	9,384	0
FIDELITY MSCI REAL ESTATE ETF	OTHER	316092857	49	1,916	SH		DFND	1	1,916	0	0
FIFTH THIRD BANCORP	COMMON STOCK	316773100	11,510	417,493	SH		DFND	1	404,464	0	13,029
FIFTH THIRD BANCORP	COMMON STOCK	316773100	4,810	174,474	SH		OTR	1	173,414	578	482
FIFTH THIRD BANCORP	COMMON STOCK	316773100	39	1,412	SH		DFND	5	1,412	0	0
51JOB INC A D R	COMMON STOCK	316827104	112	1,599	SH		DFND	1	1,144	0	455
51JOB INC A D R	COMMON STOCK	316827104	1	17	SH		OTR	1	17	0	0
FIRST BANCORP PUERTO RICO	COMMON STOCK	318672706	26	2,829	SH		DFND	1	2,829	0	0
FIRST BANCORP NC	COMMON STOCK	318910106	31	907	SH		DFND	1	355	0	552
FIRST BANCORP NC	COMMON STOCK	318910106	5	150	SH		OTR	1	150	0	0
THE FIRST BANCSHARES INC	COMMON STOCK	318916103	8	271	SH		DFND	1	271	0	0
FIRST BUSEY CORP	COMMON STOCK	319383204	140	6,505	SH		DFND	1	6,166	0	339
FIRST BUSINESS FINL SVCS INC	COMMON STOCK	319390100	37	2,000	SH		DFND	1	2,000	0	0
FIRST COMWLTH FINL CORP	COMMON STOCK	319829107	1	79	SH		DFND	1	0	0	79
FIRST FINANCIAL BANCORP	COMMON STOCK	320209109	2,523	143,931	SH		DFND	1	143,931	0	0
FIRST FINANCIAL BANCORP	COMMON STOCK	320209109	1,238	70,629	SH		OTR	1	70,629	0	0
FIRST FINL CORP IND	COMMON STOCK	320218100	6	161	SH		OTR	1	161	0	0
FIRST HORIZON CORPORATION	COMMON STOCK	320517105	1,024	80,167	SH		DFND	1	54,572	0	25,595
FIRST HORIZON CORPORATION	COMMON STOCK	320517105	67	5,253	SH		OTR	1	4,344	700	209
FIRST HORIZON NATIONAL CORP	COMMON STOCK	320517105	22	1,739	SH		DFND	5	1,739	0	0
FIRST LONG ISLAND CORP	COMMON STOCK	320734106	25	1,381	SH		DFND	1	1,381	0	0
FIRST MERCHANTS CORP	COMMON STOCK	320817109	583	15,593	SH		DFND	1	15,090	0	503
FIRST MERCHANTS CORP	COMMON STOCK	320817109	27	731	SH		OTR	1	0	731	0
FIRST MID BANCSHARES INC	COMMON STOCK	320866106	2	45	SH		DFND	1	45	0	0
FIRST MIDWEST BANCORP INC DEL	COMMON STOCK	320867104	522	32,823	SH		DFND	1	30,161	0	2,662
FIRST MIDWEST BANCORP INC DEL	COMMON STOCK	320867104	17	1,086	SH		OTR	1	1,086	0	0
FIRST SOLAR INC	COMMON STOCK	336433107	2,471	24,981	SH		DFND	1	24,000	0	981
FIRST SOLAR INC	COMMON STOCK	336433107	667	6,744	SH		OTR	1	6,695	42	7
FIRST SOLAR INC	COMMON STOCK	336433107	2	21	SH		DFND	5	21	0	0
1ST SOURCE CORP	COMMON STOCK	336901103	89	2,200	SH		DFND	1	2,200	0	0
FIRST TRUST MORN DVD LEAD IN E	OTHER	336917109	62	2,084	SH		DFND	1	2,084	0	0
FIRST TRUST MORN DVD LEAD IN E	OTHER	336917109	22	750	SH		OTR	1	750	0	0
FIRST TRUST MORNINGSTAR DIVID	OTHER	336917109	3,033	102,484	SH		DFND	5	0	0	102,484
FIRST TR GBL ENGIN AND CONST E	OTHER	336920103	58	491	SH		DFND	1	491	0	0
FIRST TRUST IPOX 100 INDX FUND	OTHER	336920103	76	640	SH		DFND	5	36	0	604
FIRST TRUST ABERDEEN GLOBAL	OTHER	337319107	13	1,225	SH		DFND	1	1,225	0	0
FIRST TR NASDQ 100 EQUAL WGHTD	OTHER	337344105	153	1,518	SH		DFND	1	818	0	700
FIRST TR NASDQ 100 EQUAL WGHTD	OTHER	337344105	151	1,500	SH		OTR	1	1,500	0	0
FIRST TRUST NASDAQ 100 TECH	OTHER	337345102	2,749	19,913	SH		DFND	5	7,299	0	12,614
FISERV INC	COMMON STOCK	337738108	27,262	239,439	SH		DFND	1	223,100	0	16,339
FISERV INC	COMMON STOCK	337738108	6,213	54,563	SH		OTR	1	41,447	10,765	2,351
FISERV INC	COMMON STOCK	337738108	80	701	SH		DFND	5	701	0	0
FLAGSTAR BANCORP INC	COMMON STOCK	337930705	1	31	SH		DFND	1	0	0	31
FIRST ENERGY CORP	COMMON STOCK	337932107	1,212	39,579	SH		DFND	1	37,919	0	1,660
FIRST ENERGY CORP	COMMON STOCK	337932107	320	10,446	SH		OTR	1	10,391	0	55
FIRSTENERGY CORP	COMMON STOCK	337932107	0	0	SH		DFND	5	0	0	0
FIVE9 INC	COMMON STOCK	338307101	1,908	10,938	SH		DFND	1	10,784	0	154
FIVE9 INC	COMMON STOCK	338307101	385	2,205	SH		OTR	1	1,052	255	898
FIVE9 INC	COMMON STOCK	338307101	3	15	SH		DFND	5	0	0	15
FLAHERTY CRUMRINE PREFERRED	OTHER	338478100	257	10,976	SH		DFND	1	10,976	0	0
FLAHERTY CRUMRINE PREFERRED	OTHER	338478100	120	5,132	SH		OTR	1	5,132	0	0
FLAHERTY CRUMRINE PREFERRED AN	OTHER	338480106	6	300	SH		DFND	1	300	0	0
FLEETCOR TECHNOLOGIES INC	COMMON STOCK	339041105	544	1,996	SH		DFND	1	1,529	0	467
FLEETCOR TECHNOLOGIES INC	COMMON STOCK	339041105	266	975	SH		OTR	1	850	41	84
FLOOR DECOR HOLDINGS INC A	COMMON STOCK	339750101	1,596	17,186	SH		DFND	1	16,920	0	266
FLOOR DECOR HOLDINGS INC A	COMMON STOCK	339750101	543	5,853	SH		OTR	1	4,715	1,138	0
FLOOR & DECOR HOLDINGS INC -A	COMMON STOCK	339750101	48	522	SH		DFND	5	494	0	28
FLUOR CORP	COMMON STOCK	343412102	192	12,019	SH		DFND	1	8,989	0	3,030
FLUOR CORP	COMMON STOCK	343412102	90	5,666	SH		OTR	1	5,521	145	0
FLOWERS FOODS INC	COMMON STOCK	343498101	982	43,405	SH		DFND	1	42,671	0	734
FLOWERS FOODS INC	COMMON STOCK	343498101	193	8,527	SH		OTR	1	4,603	15	3,909
FLUSHING FINANCIAL CORPORATION	COMMON STOCK	343873105	25	1,522	SH		DFND	1	1,373	0	149
FOLEY TRASIMENE ACQUISITION CO	COMMON STOCK	344328109	8	505	SH		DFND	1	505	0	0
FOMENTO ECONOMICO MEX SP A D R	COMMON STOCK	344419106	3,797	50,125	SH		DFND	1	48,484	0	1,641
FOMENTO ECONOMICO MEX SP A D R	COMMON STOCK	344419106	791	10,434	SH		OTR	1	9,781	645	8
FOMENTO ECONOMICO MEX SAB D CV	COMMON STOCK	344419106	7	97	SH		DFND	5	97	0	0
FOOT LOCKER INC	COMMON STOCK	344849104	5,277	130,478	SH		DFND	1	128,700	0	1,778
FOOT LOCKER INC	COMMON STOCK	344849104	1,156	28,576	SH		OTR	1	27,989	486	101
FORD MOTOR CO	COMMON STOCK	345370860	5,313	604,500	SH		DFND	1	587,644	0	16,856
FORD MOTOR CO	COMMON STOCK	345370860	419	47,684	SH		OTR	1	34,896	0	12,788
FORD MOTOR CO	COMMON STOCK	345370860	3	289	SH		DFND	5	289	0	0
FORESTAR GROUP INC	COMMON STOCK	346232101	6	279	SH		DFND	1	279	0	0
FORMFACTOR INC	COMMON STOCK	346375108	5	109	SH		DFND	1	60	0	49
FORMFACTOR INC	COMMON STOCK	346375108	235	5,475	SH		OTR	1	4,636	839	0
FORRESTER RESH INC	COMMON STOCK	346563109	31	750	SH		DFND	1	750	0	0
FORRESTER RESH INC	COMMON STOCK	346563109	12	275	SH		OTR	1	275	0	0
FORTIS INC	COMMON STOCK	349553107	479	11,730	SH		DFND	1	7,742	0	3,988
FORTIS INC	COMMON STOCK	349553107	65	1,596	SH		OTR	1	1,530	21	45
FORWARD AIR CORP	COMMON STOCK	349853101	410	5,333	SH		DFND	1	5,321	0	12
FORWARD AIR CORP	COMMON STOCK	349853101	78	1,019	SH		OTR	1	1,019	0	0
FORWARD AIR CORPORATION	COMMON STOCK	349853101	11	138	SH		DFND	5	138	0	0
FOSTER L B CO CL A	COMMON STOCK	350060109	7	457	SH		DFND	1	457	0	0
FOUNDATION BUILDING	COMMON STOCK	350392106	2	108	SH		DFND	1	108	0	0
FRANCO NEVADA CORP	COMMON STOCK	351858105	271	2,166	SH		DFND	1	1,656	0	510
FRANCO NEVADA CORP	COMMON STOCK	351858105	2,568	20,485	SH		OTR	1	20,316	18	151
FRANCO NEVADA CORP	COMMON STOCK	351858105	1	6	SH		DFND	5	6	0	0
FRANKLIN ELEC INC	COMMON STOCK	353514102	635	9,173	SH		DFND	1	9,173	0	0
FRANKLIN ELEC INC	COMMON STOCK	353514102	72	1,034	SH		OTR	1	1,034	0	0
FRANKLIN ELECTRIC CO INC	COMMON STOCK	353514102	13	195	SH		DFND	5	195	0	0
FRANKLIN RES INC	COMMON STOCK	354613101	442	17,661	SH		DFND	1	14,382	0	3,279
FRANKLIN RES INC	COMMON STOCK	354613101	88	3,514	SH		OTR	1	725	2,740	49
FRANKLIN RESOURCES INC	COMMON STOCK	354613101	2	63	SH		DFND	5	63	0	0
FRESENIUS AKTIENGESELLSCHAFT A	COMMON STOCK	358029106	704	16,942	SH		DFND	1	15,003	0	1,939
FRESENIUS AKTIENGESELLSCHAFT A	COMMON STOCK	358029106	110	2,630	SH		OTR	1	2,417	161	52
FRESENIUS MEDICAL CARE AG & CO	COMMON STOCK	358029106	1	13	SH		DFND	5	13	0	0
FRESHPET INC	COMMON STOCK	358039105	989	6,963	SH		DFND	1	6,570	0	393
FRESHPET INC	COMMON STOCK	358039105	239	1,680	SH		OTR	1	806	874	0
FUEL TECH, INC.	COMMON STOCK	359523107	945	243,476	SH		DFND	1	243,476	0	0
FULGENT GENETICS INC	COMMON STOCK	359664109	27	512	SH		DFND	1	512	0	0
FULLER H B CO	COMMON STOCK	359694106	2,445	47,128	SH		DFND	1	46,118	0	1,010
FULLER H B CO	COMMON STOCK	359694106	3,328	64,146	SH		OTR	1	63,246	0	900
FULTON FINL CORP PA	COMMON STOCK	360271100	17	1,300	SH		DFND	1	105	0	1,195
FULTON FINL CORP PA	COMMON STOCK	360271100	1	106	SH		OTR	1	106	0	0
GAMCO INVS INC	COMMON STOCK	361438104	124	7,000	SH		DFND	1	7,000	0	0
G A T X CORP	COMMON STOCK	361448103	1,371	16,487	SH		DFND	1	15,644	0	843
G A T X CORP	COMMON STOCK	361448103	345	4,148	SH		OTR	1	4,091	0	57
GATX CORP	COMMON STOCK	361448103	25	302	SH		DFND	5	302	0	0
GABELLI EQUITY TR INC	OTHER	362397101	118	18,781	SH		DFND	1	18,781	0	0
GABELLI EQUITY TR INC	OTHER	362397101	30	4,822	SH		OTR	1	4,822	0	0
ARTHUR J GALLAGHER CO	COMMON STOCK	363576109	28,903	233,632	SH		DFND	1	232,213	0	1,419
ARTHUR J GALLAGHER CO	COMMON STOCK	363576109	314	2,538	SH		OTR	1	2,538	0	0
GALLAGHER (ARTHUR J.) & CO	COMMON STOCK	363576109	173	1,400	SH		DFND	5	1,400	0	0
THE GAP INC	COMMON STOCK	364760108	130	6,429	SH		DFND	1	3,236	0	3,193
THE GAP INC	COMMON STOCK	364760108	34	1,712	SH		OTR	1	1,305	0	407
GARTNER INC	COMMON STOCK	366651107	246	1,539	SH		DFND	1	1,149	0	390
GARTNER INC	COMMON STOCK	366651107	21	132	SH		OTR	1	0	114	18
GARTNER GROUP	COMMON STOCK	366651107	58	362	SH		DFND	5	362	0	0
GENERAC HOLDINGS INC	COMMON STOCK	368736104	6,809	29,940	SH		DFND	1	27,619	0	2,321
GENERAC HOLDINGS INC	COMMON STOCK	368736104	1,103	4,849	SH		OTR	1	4,849	0	0
GENERAC HOLDINGS INC	COMMON STOCK	368736104	78	341	SH		DFND	5	327	0	14
GENERAL DYNAMICS CORP	COMMON STOCK	369550108	15,031	100,997	SH		DFND	1	96,102	0	4,895
GENERAL DYNAMICS CORP	COMMON STOCK	369550108	4,001	26,888	SH		OTR	1	23,287	2,198	1,403
GENERAL DYNAMICS CORP	COMMON STOCK	369550108	121	811	SH		DFND	5	799	0	12
GENERAL ELECTRIC CO	COMMON STOCK	369604103	34,248	3,171,055	SH		DFND	1	2,525,558	0	645,497
GENERAL ELECTRIC CO	COMMON STOCK	369604103	11,175	1,034,694	SH		OTR	1	989,611	16,609	28,474
GENERAL ELECTRIC CO	COMMON STOCK	369604103	4	327	SH		DFND	5	327	0	0
GENERAL MILLS INC	COMMON STOCK	370334104	84,347	1,434,460	SH		DFND	1	1,410,243	0	24,217
GENERAL MILLS INC	COMMON STOCK	370334104	58,975	1,002,981	SH		OTR	1	911,614	31,416	59,951
GENERAL MILLS INC	COMMON STOCK	370334104	164	2,787	SH		DFND	5	2,787	0	0
GENESCO INC	COMMON STOCK	371532102	11	366	SH		DFND	1	236	0	130
GENTEX CORP	COMMON STOCK	371901109	2,915	85,899	SH		DFND	1	85,175	0	724
GENTEX CORP	COMMON STOCK	371901109	689	20,296	SH		OTR	1	19,854	240	202
GENTEX CORP	COMMON STOCK	371901109	159	4,671	SH		DFND	5	4,671	0	0
GENESIS ENERGY LP	COMMON STOCK	371927104	9	1,500	SH		DFND	1	1,500	0	0
GENMAB A S SP A D R	COMMON STOCK	372303206	847	20,829	SH		DFND	1	14,529	0	6,300
GENMAB A S SP A D R	COMMON STOCK	372303206	86	2,099	SH		OTR	1	1,249	750	100
GENMAB A/S	COMMON STOCK	372303206	3	62	SH		DFND	5	62	0	0
GENMARK DIAGNOSTICS INC	COMMON STOCK	372309104	10	685	SH		DFND	1	685	0	0
GENUINE PARTS COMPANY	COMMON STOCK	372460105	2,299	22,894	SH		DFND	1	21,358	0	1,536
GENUINE PARTS COMPANY	COMMON STOCK	372460105	902	8,972	SH		OTR	1	7,270	1,702	0
GENUINE PARTS CO	COMMON STOCK	372460105	47	468	SH		DFND	5	468	0	0
GERDAU S A SPON A D R	COMMON STOCK	373737105	35	7,368	SH		DFND	1	697	0	6,671
GERDAU SA	COMMON STOCK	373737105	2	535	SH		DFND	5	535	0	0
GERMAN AMERICAN BANCORP	COMMON STOCK	373865104	20	591	SH		DFND	1	591	0	0
GERMAN AMERICAN BANCORP	COMMON STOCK	373865104	405	12,245	SH		OTR	1	12,245	0	0
GERON CORP	COMMON STOCK	374163103	3	1,716	SH		DFND	1	1,716	0	0
GETTY RLTY CORP NEW	COMMON STOCK	374297109	130	4,731	SH		DFND	1	4,731	0	0
GETTY RLTY CORP NEW	COMMON STOCK	374297109	46	1,673	SH		OTR	1	1,658	15	0
GETTY REALTY CORP	COMMON STOCK	374297109	6	210	SH		DFND	5	210	0	0
GEVO INC	COMMON STOCK	374396406	14	3,200	SH		DFND	1	3,200	0	0
GIBRALTAR INDS INC	COMMON STOCK	374689107	1	9	SH		DFND	1	0	0	9
GIBRALTAR INDUSTRIES INC	COMMON STOCK	374689107	75	1,040	SH		DFND	5	1,040	0	0
GILEAD SCIENCES INC	COMMON STOCK	375558103	73,517	1,261,897	SH		DFND	1	1,240,786	0	21,111
GILEAD SCIENCES INC	COMMON STOCK	375558103	10,244	175,823	SH		OTR	1	163,962	9,328	2,533
GILEAD SCIENCES INC	COMMON STOCK	375558103	349	5,992	SH		DFND	5	5,951	0	41
GILEAD SCIENCES INC	COMMON STOCK	375558103	0	1	SH	Put	DFND	1	1	0	0
GILDAN ACTIVEWEAR INC	COMMON STOCK	375916103	160	5,712	SH		DFND	1	4,527	0	1,185
GILDAN ACTIVEWEAR INC	COMMON STOCK	375916103	5	187	SH		OTR	1	0	0	187
GLADSTONE CAP CORP	COMMON STOCK	376535100	30	3,343	SH		OTR	1	3,343	0	0
GLADSTONE COMMERCIAL CORP	COMMON STOCK	376536108	15	813	SH		DFND	1	718	0	95
GLADSTONE INVESTMENT CORP	COMMON STOCK	376546107	2	155	SH		DFND	1	0	0	155
GLADSTONE LAND CORP	COMMON STOCK	376549101	146	10,000	SH		OTR	1	10,000	0	0
GLATFELTER CORPORATION	COMMON STOCK	377320106	515	31,412	SH		DFND	1	31,412	0	0
GLATFELTER CORPORATION	COMMON STOCK	377320106	14	857	SH		OTR	1	857	0	0
GLAUKOS CORP	COMMON STOCK	377322102	43	573	SH		DFND	1	573	0	0
GLAUKOS CORP	COMMON STOCK	377322102	2	32	SH		OTR	1	17	0	15
GLOBALSTAR INC	COMMON STOCK	378973408	0	96	SH		DFND	1	96	0	0
GLOBAL NET LEASE INC	COMMON STOCK	379378201	27	1,560	SH		DFND	1	1,026	0	534
GLOBUS MED INCA	COMMON STOCK	379577208	447	6,860	SH		DFND	1	4,187	0	2,673
GLOBUS MED INCA	COMMON STOCK	379577208	13	206	SH		OTR	1	175	16	15
GLOBUS MEDICAL INC	COMMON STOCK	379577208	43	662	SH		DFND	5	662	0	0
GLU MOBILE INC	COMMON STOCK	379890106	4	400	SH		OTR	1	0	0	400
GLU MOBILE INC	COMMON STOCK	379890106	0	9	SH		DFND	5	9	0	0
GODADDY INC CLASS A	COMMON STOCK	380237107	3,359	40,495	SH		DFND	1	39,858	0	637
GODADDY INC CLASS A	COMMON STOCK	380237107	1,606	19,358	SH		OTR	1	11,926	722	6,710
GODADDY INC	COMMON STOCK	380237107	336	4,052	SH		DFND	5	4,052	0	0
GOLDEN ENTERTAINMENT INC	COMMON STOCK	381013101	52	2,594	SH		DFND	1	2,535	0	59
GOLDMAN SACHS ACTIVEBETA ETF	OTHER	381430107	79	2,483	SH		DFND	1	373	0	2,110
GOLDMAN SACHS JUST U S LARGE C	OTHER	381430396	19	349	SH		DFND	1	349	0	0
GOLDMAN ACTIVEBETA US LC ETF	OTHER	381430503	137	1,808	SH		DFND	1	409	0	1,399
GOLDMAN SACHS ACCESS TREASURY	OTHER	381430529	10	100	SH		OTR	1	100	0	0
GOODYEAR TIRE RUBBER CO	COMMON STOCK	382550101	189	17,313	SH		DFND	1	11,740	0	5,573
GOODYEAR TIRE RUBBER CO	COMMON STOCK	382550101	6	576	SH		OTR	1	576	0	0
GORMAN RUPP CO	COMMON STOCK	383082104	1,768	54,490	SH		DFND	1	54,490	0	0
GRACO INC	COMMON STOCK	384109104	8,634	119,344	SH		DFND	1	112,310	0	7,034
GRACO INC	COMMON STOCK	384109104	11,430	157,974	SH		OTR	1	154,568	128	3,278
GRACO INC	COMMON STOCK	384109104	655	9,041	SH		DFND	5	9,041	0	0
GRAFTECH INTERNATIONAL LTD	COMMON STOCK	384313508	3	285	SH		DFND	1	0	0	285
GRAHAM CORP	COMMON STOCK	384556106	6	379	SH		DFND	1	379	0	0
GRAHAM HOLDINGS CO	COMMON STOCK	384637104	186	349	SH		DFND	1	252	0	97
GRAHAM HOLDINGS CO	COMMON STOCK	384637104	1	2	SH		OTR	1	2	0	0
GRAINGER W W INC	COMMON STOCK	384802104	17,337	42,458	SH		DFND	1	41,762	0	696
GRAINGER W W INC	COMMON STOCK	384802104	3,988	9,765	SH		OTR	1	8,901	67	797
W W GRAINGER INC	COMMON STOCK	384802104	474	1,157	SH		DFND	5	1,157	0	0
GRANITE CONSTRUCTION INC	COMMON STOCK	387328107	61	2,280	SH		DFND	1	2,210	0	70
GRANITE CONSTRUCTION INC	COMMON STOCK	387328107	3	119	SH		OTR	1	0	119	0
GRAPHIC PACKAGING HLDG CO	COMMON STOCK	388689101	718	42,410	SH		DFND	1	28,021	0	14,389
GRAPHIC PACKAGING HLDG CO	COMMON STOCK	388689101	218	12,857	SH		OTR	1	347	0	12,510
GRAY TELEVISION INC	COMMON STOCK	389375106	163	9,105	SH		DFND	1	5,105	0	4,000
GRAY TELEVISION INC	COMMON STOCK	389375106	13	743	SH		OTR	1	0	0	743
GREAT SOUTHERN BANCORP INC	COMMON STOCK	390905107	567	11,588	SH		DFND	1	11,588	0	0
GREAT WESTERN BANCORP INC	COMMON STOCK	391416104	68	3,254	SH		DFND	1	3,028	0	226
GREAT WESTERN BANCORP INC	COMMON STOCK	391416104	98	4,706	SH		OTR	1	4,006	0	700
GREEN BRICK PARTNERS INC	COMMON STOCK	392709101	13	583	SH		DFND	1	125	0	458
GREEN PLAINS INC.	COMMON STOCK	393222104	5	362	SH		DFND	1	150	0	212
GREENBRIER COS INC	COMMON STOCK	393657101	34	941	SH		DFND	1	682	0	259
GREENBRIER COS INC	COMMON STOCK	393657101	7	193	SH		OTR	1	0	193	0
GREIF INC CL A	COMMON STOCK	397624107	29	621	SH		DFND	1	133	0	488
GREIF INC CL A	COMMON STOCK	397624107	3	66	SH		OTR	1	0	0	66
GREIF BROS CORP CL B	COMMON STOCK	397624206	73	1,500	SH		DFND	1	0	0	1,500
GREIF BROS CORP CL B	COMMON STOCK	397624206	0	7	SH		OTR	1	0	7	0
GRIFFON CORP	COMMON STOCK	398433102	24	1,201	SH		DFND	1	1,166	0	35
GRIFOLS SA A D R	COMMON STOCK	398438408	278	15,063	SH		DFND	1	11,942	0	3,121
GRIFOLS SA A D R	COMMON STOCK	398438408	118	6,383	SH		OTR	1	4,166	1,291	926
GROUP 1 AUTOMOTIVE INC	COMMON STOCK	398905109	252	1,928	SH		DFND	1	1,669	0	259
GROUPON INC COM	COMMON STOCK	399473206	1	38	SH		DFND	1	0	0	38
GRUBHUB INC	COMMON STOCK	400110102	58	773	SH		DFND	1	578	0	195
GRUBHUB INC	COMMON STOCK	400110102	10	141	SH		OTR	1	0	124	17
GRUBHUB INC	COMMON STOCK	400110102	0	2	SH		DFND	5	2	0	0
GRUPO AEROPORTUARIO DEL A D R	COMMON STOCK	400506101	46	419	SH		DFND	1	241	0	178
GRUPO AEROPORTUARIO DEL A D R	COMMON STOCK	400506101	16	148	SH		OTR	1	148	0	0
GUESS INC	COMMON STOCK	401617105	27	1,184	SH		DFND	1	99	0	1,085
GULF IS FABRICATION INC	COMMON STOCK	402307102	0	95	SH		DFND	1	95	0	0
H E EQUIPMENT SERVICES INC	COMMON STOCK	404030108	6	205	SH		DFND	1	76	0	129
HNI CORP	COMMON STOCK	404251100	437	12,678	SH		DFND	1	12,366	0	312
HNI CORP	COMMON STOCK	404251100	1	28	SH		OTR	1	0	28	0
HSBC HOLDINGS PLC SPONS A D R	COMMON STOCK	404280406	1,065	41,109	SH		DFND	1	28,075	0	13,034
HSBC HOLDINGS PLC SPONS A D R	COMMON STOCK	404280406	191	7,366	SH		OTR	1	6,810	331	225
HACKETT GROUP INC	COMMON STOCK	404609109	6	449	SH		DFND	1	449	0	0
HACKETT GROUP INC	COMMON STOCK	404609109	1	75	SH		OTR	1	75	0	0
HAEMONETICS CORP	COMMON STOCK	405024100	305	2,566	SH		DFND	1	2,504	0	62
HAEMONETICS CORP	COMMON STOCK	405024100	4	37	SH		OTR	1	37	0	0
HAIN CELESTIAL GROUP INC	COMMON STOCK	405217100	195	4,856	SH		DFND	1	2,331	0	2,525
HAIN CELESTIAL GROUP INC	COMMON STOCK	405217100	673	16,774	SH		OTR	1	16,100	107	567
HAIN CELESTIAL GROUP INC	COMMON STOCK	405217100	1	20	SH		DFND	5	20	0	0
HALLIBURTON CO	COMMON STOCK	406216101	1,278	67,594	SH		DFND	1	58,915	0	8,679
HALLIBURTON CO	COMMON STOCK	406216101	297	15,702	SH		OTR	1	14,795	203	704
HALLIBURTON CO	COMMON STOCK	406216101	0	0	SH		DFND	5	0	0	0
HAMILTON LANE INC CLASS A	COMMON STOCK	407497106	12	158	SH		DFND	1	0	0	158
HAMILTON LANE INC CLASS A	COMMON STOCK	407497106	48	616	SH		OTR	1	0	616	0
HANCOCK WHITNEY CORP	COMMON STOCK	410120109	347	10,215	SH		DFND	1	9,875	0	340
HANCOCK WHITNEY CORP	COMMON STOCK	410120109	12	359	SH		OTR	1	251	51	57
HANESBRANDS INC	COMMON STOCK	410345102	295	20,246	SH		DFND	1	15,953	0	4,293
HANESBRANDS INC	COMMON STOCK	410345102	57	3,910	SH		OTR	1	3,775	0	135
HANOVER INS GROUP INC	COMMON STOCK	410867105	301	2,576	SH		DFND	1	882	0	1,694
HANOVER INS GROUP INC	COMMON STOCK	410867105	11	92	SH		OTR	1	92	0	0
HARLEY DAVIDSON INC	COMMON STOCK	412822108	733	19,960	SH		DFND	1	17,629	0	2,331
HARLEY DAVIDSON INC	COMMON STOCK	412822108	118	3,206	SH		OTR	1	2,999	201	6
HARLEY DAVIDSON INCORPORATED	COMMON STOCK	412822108	19	520	SH		DFND	5	520	0	0
HARMONIC INC	COMMON STOCK	413160102	70	9,434	SH		DFND	1	0	0	9,434
HARMONY GOLD MNG LTD A D R	COMMON STOCK	413216300	2	394	SH		DFND	1	0	0	394
HARSCO CORP	COMMON STOCK	415864107	1,534	85,338	SH		DFND	1	79,136	0	6,202
HARSCO CORP	COMMON STOCK	415864107	105	5,858	SH		OTR	1	5,858	0	0
HARTFORD FINANCIAL SERVICES GR	COMMON STOCK	416515104	1,028	20,981	SH		DFND	1	17,507	0	3,474
HARTFORD FINANCIAL SERVICES GR	COMMON STOCK	416515104	634	12,960	SH		OTR	1	12,675	0	285
HARTFORD FINANCIAL SVCS GRP	COMMON STOCK	416515104	2	31	SH		DFND	5	31	0	0
HASBRO INC	COMMON STOCK	418056107	4,711	50,362	SH		DFND	1	48,716	0	1,646
HASBRO INC	COMMON STOCK	418056107	1,363	14,569	SH		OTR	1	13,549	937	83
HASBRO INC	COMMON STOCK	418056107	2	17	SH		DFND	5	17	0	0
HAWAIIAN ELEC INDS INC	COMMON STOCK	419870100	403	11,378	SH		DFND	1	9,354	0	2,024
HAWAIIAN ELEC INDS INC	COMMON STOCK	419870100	5	139	SH		OTR	1	0	11	128
HAWAIIAN HLDGS INC	COMMON STOCK	419879101	18	1,003	SH		DFND	1	885	0	118
HAWKINS INC	COMMON STOCK	420261109	67	1,282	SH		DFND	1	1,269	0	13
HAWKINS INC	COMMON STOCK	420261109	14	270	SH		OTR	1	270	0	0
HAYNES INTERNATIONAL INC	COMMON STOCK	420877201	5	205	SH		OTR	1	205	0	0
HEALTHCARE SVCS GROUP INC	COMMON STOCK	421906108	851	30,265	SH		DFND	1	30,265	0	0
HEALTHCARE SVCS GROUP INC	COMMON STOCK	421906108	175	6,244	SH		OTR	1	4,104	2,140	0
HEALTHCARE SERVICES GROUP	COMMON STOCK	421906108	197	7,004	SH		DFND	5	7,004	0	0
HEALTHCARE RLTY TR	COMMON STOCK	421946104	692	23,363	SH		DFND	1	23,363	0	0
HEALTHCARE RLTY TR	COMMON STOCK	421946104	177	5,993	SH		OTR	1	5,993	0	0
HEALTHCARE REALTY TRUST	COMMON STOCK	421946104	1	35	SH		DFND	5	35	0	0
HEARTLAND EXPRESS INC	COMMON STOCK	422347104	23	1,270	SH		DFND	1	765	0	505
HECLA MNG CO	COMMON STOCK	422704106	12	1,789	SH		DFND	1	1,789	0	0
HEICO CORP	COMMON STOCK	422806109	2,845	21,487	SH		DFND	1	18,909	0	2,578
HEICO CORP	COMMON STOCK	422806109	1,446	10,922	SH		OTR	1	2,671	8,176	75
HEICO CORP	COMMON STOCK	422806109	363	2,741	SH		DFND	5	2,741	0	0
H E I C O CORPORATION CL A	COMMON STOCK	422806208	5,305	45,320	SH		DFND	1	45,122	0	198
H E I C O CORPORATION CL A	COMMON STOCK	422806208	727	6,212	SH		OTR	1	6,212	0	0
HEICO CORP	COMMON STOCK	422806208	465	3,970	SH		DFND	5	3,970	0	0
HEIDRICK & STRUGGLES INTL INC	COMMON STOCK	422819102	4	136	SH		DFND	1	136	0	0
HELMERICH PAYNE INC	COMMON STOCK	423452101	414	17,859	SH		DFND	1	16,505	0	1,354
HELMERICH PAYNE INC	COMMON STOCK	423452101	33	1,421	SH		OTR	1	283	862	276
JACK HENRY ASSOCIATES INC	COMMON STOCK	426281101	13,547	83,626	SH		DFND	1	79,934	0	3,692
JACK HENRY ASSOCIATES INC	COMMON STOCK	426281101	1,857	11,465	SH		OTR	1	6,958	1,483	3,024
HENRY JACK & ASSOCIATES INC	COMMON STOCK	426281101	135	835	SH		DFND	5	835	0	0
HERITAGE COMMERCE CORP	COMMON STOCK	426927109	9	1,054	SH		DFND	1	1,054	0	0
HERCULES CAPITAL INC	COMMON STOCK	427096508	685	47,485	SH		DFND	1	34,684	0	12,801
HERCULES CAPITAL INC	COMMON STOCK	427096508	52	3,575	SH		OTR	1	2,425	0	1,150
HERON THERAPEUTICS INC	COMMON STOCK	427746102	7	350	SH		DFND	1	350	0	0
HERSHA HOSPITALITY TRUST	COMMON STOCK	427825500	17	2,141	SH		DFND	1	1,782	0	359
HERSHEY CO THE	COMMON STOCK	427866108	60,463	396,925	SH		DFND	1	395,437	0	1,488
HERSHEY CO THE	COMMON STOCK	427866108	3,852	25,290	SH		OTR	1	23,010	85	2,195
THE HERSHEY COMPANY	COMMON STOCK	427866108	27	174	SH		DFND	5	174	0	0
HESS MIDSTREAM LP	COMMON STOCK	428103105	1	46	SH		DFND	5	46	0	0
HEXCEL CORP NEW	COMMON STOCK	428291108	1,551	31,987	SH		DFND	1	28,415	0	3,572
HEXCEL CORP NEW	COMMON STOCK	428291108	163	3,350	SH		OTR	1	370	2,902	78
HEXCEL CORP	COMMON STOCK	428291108	37	760	SH		DFND	5	760	0	0
HEXO CORP COM NEW	COMMON STOCK	428304307	0	126	SH		DFND	1	126	0	0
HIGHWOODS PROPERTIES INC	COMMON STOCK	431284108	15	378	SH		DFND	1	243	0	135
HILL ROM HOLDINGS INC	COMMON STOCK	431475102	729	7,436	SH		DFND	1	7,347	0	89
HILL ROM HOLDINGS INC	COMMON STOCK	431475102	314	3,208	SH		OTR	1	2,581	0	627
HILLENBRAND INC	COMMON STOCK	431571108	760	19,079	SH		DFND	1	17,144	0	1,935
HILLENBRAND INC	COMMON STOCK	431571108	10	262	SH		OTR	1	262	0	0
HILLENBRAND INC	COMMON STOCK	431571108	0	6	SH		DFND	5	6	0	0
HILLTOP HOLDINGS INC	COMMON STOCK	432748101	46	1,684	SH		DFND	1	734	0	950
HINGHAM INSTITUTION FOR SAVING	COMMON STOCK	433323102	64	298	SH		DFND	1	295	0	3
HOLLYFRONTIER CORP	COMMON STOCK	436106108	157	6,059	SH		DFND	1	3,548	0	2,511
HOLLYFRONTIER CORP	COMMON STOCK	436106108	101	3,895	SH		OTR	1	536	3,359	0
HOLOGIC INC	COMMON STOCK	436440101	1,423	19,542	SH		DFND	1	17,629	0	1,913
HOLOGIC INC	COMMON STOCK	436440101	447	6,137	SH		OTR	1	2,532	3,563	42
HOME BANCSHARES INC	COMMON STOCK	436893200	37	1,922	SH		DFND	1	1,922	0	0
HOME BANCSHARES INC	COMMON STOCK	436893200	58	2,958	SH		OTR	1	2,827	0	131
HOME BANCSHARES INC	COMMON STOCK	436893200	11	561	SH		DFND	5	561	0	0
HOME DEPOT INC	COMMON STOCK	437076102	231,065	869,910	SH		DFND	1	824,947	0	44,963
HOME DEPOT INC	COMMON STOCK	437076102	76,042	286,282	SH		OTR	1	263,684	11,011	11,587
HOME DEPOT INC	COMMON STOCK	437076102	956	3,591	SH		DFND	5	3,441	0	150
HOME DEPOT INC	COMMON STOCK	437076102	4	4	SH	Put	OTR	1	4	0	0
HOMOLOGY MEDICINES INC	COMMON STOCK	438083107	14	1,215	SH		DFND	1	985	0	230
HONDA MOTOR CO LTD A D R	COMMON STOCK	438128308	1,409	49,850	SH		DFND	1	39,571	0	10,279
HONDA MOTOR CO LTD A D R	COMMON STOCK	438128308	312	11,039	SH		OTR	1	7,660	680	2,699
HONDA MOTOR CO LTD	COMMON STOCK	438128308	1	33	SH		DFND	5	33	0	0
HONEYWELL INTERNATIONAL INC	COMMON STOCK	438516106	114,530	538,460	SH		DFND	1	511,220	0	27,240
HONEYWELL INTERNATIONAL INC	COMMON STOCK	438516106	58,845	276,657	SH		OTR	1	253,342	13,115	10,200
HONEYWELL INTERNATIONAL INC	COMMON STOCK	438516106	245	1,151	SH		DFND	5	1,118	0	33
HONEYWELL INTERNATIONAL INC	COMMON STOCK	438516106	1	19	SH	Put	OTR	1	19	0	0
HOOKER FURNITURE CORP	COMMON STOCK	439038100	32	982	SH		DFND	1	982	0	0
HORACE MANN EDUCATORS CORP	COMMON STOCK	440327104	66	1,561	SH		DFND	1	1,523	0	38
HORIZON BANCORP INC	COMMON STOCK	440407104	171	10,755	SH		DFND	1	10,708	0	47
HORIZON BANCORP INC	COMMON STOCK	440407104	7	450	SH		OTR	1	450	0	0
HORMEL FOODS CORP	COMMON STOCK	440452100	49,029	1,051,895	SH		DFND	1	750,315	0	301,580
HORMEL FOODS CORP	COMMON STOCK	440452100	9,074	194,692	SH		OTR	1	185,161	6,379	3,152
HORMEL FOODS CORPORATION	COMMON STOCK	440452100	151	3,245	SH		DFND	5	3,245	0	0
HOULIHAN LOKEY INC	COMMON STOCK	441593100	1,170	17,398	SH		DFND	1	16,806	0	592
HOULIHAN LOKEY INC	COMMON STOCK	441593100	384	5,706	SH		OTR	1	5,706	0	0
HOULIHAN LOKEY INC	COMMON STOCK	441593100	26	391	SH		DFND	5	391	0	0
HOWMET AEROSPACE INC COM	COMMON STOCK	443201108	416	14,565	SH		DFND	1	13,402	0	1,163
HOWMET AEROSPACE INC COM	COMMON STOCK	443201108	99	3,501	SH		OTR	1	3,401	100	0
HOWMET AEROSPACE INC	COMMON STOCK	443201108	37	1,280	SH		DFND	5	1,280	0	0
HUANENG POWER INTL SPON A D R	COMMON STOCK	443304100	14	978	SH		DFND	1	283	0	695
HUB GROUP INC CL A	COMMON STOCK	443320106	521	9,148	SH		DFND	1	9,046	0	102
HUB GROUP INC CL A	COMMON STOCK	443320106	77	1,346	SH		OTR	1	1,346	0	0
HUBBELL INC	COMMON STOCK	443510607	1,862	11,877	SH		DFND	1	11,505	0	372
HUBBELL INC	COMMON STOCK	443510607	1,948	12,426	SH		OTR	1	9,955	2,397	74
HUBSPOT INC	COMMON STOCK	443573100	424	1,069	SH		DFND	1	861	0	208
HUBSPOT INC	COMMON STOCK	443573100	128	324	SH		OTR	1	300	15	9
HUBSPOT INC	COMMON STOCK	443573100	96	241	SH		DFND	5	241	0	0
HUDSON PACIFIC PROPERTIES INC	COMMON STOCK	444097109	1,113	46,324	SH		DFND	1	46,300	0	24
HUDSON PACIFIC PROPERTIES INC	COMMON STOCK	444097109	17	695	SH		OTR	1	695	0	0
HUDSON PACIFIC PROPERTIES INC	COMMON STOCK	444097109	35	1,442	SH		DFND	5	1,442	0	0
HUMANA INC	COMMON STOCK	444859102	7,766	18,931	SH		DFND	1	17,601	0	1,330
HUMANA INC	COMMON STOCK	444859102	6,698	16,324	SH		OTR	1	11,578	4,382	364
HUMANA INC	COMMON STOCK	444859102	12	30	SH		DFND	5	21	0	9
HUNT J B TRANS SVCS INC	COMMON STOCK	445658107	2,703	19,779	SH		DFND	1	18,750	0	1,029
HUNT J B TRANS SVCS INC	COMMON STOCK	445658107	290	2,124	SH		OTR	1	2,056	68	0
J B HUNT TRANSPORT SERVICES	COMMON STOCK	445658107	30	223	SH		DFND	5	223	0	0
HUNTINGTON BANCSHARES INC	COMMON STOCK	446150104	1,851	146,520	SH		DFND	1	131,627	0	14,893
HUNTINGTON BANCSHARES INC	COMMON STOCK	446150104	281	22,241	SH		OTR	1	20,790	0	1,451
HUNTINGTON BANCSHARES INC	COMMON STOCK	446150104	284	22,494	SH		DFND	5	22,494	0	0
HUNTINGTON INGALLS INDUSTRIE	COMMON STOCK	446413106	1,444	8,470	SH		DFND	1	8,037	0	433
HUNTINGTON INGALLS INDUSTRIE	COMMON STOCK	446413106	548	3,217	SH		OTR	1	3,181	23	13
HUNTINGTON INGALLS INDUSTRIES	COMMON STOCK	446413106	67	392	SH		DFND	5	385	0	7
HUNTSMAN CORP	COMMON STOCK	447011107	68,966	2,743,242	SH		DFND	1	14,430	0	2,728,812
HUNTSMAN CORP	COMMON STOCK	447011107	92	3,663	SH		OTR	1	0	3,418	245
HURON CONSULTING GROUP INC	COMMON STOCK	447462102	133	2,254	SH		DFND	1	0	0	2,254
HYATT HOTELS CORP CL A	COMMON STOCK	448579102	616	8,297	SH		DFND	1	8,047	0	250
HYATT HOTELS CORP CL A	COMMON STOCK	448579102	205	2,763	SH		OTR	1	0	0	2,763
HYSTER YALE MATERIALS W I	COMMON STOCK	449172105	4	62	SH		DFND	1	62	0	0
IAA SPINCO INC	COMMON STOCK	449253103	394	6,068	SH		DFND	1	4,843	0	1,225
IAA SPINCO INC	COMMON STOCK	449253103	149	2,293	SH		OTR	1	1,858	147	288
IGM BIOSCIENCES INC	COMMON STOCK	449585108	273	3,094	SH		DFND	1	3,094	0	0
IGM BIOSCIENCES INC	COMMON STOCK	449585108	3	36	SH		OTR	1	36	0	0
IRHYTHM TECHNOLOGIES INC	COMMON STOCK	450056106	59	246	SH		DFND	1	233	0	13
IRHYTHM TECHNOLOGIES INC	COMMON STOCK	450056106	303	1,279	SH		OTR	1	0	0	1,279
IAMGOLD CORPORATION	COMMON STOCK	450913108	6	1,648	SH		DFND	1	1,360	0	288
IBIO INC	COMMON STOCK	451033203	25	24,100	SH		DFND	1	24,100	0	0
ICAHN ENTERPRISES LP	COMMON STOCK	451100101	82	1,612	SH		DFND	1	1,295	0	317
IDACORP INC	COMMON STOCK	451107106	1,344	13,993	SH		DFND	1	12,221	0	1,772
IDACORP INC	COMMON STOCK	451107106	265	2,761	SH		OTR	1	2,739	11	11
IDACORP INC	COMMON STOCK	451107106	58	605	SH		DFND	5	605	0	0
IDEAL POWER INC	COMMON STOCK	451622203	4	500	SH		DFND	1	500	0	0
ILLINOIS TOOL WORKS INC	COMMON STOCK	452308109	55,004	269,788	SH		DFND	1	251,020	0	18,768
ILLINOIS TOOL WORKS INC	COMMON STOCK	452308109	21,940	107,614	SH		OTR	1	100,387	4,274	2,953
ILLINOIS TOOL WORKS INC	COMMON STOCK	452308109	6	27	SH		DFND	5	27	0	0
ILLINOIS TOOL WORKS INC	COMMON STOCK	452308109	19	120	SH	Put	OTR	1	120	0	0
ILLUMINA INC	COMMON STOCK	452327109	8,036	21,720	SH		DFND	1	20,837	0	883
ILLUMINA INC	COMMON STOCK	452327109	419	1,133	SH		OTR	1	967	117	49
ILLUMINA INC	COMMON STOCK	452327109	17	47	SH		DFND	5	47	0	0
IMEDIA BRANDS INC	COMMON STOCK	452465206	0	70	SH		OTR	1	70	0	0
IMPERIAL OIL LTD	COMMON STOCK	453038408	295	15,539	SH		DFND	1	10,635	0	4,904
IMPERIAL OIL LTD	COMMON STOCK	453038408	88	4,600	SH		OTR	1	4,500	0	100
IMPINJ INC	COMMON STOCK	453204109	70	1,666	SH		DFND	1	1,666	0	0
INDEPENDENCE HOLDINGS CO	COMMON STOCK	453440307	9	222	SH		DFND	1	222	0	0
INDEPENDENT BK CORP MASS	COMMON STOCK	453836108	2,085	28,544	SH		DFND	1	28,513	0	31
INDEPENDENT BK CORP MASS	COMMON STOCK	453836108	59	807	SH		OTR	1	807	0	0
INDUSTRIAL LOGISTICS PROPERT	COMMON STOCK	456237106	281	12,051	SH		DFND	1	368	0	11,683
INDUSTRIAL LOGISTICS PROPERT	COMMON STOCK	456237106	5	220	SH		OTR	1	220	0	0
INDUSTRIAS BACHOCO SAB DE CV A	COMMON STOCK	456463108	2,878	63,724	SH		DFND	1	63,535	0	189
INDUSTRIAS BACHOCO SAB DE CV A	COMMON STOCK	456463108	696	15,412	SH		OTR	1	15,412	0	0
INFOSYS LTD SP A D R	COMMON STOCK	456788108	2,191	129,242	SH		DFND	1	104,667	0	24,575
INFOSYS LTD SP A D R	COMMON STOCK	456788108	618	36,456	SH		OTR	1	29,190	0	7,266
INFOSYS LIMITED	COMMON STOCK	456788108	5	280	SH		DFND	5	280	0	0
I N G GROEP N V SPONSORED A D	COMMON STOCK	456837103	575	60,917	SH		DFND	1	34,556	0	26,361
I N G GROEP N V SPONSORED A D	COMMON STOCK	456837103	115	12,179	SH		OTR	1	9,799	1,824	556
INGLES MARKETS INC CL A	COMMON STOCK	457030104	120	2,811	SH		DFND	1	2,682	0	129
INGREDION INC	COMMON STOCK	457187102	1,381	17,554	SH		DFND	1	15,511	0	2,043
INGREDION INC	COMMON STOCK	457187102	86	1,092	SH		OTR	1	1,092	0	0
INGREDION INC	COMMON STOCK	457187102	38	478	SH		DFND	5	478	0	0
INSMED INC	COMMON STOCK	457669307	15	461	SH		DFND	1	406	0	55
INSPIRE MEDICAL SYSTEMS INC	COMMON STOCK	457730109	404	2,150	SH		DFND	1	2,150	0	0
INSPIRE MEDICAL SYSTEMS INC	COMMON STOCK	457730109	1	6	SH		OTR	1	0	0	6
INTEGRA LIFESCIENCES HOLDINGS	COMMON STOCK	457985208	587	9,050	SH		DFND	1	6,657	0	2,393
INTEGRA LIFESCIENCES HOLDINGS	COMMON STOCK	457985208	117	1,804	SH		OTR	1	1,804	0	0
INTEGRA LIFESCIENCES CORP	COMMON STOCK	457985208	17	266	SH		DFND	5	266	0	0
INTEL CORP	COMMON STOCK	458140100	163,495	3,281,718	SH		DFND	1	3,217,806	0	63,912
INTEL CORP	COMMON STOCK	458140100	74,397	1,493,320	SH		OTR	1	1,414,747	28,377	50,196
INTEL CORP	COMMON STOCK	458140100	57	1,140	SH		DFND	5	679	0	461
INTER PARFUMS INC	COMMON STOCK	458334109	30	492	SH		DFND	1	492	0	0
INTER PARFUMS INC	COMMON STOCK	458334109	132	2,178	SH		OTR	1	58	2,120	0
INTERFACE INC	COMMON STOCK	458665304	94	8,921	SH		DFND	1	8,565	0	356
INTERFACE INC	COMMON STOCK	458665304	1	117	SH		OTR	1	0	117	0
INTERNATIONAL BANCSHARES CORP	COMMON STOCK	459044103	12	319	SH		DFND	1	132	0	187
INTERNATIONAL BANCSHARES CORP	COMMON STOCK	459044103	1	40	SH		OTR	1	40	0	0
INTL BUSINESS MACHINES CORP	COMMON STOCK	459200101	47,430	376,786	SH		DFND	1	367,968	0	8,818
INTL BUSINESS MACHINES CORP	COMMON STOCK	459200101	27,618	219,396	SH		OTR	1	197,514	8,266	13,616
INTL BUSINESS MACHINES CORP	COMMON STOCK	459200101	234	1,861	SH		DFND	5	1,843	0	18
INTL FLAVORS FRAGRANCES	COMMON STOCK	459506101	1,051	9,654	SH		DFND	1	9,020	0	634
INTL FLAVORS FRAGRANCES	COMMON STOCK	459506101	638	5,857	SH		OTR	1	5,835	0	22
INTL FLAVORS & FRAGRANCES	COMMON STOCK	459506101	2	16	SH		DFND	5	16	0	0
INTERNATIONAL PAPER CO	COMMON STOCK	460146103	7,439	149,623	SH		DFND	1	143,972	0	5,651
INTERNATIONAL PAPER CO	COMMON STOCK	460146103	2,292	46,083	SH		OTR	1	41,088	436	4,559
INTERNATIONAL PAPER CO	COMMON STOCK	460146103	8	162	SH		DFND	5	103	0	59
INTERPUBLIC GROUP COS INC	COMMON STOCK	460690100	453	19,227	SH		DFND	1	11,907	0	7,320
INTERPUBLIC GROUP COS INC	COMMON STOCK	460690100	134	5,694	SH		OTR	1	0	5,657	37
INTERPUBLIC GROUP COS	COMMON STOCK	460690100	43	1,822	SH		DFND	5	1,721	0	101
INTUIT INC	COMMON STOCK	461202103	59,291	156,091	SH		DFND	1	151,824	0	4,267
INTUIT INC	COMMON STOCK	461202103	13,294	34,997	SH		OTR	1	29,408	667	4,922
INTUIT	COMMON STOCK	461202103	262	687	SH		DFND	5	630	0	57
IONIS PHARMACEUTICALS INC	COMMON STOCK	462222100	101	1,789	SH		DFND	1	850	0	939
IONIS PHARMACEUTICALS INC	COMMON STOCK	462222100	41	729	SH		OTR	1	600	0	129
IOVANCE BIOTHERAPEUTICS INC	COMMON STOCK	462260100	926	19,948	SH		DFND	1	19,743	0	205
IOVANCE BIOTHERAPEUTICS INC	COMMON STOCK	462260100	38	825	SH		OTR	1	610	185	30
IROBOT CORP	COMMON STOCK	462726100	130	1,626	SH		DFND	1	1,551	0	75
IROBOT CORP	COMMON STOCK	462726100	33	405	SH		OTR	1	405	0	0
ISHARES GOLD TRUST ETF	OTHER	464285105	1,059	58,394	SH		DFND	1	57,356	0	1,038
ISHARES GOLD TRUST ETF	OTHER	464285105	836	46,129	SH		OTR	1	39,649	0	6,480
ISHARES COMEX GOLD TRUST	OTHER	464285105	223	12,306	SH		DFND	5	951	0	11,355
ISHARES J.P. MORGAN EM HIGH ET	OTHER	464286285	9	196	SH		DFND	1	196	0	0
ISHARES EMERGING MARKETS	OTHER	464286285	77	1,651	SH		DFND	5	0	0	1,651
ISHARES EMERGING MARKETS ETF	OTHER	464286319	29	790	SH		DFND	1	790	0	0
ISHARES MSCI GLBL AGRICULTURE	OTHER	464286350	1	31	SH		DFND	1	31	0	0
ISHARES MSCI BRAZIL ETF	OTHER	464286400	1	20	SH		DFND	1	20	0	0
ISHARES MSCI EMERGING MARKETS	OTHER	464286426	9	102	SH		DFND	1	102	0	0
ISHARES MSCI EMERGING MARKETS	OTHER	464286426	67	765	SH		DFND	5	765	0	0
ISHARES MSCI GLOBAL MIN VOL ET	OTHER	464286525	178	1,840	SH		DFND	1	1,840	0	0
ISHARES MSCI ALL COUNTRY WORLD	OTHER	464286525	60	618	SH		DFND	5	570	0	48
ISHARES MSCI EMERGING MARKETS	OTHER	464286533	58	942	SH		DFND	1	942	0	0
ISHARES MSCI EMRG MKT MINIMUM	OTHER	464286533	17	272	SH		DFND	5	0	0	272
ISHARES MSCI EUROZONE ETF	OTHER	464286608	168	3,803	SH		DFND	1	3,803	0	0
ISHARES MSCI EUROZONE ETF	OTHER	464286608	61	1,391	SH		OTR	1	1,391	0	0
ISHARES MSCI THAILAND ETF	OTHER	464286624	35	450	SH		OTR	1	0	0	450
ISHARES MSCI ISRAEL ETF	OTHER	464286632	122	1,902	SH		DFND	1	1,902	0	0
ISHARES MSCI ISRAEL ETF	OTHER	464286632	13	200	SH		OTR	1	200	0	0
I SHARES MSCI PACIFIC EX JAPAN	OTHER	464286665	239	4,989	SH		DFND	1	4,989	0	0
I SHARES MSCI PACIFIC EX JAPAN	OTHER	464286665	431	9,000	SH		OTR	1	9,000	0	0
ISHARES MSCI SWITZERLAND ETF	OTHER	464286749	292	6,539	SH		DFND	1	6,539	0	0
ISHARES MSCI SOUTH KOREA ETF	OTHER	464286772	589	6,844	SH		DFND	1	6,844	0	0
ISHARES MSCI HONG KONG ETF	OTHER	464286871	7	300	SH		DFND	1	300	0	0
ISHARES MSCI HONG KONG ETF	OTHER	464286871	37	1,500	SH		OTR	1	1,500	0	0
ISHARES S P 100 ETF	OTHER	464287101	3,657	21,307	SH		DFND	1	21,307	0	0
ISHARES S P 100 ETF	OTHER	464287101	5,424	31,604	SH		OTR	1	25,444	5,024	1,136
ISHARES MORNINGSTAR LG CAP GRW	OTHER	464287119	559	1,929	SH		DFND	1	1,929	0	0
ISHARES MORNINGSTAR LG CAP GRW	OTHER	464287119	17	60	SH		OTR	1	60	0	0
ISHARES CORE S P TOTL US STK M	OTHER	464287150	1,279	14,839	SH		DFND	1	14,839	0	0
ISHARES CORE S P TOTL US STK M	OTHER	464287150	313	3,635	SH		OTR	1	3,635	0	0
ISHARES S&P 1500 INDEX	OTHER	464287150	37	430	SH		DFND	5	430	0	0
ISHARES SELECT DIVIDEND ETF	OTHER	464287168	117,987	1,226,733	SH		DFND	1	1,200,100	0	26,633
ISHARES SELECT DIVIDEND ETF	OTHER	464287168	24,310	252,754	SH		OTR	1	243,765	1,539	7,450
ISHARES DJ SELECT DIVID INDEX	OTHER	464287168	8	85	SH		DFND	5	0	0	85
ISHARES TIPS BOND ETF	OTHER	464287176	17,414	136,415	SH		DFND	1	135,215	0	1,200
ISHARES TIPS BOND ETF	OTHER	464287176	4,111	32,202	SH		OTR	1	31,074	978	150
ISHARES BARCLAYS US TREASURY	OTHER	464287176	235	1,839	SH		DFND	5	722	0	1,117
ISHARES CHINA LARGE CAP ETF	OTHER	464287184	37	792	SH		DFND	1	792	0	0
ISHARES FTSE CHINA 25 INDEX	OTHER	464287184	8	175	SH		DFND	5	0	0	175
ISHARES TRANSPORTATION AVERAGE	OTHER	464287192	2	10	SH		OTR	1	10	0	0
ISHARES CORE S P 500 ETF	OTHER	464287200	978,334	2,606,179	SH		DFND	1	2,354,413	0	251,766
ISHARES CORE S P 500 ETF	OTHER	464287200	42,076	112,084	SH		OTR	1	101,466	8,200	2,418
ISHARES S&P 500 INDEX	OTHER	464287200	573,556	1,527,818	SH		DFND	5	13,736	0	1,514,082
ISHARES CORE U.S. AGGREGATE BO	OTHER	464287226	912,105	7,717,280	SH		DFND	1	7,120,855	0	596,425
ISHARES CORE U.S. AGGREGATE BO	OTHER	464287226	57,515	486,643	SH		OTR	1	481,035	1,555	4,053
ISHARES CORE US AGGREGATE BOND	OTHER	464287226	335,733	2,840,543	SH		DFND	5	177,298	0	2,663,245
ISHARES MSCI EMERGING MARKETS	OTHER	464287234	184,308	3,567,004	SH		DFND	1	3,530,472	0	36,532
ISHARES MSCI EMERGING MARKETS	OTHER	464287234	51,294	992,728	SH		OTR	1	963,141	22,640	6,947
ISHARES MSCI EMERGING MKTS IDX	OTHER	464287234	7,731	149,628	SH		DFND	5	112,496	0	37,132
ISHARES MSCI EMERGING MARKETS ETF	OTHER	464287234	58	271	SH	Put	DFND	1	271	0	0
ISHARES IBOXX INVST GRADE CRP	OTHER	464287242	45,893	332,244	SH		DFND	1	329,842	0	2,402
ISHARES IBOXX INVST GRADE CRP	OTHER	464287242	5,722	41,423	SH		OTR	1	38,863	1,725	835
ISHARES IBOXX $ INV GRADE CORP	OTHER	464287242	2,135	15,456	SH		DFND	5	0	0	15,456
ISHARES GLOBAL COMM SERVICES E	OTHER	464287275	218	2,941	SH		DFND	1	2,941	0	0
ISHARES GLOBAL COMM SERVICES E	OTHER	464287275	120	1,612	SH		OTR	1	262	0	1,350
ISHARES GLOBAL TECH ETF	OTHER	464287291	682	2,274	SH		DFND	1	2,274	0	0
ISHARES S P 500 GROWTH ETF	OTHER	464287309	105,609	1,654,793	SH		DFND	1	1,306,028	0	348,765
ISHARES S P 500 GROWTH ETF	OTHER	464287309	3,398	53,240	SH		OTR	1	51,864	0	1,376
ISHARES S&P 500 GROWTH INDEX	OTHER	464287309	5,040	78,967	SH		DFND	5	12,471	0	66,496
ISHARES GLOBAL HEALTHCARE ETF	OTHER	464287325	3	43	SH		DFND	1	43	0	0
ISHARES GLOBAL HEALTHCARE ETF	OTHER	464287325	75	980	SH		OTR	1	980	0	0
ISHARES S&P GLOBAL ENERGY ETF	OTHER	464287341	6	300	SH		DFND	1	300	0	0
ISHARES S&P GLOBAL ENERGY ETF	OTHER	464287341	9	450	SH		OTR	1	450	0	0
ISHARES FUNDS S&P GLBL ENERGY	OTHER	464287341	50	2,439	SH		DFND	5	2,374	0	65
ISHARES S P LATIN AMERICA 40 E	OTHER	464287390	37	1,248	SH		DFND	1	1,248	0	0
ISHARES S P 500 VALUE ETF	OTHER	464287408	123,154	961,994	SH		DFND	1	659,331	0	302,663
ISHARES S P 500 VALUE ETF	OTHER	464287408	2,093	16,351	SH		OTR	1	16,351	0	0
ISHARES S&P 500 VALUE INDEX	OTHER	464287408	60	465	SH		DFND	5	6	0	459
ISHARES 20 YEAR TREASURY BOND	OTHER	464287432	39,374	249,625	SH		DFND	1	249,422	0	203
ISHARES 20 YEAR TREASURY BOND	OTHER	464287432	3,412	21,634	SH		OTR	1	21,634	0	0
ISHARES BARCLAYS 20 YRS	OTHER	464287432	162	1,027	SH		DFND	5	250	0	777
ISHARES 7 10 YEAR TREASURY BON	OTHER	464287440	8,664	72,230	SH		DFND	1	71,397	0	833
ISHARES 7 10 YEAR TREASURY BON	OTHER	464287440	1,357	11,310	SH		OTR	1	11,081	0	229
ISHARES BARLAYS 7-10 YRS TREAS	OTHER	464287440	18	151	SH		DFND	5	151	0	0
ISHARES 1 3 YEAR TREASURY BOND	OTHER	464287457	33,762	390,854	SH		DFND	1	389,854	0	1,000
ISHARES 1 3 YEAR TREASURY BOND	OTHER	464287457	414	4,794	SH		OTR	1	4,794	0	0
ISHARES BARCLAYS 1-3 YEARS	OTHER	464287457	249	2,887	SH		DFND	5	9	0	2,878
ISHARES MSCI EAFE ETF	OTHER	464287465	361,174	4,950,303	SH		DFND	1	4,905,860	0	44,443
ISHARES MSCI EAFE ETF	OTHER	464287465	108,270	1,483,951	SH		OTR	1	1,443,678	24,493	15,780
ISHARES MSCI EAFE INDEX	OTHER	464287465	15,619	214,074	SH		DFND	5	173,987	0	40,087
ISHARES MSCI EAFE ETF	OTHER	464287465	32	38	SH	Call	DFND	1	38	0	0
ISHARES MSCI EAFE ETF	OTHER	464287465	129	827	SH	Put	DFND	1	827	0	0
ISHARES RUSSELL MID CAP VALUE	OTHER	464287473	119,625	1,233,757	SH		DFND	1	1,223,687	0	10,070
ISHARES RUSSELL MID CAP VALUE	OTHER	464287473	13,149	135,613	SH		OTR	1	129,192	1,403	5,018
ISHARES RUSSELL MIDCP VAL INDX	OTHER	464287473	70,307	725,044	SH		DFND	5	17,594	0	707,450
ISHARES RUSSELL MID CAP GROWTH	OTHER	464287481	110,968	1,081,031	SH		DFND	1	1,063,387	0	17,644
ISHARES RUSSELL MID CAP GROWTH	OTHER	464287481	26,559	258,730	SH		OTR	1	249,166	428	9,136
ISHARES RUSSELL MIDCAP GROWTH	OTHER	464287481	2,439	23,760	SH		DFND	5	14,038	0	9,722
ISHARES RUSSELL MID CAP ETF	OTHER	464287499	700,498	10,218,792	SH		DFND	1	9,252,993	0	965,799
ISHARES RUSSELL MID CAP ETF	OTHER	464287499	48,110	701,826	SH		OTR	1	680,236	17,700	3,890
ISHARES RUSSELL MIDCAP INDEX	OTHER	464287499	235,185	3,430,780	SH		DFND	5	29,616	0	3,401,164
ISHARES CORE S P MID CAP ETF	OTHER	464287507	83,060	361,397	SH		DFND	1	358,042	0	3,355
ISHARES CORE S P MID CAP ETF	OTHER	464287507	13,570	59,043	SH		OTR	1	47,773	10,650	620
ISHARES S&P MIDCAP 400 INDEX	OTHER	464287507	2,851	12,403	SH		DFND	5	7,735	0	4,668
ISHARES NORTH AMERICAN TECH SF	OTHER	464287515	22	62	SH		DFND	1	62	0	0
ISHARES NORTH AMERICAN TECH SF	OTHER	464287515	57	160	SH		OTR	1	160	0	0
ISHARES S&P NORTH AMERICAN	OTHER	464287515	562	1,587	SH		DFND	5	0	0	1,587
ISHARES PHLX SEMICONDUCTOR ETF	OTHER	464287523	12	31	SH		DFND	1	31	0	0
ISHARES PHLX SEMICONDUCTOR ETF	OTHER	464287523	35	93	SH		OTR	1	93	0	0
ISHARES PHLX SEMICONDUCTOR	OTHER	464287523	396	1,044	SH		DFND	5	0	0	1,044
ISHARES NORTH AMERICAN TECH ET	OTHER	464287549	157	448	SH		DFND	1	448	0	0
ISHARES NORTH AMERICAN TECH ET	OTHER	464287549	236	675	SH		OTR	1	675	0	0
ISHARES S&P NORTH AMERICAN	OTHER	464287549	221	631	SH		DFND	5	0	0	631
ISHARES NASDAQ BIOTECHNOLOGY E	OTHER	464287556	1,519	10,028	SH		DFND	1	9,058	0	970
ISHARES NASDAQ BIOTECHNOLOGY E	OTHER	464287556	1,300	8,582	SH		OTR	1	8,532	50	0
ISHARES NASDAQ BIOTECH	OTHER	464287556	324	2,139	SH		DFND	5	0	0	2,139
ISHARES COHEN STEERS REIT ETF	OTHER	464287564	921	17,094	SH		DFND	1	15,694	0	1,400
ISHARES COHEN & STREERS REALTY	OTHER	464287564	123	2,280	SH		DFND	5	2,252	0	28
ISHARES GLOBAL 100 ETF	OTHER	464287572	423	6,724	SH		DFND	1	6,724	0	0
ISHARES RUSSELL 1000 VALUE ETF	OTHER	464287598	97,550	713,450	SH		DFND	1	667,822	0	45,628
ISHARES RUSSELL 1000 VALUE ETF	OTHER	464287598	12,372	90,487	SH		OTR	1	89,887	500	100
ISHARES RUSSELL 1000 VALUE IDX	OTHER	464287598	2,910	21,280	SH		DFND	5	600	0	20,680
ISHARES S P MID CAP 400 GROWTH	OTHER	464287606	3,135	43,422	SH		DFND	1	43,422	0	0
ISHARES S P MID CAP 400 GROWTH	OTHER	464287606	824	11,410	SH		OTR	1	11,410	0	0
ISHARES S&P MID CAP 400/GROWTH	OTHER	464287606	3,126	43,278	SH		DFND	5	0	0	43,278
ISHARES RUSSELL 1000 GROWTH ET	OTHER	464287614	238,382	988,564	SH		DFND	1	955,502	0	33,062
ISHARES RUSSELL 1000 GROWTH ET	OTHER	464287614	45,534	188,828	SH		OTR	1	178,994	5,456	4,378
ISHARES RUSSELL 1000 GROW INDX	OTHER	464287614	5,142	21,323	SH		DFND	5	271	0	21,052
ISHARES RUSSELL 1000 ETF	OTHER	464287622	110,388	521,092	SH		DFND	1	322,028	0	199,064
ISHARES RUSSELL 1000 ETF	OTHER	464287622	8,287	39,120	SH		OTR	1	38,685	134	301
ISHARES RUSSELL 1000 INDEX	OTHER	464287622	72,412	341,752	SH		DFND	5	3,930	0	337,822
ISHARES RUSSELL 2000 VALUE ETF	OTHER	464287630	58,258	442,183	SH		DFND	1	376,125	0	66,058
ISHARES RUSSELL 2000 VALUE ETF	OTHER	464287630	2,945	22,353	SH		OTR	1	20,734	0	1,619
ISHARES RUSSELL 2000 VAL INDEX	OTHER	464287630	2	15	SH		DFND	5	15	0	0
ISHARES RUSSELL 2000 GROWTH ET	OTHER	464287648	83,571	291,492	SH		DFND	1	258,740	0	32,752
ISHARES RUSSELL 2000 GROWTH ET	OTHER	464287648	9,746	33,996	SH		OTR	1	28,306	0	5,690
ISHARES RUSSELL 2000 GROW INDX	OTHER	464287648	159	556	SH		DFND	5	34	0	522
ISHARES RUSSELL 2000 ETF	OTHER	464287655	253,948	1,295,251	SH		DFND	1	1,185,817	0	109,434
ISHARES RUSSELL 2000 ETF	OTHER	464287655	30,750	156,838	SH		OTR	1	138,564	9,242	9,032
ISHARES RUSSELL 2000 INDEX	OTHER	464287655	107,949	550,591	SH		DFND	5	4,118	0	546,473
ISHARES CORE S P US VALUE ETF	OTHER	464287663	4,525	72,764	SH		DFND	1	72,764	0	0
ISHARES CORE S P US VALUE ETF	OTHER	464287663	3,586	57,668	SH		OTR	1	57,668	0	0
ISHARES CORE S P US GROWTH ETF	OTHER	464287671	6,380	71,931	SH		DFND	1	71,931	0	0
ISHARES CORE S P US GROWTH ETF	OTHER	464287671	1,196	13,480	SH		OTR	1	13,230	250	0
ISHARES RUSSEL 3000 ETF	OTHER	464287689	21,203	94,846	SH		DFND	1	93,130	0	1,716
ISHARES RUSSEL 3000 ETF	OTHER	464287689	4,548	20,343	SH		OTR	1	20,343	0	0
ISHARES RUSSELL 3000 INDEX FD	OTHER	464287689	650	2,909	SH		DFND	5	0	0	2,909
ISHARES US UTILITES ETF	OTHER	464287697	139	1,792	SH		DFND	1	1,792	0	0
ISHARES S P MID CAP 400 VALUE	OTHER	464287705	2,339	27,104	SH		DFND	1	26,954	0	150
ISHARES S P MID CAP 400 VALUE	OTHER	464287705	372	4,307	SH		OTR	1	4,307	0	0
ISHARES S&P MIDCAP 400 VALUE	OTHER	464287705	3,288	38,047	SH		DFND	5	0	0	38,047
ISHARES US TELECOMMUNICATIONS	OTHER	464287713	193	6,379	SH		DFND	1	6,379	0	0
ISHARES US TECHNOLOGY ETF	OTHER	464287721	763	8,964	SH		DFND	1	8,964	0	0
ISHARES US TECHNOLOGY ETF	OTHER	464287721	2,063	24,252	SH		OTR	1	22,572	0	1,680
ISHARES US REAL ESTATE ETF	OTHER	464287739	39,869	465,484	SH		DFND	1	461,102	0	4,382
ISHARES US REAL ESTATE ETF	OTHER	464287739	16,471	192,299	SH		OTR	1	189,470	1,579	1,250
ISHARES DJ US REAL ESTATE INDX	OTHER	464287739	112	1,302	SH		DFND	5	1,056	0	246
ISHARES US INDUSTRIALS ETF	OTHER	464287754	241	2,480	SH		DFND	1	2,480	0	0
ISHARES US HEALTHCARE ETF	OTHER	464287762	570	2,319	SH		DFND	1	2,319	0	0
ISHARES DJ US HLTHCARE SECTOR	OTHER	464287762	878	3,562	SH		DFND	5	0	0	3,562
ISHARES US FINANCIAL SERVICES	OTHER	464287770	32	214	SH		DFND	1	214	0	0
ISHARES US FINANCIAL SERVICES	OTHER	464287770	39	260	SH		OTR	1	260	0	0
ISHARES DJ US FIN SERVICS INDX	OTHER	464287770	196	1,303	SH		DFND	5	0	0	1,303
ISHARES US FINANCIALS ETF	OTHER	464287788	29	440	SH		DFND	1	440	0	0
ISHARES DJ US FIN SECTOR INDX	OTHER	464287788	3	38	SH		DFND	5	0	0	38
ISHARES US ENERGY ETF	OTHER	464287796	8	375	SH		DFND	1	375	0	0
ISHARES CORE S P SMALL CAP ETF	OTHER	464287804	327,940	3,568,451	SH		DFND	1	3,539,287	0	29,164
ISHARES CORE S P SMALL CAP ETF	OTHER	464287804	67,785	737,594	SH		OTR	1	703,290	22,316	11,988
ISHARES S&P SMALLCAP 600 INDEX	OTHER	464287804	2,716	29,556	SH		DFND	5	14,624	0	14,932
ISHARES CORE S P SMALL CAP ETF	OTHER	464287804	22	49	SH	Put	DFND	1	49	0	0
ISHARES US BASIC MATERIALS ETF	OTHER	464287838	86	757	SH		DFND	1	757	0	0
ISHARES DOW JONES US ETF	OTHER	464287846	254	2,694	SH		DFND	1	2,694	0	0
ISHARES DJ TOTAL MARKET INDEX	OTHER	464287846	26	278	SH		DFND	5	278	0	0
ISHARES EUROPE ETF	OTHER	464287861	30	635	SH		DFND	1	635	0	0
ISHARES S P SMALL CAP 600 VALU	OTHER	464287879	3,622	44,549	SH		DFND	1	43,764	0	785
ISHARES S P SMALL CAP 600 VALU	OTHER	464287879	524	6,446	SH		OTR	1	3,726	0	2,720
ISHARES S&P SMALLCAP 600 VALUE	OTHER	464287879	1,513	18,608	SH		DFND	5	0	0	18,608
ISHARES S P SMALL CAP 600 GROW	OTHER	464287887	5,800	50,792	SH		DFND	1	50,792	0	0
ISHARES S P SMALL CAP 600 GROW	OTHER	464287887	492	4,310	SH		OTR	1	4,310	0	0
ISHARES S&P SMALL CAP 600	OTHER	464287887	2,692	23,578	SH		DFND	5	0	0	23,578
ISHARES SHORT TERM NATL MUNI B	OTHER	464288158	85,120	787,350	SH		DFND	1	787,088	0	262
ISHARES SHORT TERM NATL MUNI B	OTHER	464288158	10,032	92,796	SH		OTR	1	84,640	6,389	1,767
ISHARES S&P SHORT TERM NATL	OTHER	464288158	4,746	43,900	SH		DFND	5	43,900	0	0
ISHARES AGENCY BOND ETF	OTHER	464288166	19,441	162,616	SH		DFND	1	162,616	0	0
ISHARES AGENCY BOND ETF	OTHER	464288166	793	6,637	SH		OTR	1	6,637	0	0
ISHARES MORNINGSTAR MID CAP ET	OTHER	464288208	64	275	SH		DFND	1	275	0	0
ISHARES MORNINGSTAR MID CORE	OTHER	464288208	100	427	SH		DFND	5	0	0	427
ISHARES GLOBAL CLEAN ENERGY ET	OTHER	464288224	847	29,997	SH		DFND	1	29,997	0	0
ISHARES GLOBAL CLEAN ENERGY ET	OTHER	464288224	99	3,511	SH		OTR	1	3,511	0	0
ISHARES GLOBAL CLEAN ENRGY	OTHER	464288224	327	11,596	SH		DFND	5	0	0	11,596
ISHARES MSCI ACWI EX US ETF	OTHER	464288240	17,677	333,284	SH		DFND	1	332,619	0	665
ISHARES MSCI ACWI EX US ETF	OTHER	464288240	3,156	59,494	SH		OTR	1	55,094	4,400	0
ISHARES MSCI ACWI ETF	OTHER	464288257	16,111	177,588	SH		DFND	1	563	0	177,025
ISHARES MSCI EAFE SMALL CAP ET	OTHER	464288273	9,631	140,902	SH		DFND	1	140,902	0	0
ISHARES MSCI EAFE SMALL CAP ET	OTHER	464288273	4,565	66,781	SH		OTR	1	65,976	0	805
ISHARES MSCI EAFE SMALL CAP	OTHER	464288273	2,604	38,101	SH		DFND	5	0	0	38,101
ISHARES JPMORGAN USD EMERGING	OTHER	464288281	15,523	133,919	SH		DFND	1	133,328	0	591
ISHARES JPMORGAN USD EMERGING	OTHER	464288281	3,401	29,341	SH		OTR	1	29,028	0	313
ISHARES JP MORGAN USD EMERGING	OTHER	464288281	356	3,072	SH		DFND	5	2,232	0	840
ISHARES MORNINGSTAR MID CAP ET	OTHER	464288307	36	95	SH		DFND	1	95	0	0
ISHARES MORNINGSTAR MID GROWTH	OTHER	464288307	750	1,960	SH		DFND	5	0	0	1,960
ISHARES S&P CA AMT FREE BD FD	OTHER	464288356	126	2,012	SH		DFND	5	2,012	0	0
ISHARES GLOBAL INFRASTRUCTURE	OTHER	464288372	8	175	SH		DFND	1	175	0	0
ISHARES S&P GLOBAL INFRASTR	OTHER	464288372	27	628	SH		DFND	5	612	0	16
ISHARES MORNINGSTAR MID CAP VA	OTHER	464288406	84	538	SH		DFND	1	538	0	0
ISHARES NATIONAL MUNI BOND ETF	OTHER	464288414	300,535	2,564,292	SH		DFND	1	2,555,982	0	8,310
ISHARES NATIONAL MUNI BOND ETF	OTHER	464288414	33,358	284,628	SH		OTR	1	258,722	22,871	3,035
ISHARES S&P NATL AMT FREE MUNI	OTHER	464288414	8,346	71,209	SH		DFND	5	70,204	0	1,005
ISHARES INTERNATIONAL DEVELOPE	OTHER	464288422	8	222	SH		OTR	1	222	0	0
ISHARES ASIA 50 ETF	OTHER	464288430	20	235	SH		DFND	1	235	0	0
ISHARES INTERNATIONAL SELECT E	OTHER	464288448	40,590	1,377,318	SH		DFND	1	1,370,659	0	6,659
ISHARES INTERNATIONAL SELECT E	OTHER	464288448	8,741	296,603	SH		OTR	1	293,878	1,325	1,400
ISHARES DJ INTL SELECT DIV	OTHER	464288448	24	818	SH		DFND	5	0	0	818
ISHARES IBOXX HIGH YIELD ETF	OTHER	464288513	55,909	640,428	SH		DFND	1	637,298	0	3,130
ISHARES IBOXX HIGH YIELD ETF	OTHER	464288513	14,744	168,890	SH		OTR	1	167,376	304	1,210
ISHARES IBOXX HI YLD CORP BOND	OTHER	464288513	4,792	54,893	SH		DFND	5	46,016	0	8,877
ISHARES CORE US REIT ETF	OTHER	464288521	13,137	271,372	SH		DFND	1	269,178	0	2,194
ISHARES CORE US REIT ETF	OTHER	464288521	2,410	49,787	SH		OTR	1	49,787	0	0
ISHARS FTSE NAREIT REAL ESTATE	OTHER	464288521	3	54	SH		DFND	5	54	0	0
ISHARES RESIDENTIAL AND ETF	OTHER	464288562	12	180	SH		DFND	1	180	0	0
ISHARES MSCI KLD 400 SOCIAL ET	OTHER	464288570	14,046	196,340	SH		DFND	1	196,340	0	0
ISHARES MSCI KLD 400 SOCIAL ET	OTHER	464288570	388	5,416	SH		OTR	1	5,416	0	0
ISHARES MSCI KLD 400 SOCIAL	OTHER	464288570	31	435	SH		DFND	5	0	0	435
ISHARES MBS ETF	OTHER	464288588	33,354	302,859	SH		DFND	1	299,450	0	3,409
ISHARES MBS ETF	OTHER	464288588	565	5,131	SH		OTR	1	5,131	0	0
ISHARES BARCLAYS MBS BOND FUND	OTHER	464288588	102	923	SH		DFND	5	923	0	0
ISHARES GOVERNMENT CREDIT BOND	OTHER	464288596	2,423	19,243	SH		DFND	1	19,243	0	0
ISHARES GOVERNMENT CREDIT BOND	OTHER	464288596	224	1,778	SH		OTR	1	1,460	318	0
ISHARES MORNINGSTAR SMALL	OTHER	464288604	65	213	SH		DFND	5	213	0	0
ISHARES INTERMEDIATE GOVT CRDT	OTHER	464288612	74,028	629,274	SH		DFND	1	629,189	0	85
ISHARES INTERMEDIATE GOVT CRDT	OTHER	464288612	5,218	44,352	SH		OTR	1	44,352	0	0
ISHARES BARCLAYS INTERMEDIATE	OTHER	464288612	6,757	57,439	SH		DFND	5	3,940	0	53,499
ISHARES BROAD USD INVESTMENT E	OTHER	464288620	2,366	38,181	SH		DFND	1	6,356	0	31,825
ISHARES BROAD USD INVESTMENT E	OTHER	464288620	378	6,100	SH		OTR	1	6,100	0	0
ISHARES BARCLAYS CREDIT BOND	OTHER	464288620	90	1,451	SH		DFND	5	0	0	1,451
ISHARES TRUST ISHARES 5 10 YEA	OTHER	464288638	37,742	610,816	SH		DFND	1	596,778	0	14,038
ISHARES TRUST ISHARES 5 10 YEA	OTHER	464288638	3,608	58,384	SH		OTR	1	57,234	1,150	0
ISHARES BARCLAYS INTERMEDIATE	OTHER	464288638	83	1,343	SH		DFND	5	1,276	0	67
ISHARES TRUST ISHARES 1 5 YEAR	OTHER	464288646	87,601	1,587,830	SH		DFND	1	1,567,550	0	20,280
ISHARES TRUST ISHARES 1 5 YEAR	OTHER	464288646	5,818	105,450	SH		OTR	1	105,450	0	0
ISHARES TR 1-5 YR INVT GRADE	OTHER	464288646	173	3,133	SH		DFND	5	1,741	0	1,392
ISHARES LEHMAN 10-20 YR T BOND	OTHER	464288653	231	1,449	SH		DFND	5	259	0	1,190
ISHARES 3 7 YEAR TREASURY BOND	OTHER	464288661	26,343	198,103	SH		DFND	1	197,849	0	254
ISHARES 3 7 YEAR TREASURY BOND	OTHER	464288661	261	1,966	SH		OTR	1	1,452	514	0
ISHARES SHORT TREASURY BOND ET	OTHER	464288679	11,807	106,824	SH		DFND	1	106,824	0	0
ISHARES SHORT TREASURY BOND ET	OTHER	464288679	1,933	17,492	SH		OTR	1	17,492	0	0
ISHARES SHORT TREASURY BOND	OTHER	464288679	378	3,420	SH		DFND	5	3,399	0	21
ISHARES TRUST ISHARES PREFERRE	OTHER	464288687	59,308	1,540,071	SH		DFND	1	1,530,048	0	10,023
ISHARES TRUST ISHARES PREFERRE	OTHER	464288687	15,505	402,624	SH		OTR	1	356,530	2,863	43,231
ISHARES S&P US PFD STK INDEX	OTHER	464288687	135	3,502	SH		DFND	5	3,192	0	310
ISHARES MORNINGSTAR SMALL CAP	OTHER	464288703	17	123	SH		DFND	1	123	0	0
ISHARES GLOBAL CONSUMER STAPLE	OTHER	464288737	4	76	SH		DFND	1	76	0	0
ISHARES GLOBAL CONSUMER ETF	OTHER	464288745	6	40	SH		DFND	1	40	0	0
ISHARES HOME CONSTRUCTION ETF	OTHER	464288752	81	1,450	SH		DFND	1	1,450	0	0
ISHARES HOME CONSTRUCTION ETF	OTHER	464288752	19	346	SH		OTR	1	346	0	0
ISHARES US AEROSPACE DEFENSE E	OTHER	464288760	229	2,418	SH		DFND	1	2,418	0	0
ISHARES US REGIONAL BANKS ETF	OTHER	464288778	7,050	155,491	SH		DFND	1	155,056	0	435
ISHARES US REGIONAL BANKS ETF	OTHER	464288778	309	6,818	SH		OTR	1	6,818	0	0
ISHARES US INSURANCE ETF	OTHER	464288786	122	1,800	SH		DFND	1	1,800	0	0
ISHARES MSCI USA ESG SELECT ET	OTHER	464288802	943	11,448	SH		DFND	1	11,448	0	0
ISHARES MSCI USA ESG SELECT ET	OTHER	464288802	58	700	SH		OTR	1	700	0	0
ISHARES MSCI USA ESG SELECT	OTHER	464288802	21	252	SH		DFND	5	0	0	252
ISHARES US MEDICAL DEVICES ETF	OTHER	464288810	164	500	SH		DFND	1	500	0	0
ISHARES US MEDICAL DEVICES ETF	OTHER	464288810	38	115	SH		OTR	1	115	0	0
ISHARES DJ US MED EQUIPMT INDX	OTHER	464288810	3,808	11,635	SH		DFND	5	6,802	0	4,833
ISHARES U.S. HEALTHCARE ETF	OTHER	464288828	5	20	SH		DFND	1	20	0	0
ISHARES US PHARMACEUTICALS ETF	OTHER	464288836	19	104	SH		DFND	1	50	0	54
ISHARES US OIL GAS EXPLORATION	OTHER	464288851	1	20	SH		DFND	1	20	0	0
ISHARES MICRO CAP ETF	OTHER	464288869	19	162	SH		DFND	1	162	0	0
ISHARES MSCI EAFE VALUE ETF	OTHER	464288877	95,506	2,023,433	SH		DFND	1	2,002,888	0	20,545
ISHARES MSCI EAFE VALUE ETF	OTHER	464288877	16,523	350,058	SH		OTR	1	345,894	3,181	983
ISHARES MSCI EAFE VALUE	OTHER	464288877	71,583	1,516,517	SH		DFND	5	9,004	0	1,507,513
ISHARES MSCI EAFE GROWTH ETF	OTHER	464288885	93,541	926,972	SH		DFND	1	903,386	0	23,586
ISHARES MSCI EAFE GROWTH ETF	OTHER	464288885	22,113	219,140	SH		OTR	1	214,629	903	3,608
ISHARES MSCI EAFE GROWTH	OTHER	464288885	6,255	61,985	SH		DFND	5	73	0	61,912
ISHARES MSIC NEW ZEALAND ETF	OTHER	464289123	169	2,477	SH		DFND	1	2,477	0	0
ISHARES MSCI EUROPE FINANCIALS	OTHER	464289180	25	1,400	SH		DFND	1	1,400	0	0
ISHARES RUSSEL TOP 200 GROWTH	OTHER	464289438	3,963	29,787	SH		DFND	5	0	0	29,787
ISHARES RUSSELL TOP 200 INDEX	OTHER	464289446	2	23	SH		DFND	5	23	0	0
ISHARES 10 YEAR INVESTMENT GRA	OTHER	464289511	2,650	36,293	SH		DFND	1	36,293	0	0
ISHARES 10 YEAR INVESTMENT GRA	OTHER	464289511	249	3,414	SH		OTR	1	3,414	0	0
ISHARES 10 YR CREDIT BOND	OTHER	464289511	65	893	SH		DFND	5	893	0	0
ISHARES S&P INDIA NIFTY 50	OTHER	464289529	14	332	SH		DFND	5	0	0	332
ISHARES CORE AGGRESSIVE ETF	OTHER	464289859	126	1,967	SH		DFND	1	1,967	0	0
ISHARES S & P AGGRESSIVE ALLOC	OTHER	464289859	29	459	SH		DFND	5	0	0	459
ITAU UNIBANCO HOLDINGS SA A D	COMMON STOCK	465562106	4,771	783,351	SH		DFND	1	780,782	0	2,569
ITAU UNIBANCO HOLDINGS SA A D	COMMON STOCK	465562106	900	147,805	SH		OTR	1	142,292	5,513	0
ITAU UNIBANCO HOLDING SA	COMMON STOCK	465562106	23	3,802	SH		DFND	5	3,802	0	0
ITRON INC	COMMON STOCK	465741106	26	266	SH		DFND	1	185	0	81
J & J SNACK FOODS CORP	COMMON STOCK	466032109	786	5,057	SH		DFND	1	3,944	0	1,113
J & J SNACK FOODS CORP	COMMON STOCK	466032109	89	572	SH		OTR	1	572	0	0
J & J SNACK FOODS CORP	COMMON STOCK	466032109	13	81	SH		DFND	5	81	0	0
JABIL INC	COMMON STOCK	466313103	427	10,045	SH		DFND	1	7,507	0	2,538
JABIL CIRCUIT INC	COMMON STOCK	466313103	0	0	SH		DFND	5	0	0	0
JACK IN THE BOX INC	COMMON STOCK	466367109	667	7,187	SH		DFND	1	6,935	0	252
JACK IN THE BOX INC	COMMON STOCK	466367109	134	1,440	SH		OTR	1	1,402	0	38
JACOBS ENGR GROUP INC	COMMON STOCK	469814107	4,991	45,803	SH		DFND	1	43,950	0	1,853
JACOBS ENGR GROUP INC	COMMON STOCK	469814107	1,182	10,851	SH		OTR	1	10,788	0	63
JACOBS ENGINEERING GROUP INC	COMMON STOCK	469814107	6	51	SH		DFND	5	51	0	0
JETBLUE AIRWAYS CORP	COMMON STOCK	477143101	340	23,349	SH		DFND	1	22,750	0	599
JETBLUE AIRWAYS CORP	COMMON STOCK	477143101	1	50	SH		OTR	1	50	0	0
JOHN BEAN TECHNOLOGIES CORP	COMMON STOCK	477839104	336	2,953	SH		DFND	1	2,838	0	115
JOHN BEAN TECHNOLOGIES CORP	COMMON STOCK	477839104	262	2,308	SH		OTR	1	165	618	1,525
JOHNSON JOHNSON	COMMON STOCK	478160104	353,037	2,243,215	SH		DFND	1	2,158,244	0	84,971
JOHNSON JOHNSON	COMMON STOCK	478160104	176,286	1,120,132	SH		OTR	1	1,053,717	19,722	46,693
JOHNSON & JOHNSON	COMMON STOCK	478160104	369	2,345	SH		DFND	5	2,292	0	53
JOHNSON JOHNSON	COMMON STOCK	478160104	0	9	SH	Put	DFND	1	9	0	0
JOHNSON JOHNSON	COMMON STOCK	478160104	0	12	SH	Put	OTR	1	0	12	0
JOHNSON OUTDOORS INC CL A	COMMON STOCK	479167108	379	3,361	SH		DFND	1	3,361	0	0
JOHNSON OUTDOORS INC CL A	COMMON STOCK	479167108	23	207	SH		OTR	1	207	0	0
KLA CORPORATION	COMMON STOCK	482480100	12,316	47,568	SH		DFND	1	45,043	0	2,525
KLA CORPORATION	COMMON STOCK	482480100	8,248	31,858	SH		OTR	1	31,610	142	106
KLA-TENCOR CORP	COMMON STOCK	482480100	126	487	SH		DFND	5	487	0	0
KAISER ALUMINUM CORP	COMMON STOCK	483007704	145	1,473	SH		DFND	1	1,473	0	0
KAISER ALUMINUM CORP	COMMON STOCK	483007704	112	1,136	SH		OTR	1	1,136	0	0
KAISER ALUMINUM CORP	COMMON STOCK	483007704	13	135	SH		DFND	5	135	0	0
KALA PHARMACEUTICALS INC	COMMON STOCK	483119103	1	160	SH		DFND	1	160	0	0
KALEIDO BIOSCIENCES INC	COMMON STOCK	483347100	18	2,000	SH		DFND	1	2,000	0	0
KAMAN CORP	COMMON STOCK	483548103	45	789	SH		DFND	1	324	0	465
KAMAN CORP	COMMON STOCK	483548103	87	1,523	SH		OTR	1	0	1,523	0
KANSAS CITY SOUTHERN	COMMON STOCK	485170302	5,569	27,281	SH		DFND	1	24,709	0	2,572
KANSAS CITY SOUTHERN	COMMON STOCK	485170302	1,378	6,752	SH		OTR	1	6,497	177	78
KANSAS CITY SOUTHERN	COMMON STOCK	485170302	1	5	SH		DFND	5	5	0	0
KAYNE ANDERSON ENERGY INFRASTR	OTHER	486606106	130	22,671	SH		DFND	1	11,671	0	11,000
KAYNE ANDERSON ENERGY INFRASTR	OTHER	486606106	79	13,816	SH		OTR	1	13,816	0	0
KELLOGG CO	COMMON STOCK	487836108	3,690	59,292	SH		DFND	1	56,658	0	2,634
KELLOGG CO	COMMON STOCK	487836108	968	15,556	SH		OTR	1	13,845	79	1,632
KELLOGG CO	COMMON STOCK	487836108	29	466	SH		DFND	5	466	0	0
KELLY SVCS INC CL A	COMMON STOCK	488152208	147	7,136	SH		DFND	1	920	0	6,216
KEMPER CORP	COMMON STOCK	488401100	692	9,005	SH		DFND	1	7,733	0	1,272
KEMPER CORP	COMMON STOCK	488401100	261	3,399	SH		OTR	1	1,959	0	1,440
KEMPER CORP	COMMON STOCK	488401100	39	513	SH		DFND	5	513	0	0
KENNAMETAL INC	COMMON STOCK	489170100	1,615	44,556	SH		DFND	1	43,938	0	618
KENNAMETAL INC	COMMON STOCK	489170100	39	1,071	SH		OTR	1	1,071	0	0
KENNAMETAL INC	COMMON STOCK	489170100	0	0	SH		DFND	5	0	0	0
KENNEDY WILSON HOLDINGS INC	COMMON STOCK	489398107	33	1,852	SH		DFND	1	1,598	0	254
KENNEDY WILSON HOLDINGS INC	COMMON STOCK	489398107	8	467	SH		DFND	5	467	0	0
KEWAUNEE SCIENTIFIC CORP	COMMON STOCK	492854104	7	534	SH		DFND	1	34	0	500
KEYCORP	COMMON STOCK	493267108	3,228	196,677	SH		DFND	1	179,316	0	17,361
KEYCORP	COMMON STOCK	493267108	1,598	97,366	SH		OTR	1	93,476	594	3,296
KEYCORP	COMMON STOCK	493267108	21	1,250	SH		DFND	5	1,250	0	0
K FORCE INC	COMMON STOCK	493732101	347	8,247	SH		DFND	1	7,641	0	606
K FORCE INC	COMMON STOCK	493732101	57	1,358	SH		OTR	1	1,358	0	0
KFORCE INC	COMMON STOCK	493732101	11	258	SH		DFND	5	258	0	0
KIMBALL INTERNATIONAL B	COMMON STOCK	494274103	101	8,482	SH		DFND	1	8,482	0	0
KIMBERLY CLARK CORP	COMMON STOCK	494368103	32,314	239,666	SH		DFND	1	231,517	0	8,149
KIMBERLY CLARK CORP	COMMON STOCK	494368103	15,790	117,114	SH		OTR	1	113,794	1,273	2,047
KIMBERLY-CLARK CORP	COMMON STOCK	494368103	311	2,303	SH		DFND	5	2,253	0	50
KIMBERLY CLARK CORP	COMMON STOCK	494368103	4	5	SH	Put	OTR	1	5	0	0
KINROSS GOLD CORP	COMMON STOCK	496902404	217	29,435	SH		DFND	1	18,557	0	10,878
KINROSS GOLD CORP	COMMON STOCK	496902404	107	14,522	SH		OTR	1	2,021	8,971	3,530
KINROSS GOLD CORP	COMMON STOCK	496902404	8	1,034	SH		DFND	5	1,034	0	0
KIRBY CORP	COMMON STOCK	497266106	62	1,189	SH		DFND	1	431	0	758
KIRBY CORP	COMMON STOCK	497266106	17	325	SH		OTR	1	300	0	25
KIRKLANDS INC	COMMON STOCK	497498105	4,470	250,000	SH		DFND	1	250,000	0	0
KNOLL INC	COMMON STOCK	498904200	227	15,476	SH		DFND	1	15,361	0	115
KNOLL INC	COMMON STOCK	498904200	88	5,967	SH		OTR	1	5,899	68	0
KNIGHT SWIFT TRANSPORTATION	COMMON STOCK	499049104	609	14,560	SH		DFND	1	4,405	0	10,155
KNIGHT SWIFT TRANSPORTATION	COMMON STOCK	499049104	10	240	SH		OTR	1	79	21	140
KOHLS CORP	COMMON STOCK	500255104	742	18,233	SH		DFND	1	15,951	0	2,282
KOHLS CORP	COMMON STOCK	500255104	79	1,926	SH		OTR	1	1,400	438	88
KONINKLIJKE PHILIPS NVR NY A D	COMMON STOCK	500472303	14,489	267,460	SH		DFND	1	263,223	0	4,237
KONINKLIJKE PHILIPS NVR NY A D	COMMON STOCK	500472303	3,819	70,491	SH		OTR	1	70,102	287	102
KONINKLIJKE PHILIPS NV	COMMON STOCK	500472303	53	969	SH		DFND	5	969	0	0
KOPIN CORP	COMMON STOCK	500600101	2	1,000	SH		OTR	1	1,000	0	0
KOREA ELEC POWER SPON A D R	COMMON STOCK	500631106	351	28,609	SH		DFND	1	14,392	0	14,217
KOREA ELEC POWER SPON A D R	COMMON STOCK	500631106	3	220	SH		OTR	1	0	94	126
KORN FERRY INTL	COMMON STOCK	500643200	732	16,815	SH		DFND	1	14,356	0	2,459
KORN FERRY INTL	COMMON STOCK	500643200	145	3,328	SH		OTR	1	3,328	0	0
KORN/FERRY INTERNATIONAL	COMMON STOCK	500643200	6	149	SH		DFND	5	149	0	0
THE KRAFT HEINZ CO	COMMON STOCK	500754106	2,175	62,762	SH		DFND	1	55,297	0	7,465
THE KRAFT HEINZ CO	COMMON STOCK	500754106	2,109	60,855	SH		OTR	1	58,944	1,576	335
KRAFT HEINZ CO (THE)	COMMON STOCK	500754106	2	48	SH		DFND	5	48	0	0
KRANESHARES CSI CHINA INTERNET	OTHER	500767306	23	300	SH		DFND	1	300	0	0
KROGER CO	COMMON STOCK	501044101	26,823	844,534	SH		DFND	1	638,120	0	206,414
KROGER CO	COMMON STOCK	501044101	1,688	53,152	SH		OTR	1	46,156	500	6,496
KROGER CO	COMMON STOCK	501044101	45	1,410	SH		DFND	5	1,347	0	63
KRYSTAL BIOTECH INC	COMMON STOCK	501147102	12	205	SH		DFND	1	182	0	23
KULICKE & SOFFA INDS INC	COMMON STOCK	501242101	17	526	SH		DFND	1	246	0	280
KULICKE & SOFFA INDS INC	COMMON STOCK	501242101	147	4,635	SH		OTR	1	2,900	1,735	0
L BRANDS INC	COMMON STOCK	501797104	2,382	64,050	SH		DFND	1	62,213	0	1,837
L BRANDS INC	COMMON STOCK	501797104	12	332	SH		OTR	1	108	224	0
L BRANDS INC	COMMON STOCK	501797104	32	845	SH		DFND	5	845	0	0
LKQ CORPORATION	COMMON STOCK	501889208	1,870	53,070	SH		DFND	1	51,626	0	1,444
LKQ CORPORATION	COMMON STOCK	501889208	152	4,313	SH		OTR	1	2,218	2,095	0
LKQ CORP	COMMON STOCK	501889208	58	1,641	SH		DFND	5	1,641	0	0
L T C PPTYS INC	COMMON STOCK	502175102	306	7,854	SH		DFND	1	7,854	0	0
L T C PPTYS INC	COMMON STOCK	502175102	331	8,514	SH		OTR	1	8,514	0	0
L3HARRIS TECHNOLOGIES INC	COMMON STOCK	502431109	8,196	43,357	SH		DFND	1	39,662	0	3,695
L3HARRIS TECHNOLOGIES INC	COMMON STOCK	502431109	1,563	8,271	SH		OTR	1	7,445	102	724
L3HARRIS TECHNOLOGIES INC	COMMON STOCK	502431109	136	715	SH		DFND	5	656	0	59
LA Z BOY INC	COMMON STOCK	505336107	513	12,884	SH		DFND	1	12,865	0	19
LA Z BOY INC	COMMON STOCK	505336107	121	3,039	SH		OTR	1	2,145	894	0
LA-Z BOY INCORPORATED	COMMON STOCK	505336107	12	313	SH		DFND	5	313	0	0
LADDER CAPITAL CORP REIT CL A	COMMON STOCK	505743104	9	872	SH		DFND	1	812	0	60
LAKELAND FINANCIAL CORP	COMMON STOCK	511656100	134	2,495	SH		DFND	1	2,219	0	276
LAKELAND FINANCIAL CORP	COMMON STOCK	511656100	9	172	SH		DFND	5	172	0	0
LAKELAND INDUSTRIES INC	COMMON STOCK	511795106	55	2,009	SH		DFND	1	2,009	0	0
LAM RESEARCH CORP	COMMON STOCK	512807108	9,132	19,337	SH		DFND	1	17,990	0	1,347
LAM RESEARCH CORP	COMMON STOCK	512807108	12,274	25,989	SH		OTR	1	22,821	2,841	327
LAM RESEARCH CORPORATION	COMMON STOCK	512807108	244	515	SH		DFND	5	507	0	8
LAMAR ADVERTISING CO A	COMMON STOCK	512816109	653	7,846	SH		DFND	1	7,433	0	413
LAMAR ADVERTISING CO A	COMMON STOCK	512816109	140	1,689	SH		OTR	1	1,624	0	65
LAMB WESTON HOLDINGS INC	COMMON STOCK	513272104	7,254	92,117	SH		DFND	1	90,861	0	1,256
LAMB WESTON HOLDINGS INC	COMMON STOCK	513272104	1,370	17,410	SH		OTR	1	16,593	449	368
LAMB WESTON HOLDING INC	COMMON STOCK	513272104	57	728	SH		DFND	5	706	0	22
LANCASTER COLONY CORP	COMMON STOCK	513847103	1,396	7,596	SH		DFND	1	7,486	0	110
LANCASTER COLONY CORP	COMMON STOCK	513847103	137	749	SH		OTR	1	517	224	8
LANCASTER COLONY CORP	COMMON STOCK	513847103	49	265	SH		DFND	5	265	0	0
LANDSTAR SYS INC	COMMON STOCK	515098101	403	2,994	SH		DFND	1	2,855	0	139
LANDSTAR SYS INC	COMMON STOCK	515098101	358	2,657	SH		OTR	1	0	41	2,616
LANDSTAR SYSTEM INC	COMMON STOCK	515098101	9	69	SH		DFND	5	69	0	0
LANTRONIX INC	COMMON STOCK	516548203	2	500	SH		DFND	1	500	0	0
LAREDO PETE HLDGS INC	COMMON STOCK	516806205	0	10	SH		DFND	1	10	0	0
LARIMAR THERAPEUTICS INC	COMMON STOCK	517125100	25	1,175	SH		OTR	1	1,175	0	0
LAS VEGAS SANDS CORP	COMMON STOCK	517834107	2,409	40,405	SH		DFND	1	38,669	0	1,736
LAS VEGAS SANDS CORP	COMMON STOCK	517834107	314	5,274	SH		OTR	1	4,770	504	0
LAS VEGAS SANDS CORP	COMMON STOCK	517834107	23	383	SH		DFND	5	383	0	0
LATTICE SEMICONDUCTOR CORP	COMMON STOCK	518415104	9	186	SH		DFND	1	0	0	186
LATTICE SEMICONDUCTOR CORP	COMMON STOCK	518415104	40	876	SH		OTR	1	0	876	0
HARTFORD MULTIFACTOR DEVELOPED	OTHER	518416102	9	303	SH		DFND	1	303	0	0
HARTFORD MULTIFACTOR DEV MKTS	OTHER	518416102	2	62	SH		DFND	5	0	0	62
LAUDER ESTEE COS INC CL A	COMMON STOCK	518439104	36,860	138,474	SH		DFND	1	134,321	0	4,153
LAUDER ESTEE COS INC CL A	COMMON STOCK	518439104	7,762	29,161	SH		OTR	1	24,031	949	4,181
ESTEE LAUDER COMPANIES INC	COMMON STOCK	518439104	168	627	SH		DFND	5	584	0	43
LEAR CORP	COMMON STOCK	521865204	1,585	9,970	SH		DFND	1	9,736	0	234
LEAR CORP	COMMON STOCK	521865204	291	1,828	SH		OTR	1	1,808	20	0
LEAR CORP	COMMON STOCK	521865204	15	95	SH		DFND	5	95	0	0
LEGGETT PLATT INC	COMMON STOCK	524660107	85,375	1,927,199	SH		DFND	1	1,923,764	0	3,435
LEGGETT PLATT INC	COMMON STOCK	524660107	1,456	32,871	SH		OTR	1	32,546	0	325
LEIDOS HOLDINGS INC	COMMON STOCK	525327102	759	7,217	SH		DFND	1	6,287	0	930
LEIDOS HOLDINGS INC	COMMON STOCK	525327102	64	606	SH		OTR	1	301	158	147
LEIDOS HLDGS INC	COMMON STOCK	525327102	30	285	SH		DFND	5	285	0	0
LEMAITRE VASCULAR	COMMON STOCK	525558201	159	3,926	SH		DFND	1	1,599	0	2,327
LEMAITRE VASCULAR	COMMON STOCK	525558201	133	3,275	SH		OTR	1	215	0	3,060
LEMAITRE VASCULAR INC	COMMON STOCK	525558201	13	310	SH		DFND	5	310	0	0
LENNAR CORP CL A	COMMON STOCK	526057104	547	7,173	SH		DFND	1	5,213	0	1,960
LENNAR CORP CL A	COMMON STOCK	526057104	103	1,348	SH		OTR	1	973	216	159
LENNAR CORP	COMMON STOCK	526057104	27	349	SH		DFND	5	349	0	0
LENNAR CORP CL B	COMMON STOCK	526057302	21	347	SH		DFND	1	347	0	0
LENNAR CORP CL B	COMMON STOCK	526057302	0	7	SH		OTR	1	2	4	1
LENNOX INTERNATIONAL INC	COMMON STOCK	526107107	2,845	10,384	SH		DFND	1	9,754	0	630
LENNOX INTERNATIONAL INC	COMMON STOCK	526107107	459	1,675	SH		OTR	1	1,347	328	0
LENNOX INTERNATIONAL INC	COMMON STOCK	526107107	132	482	SH		DFND	5	482	0	0
LEXINFINTECH HLDGS LTD A D R	COMMON STOCK	528877103	47	7,030	SH		DFND	1	6,754	0	276
LEXINGTON REALTY TRUST	COMMON STOCK	529043101	350	33,014	SH		DFND	1	26,805	0	6,209
LEXINGTON REALTY TRUST	COMMON STOCK	529043101	56	5,278	SH		OTR	1	609	44	4,625
LIBERTY BROADBAND A	COMMON STOCK	530307107	119	755	SH		DFND	1	507	0	248
LIBERTY BROADBAND A	COMMON STOCK	530307107	462	2,934	SH		OTR	1	0	0	2,934
LIBERTY BROADBAND C	COMMON STOCK	530307305	374	2,358	SH		DFND	1	2,108	0	250
LIBERTY BROADBAND C	COMMON STOCK	530307305	76	480	SH		OTR	1	388	35	57
LIBERTY MEDIA CORP DELAWARE	COMMON STOCK	531229409	99	2,291	SH		DFND	1	1,008	0	1,283
LIBERTY MEDIA CORP DELAWARE	COMMON STOCK	531229607	56	1,290	SH		DFND	1	1,290	0	0
LIBERTY MEDIA CORP DELAWARE	COMMON STOCK	531229607	467	10,737	SH		OTR	1	0	0	10,737
LIBERTY FORMULA ONE COMMON STO	COMMON STOCK	531229854	7	166	SH		DFND	1	166	0	0
LIBERTY FORMULA ONE COMMON STO	COMMON STOCK	531229854	579	13,597	SH		OTR	1	0	0	13,597
LIBERTY FORMULA ONE COMMON STO	COMMON STOCK	531229870	27	714	SH		DFND	1	270	0	444
LIBERTY FORMULA ONE COMMON STO	COMMON STOCK	531229870	5	128	SH		OTR	1	76	0	52
LIBERTY MEDIA CORP DELAWARE	COMMON STOCK	531229888	26	1,026	SH		DFND	1	1,026	0	0
LIBERTY MEDIA CORP DELAWARE	COMMON STOCK	531229888	16	653	SH		OTR	1	0	653	0
ELI LILLY CO	COMMON STOCK	532457108	351,115	2,079,575	SH		DFND	1	2,062,232	0	17,343
ELI LILLY CO	COMMON STOCK	532457108	14,047	83,195	SH		OTR	1	69,899	2,356	10,940
ELI LILLY & CO	COMMON STOCK	532457108	104	617	SH		DFND	5	587	0	30
ELI LILLY CO	COMMON STOCK	532457108	5	8	SH	Put	DFND	1	8	0	0
LINCOLN ELEC HLDGS INC	COMMON STOCK	533900106	691	5,942	SH		DFND	1	5,774	0	168
LINCOLN ELEC HLDGS INC	COMMON STOCK	533900106	740	6,365	SH		OTR	1	4,764	1,601	0
LINCOLN NATIONAL CORP	COMMON STOCK	534187109	3,135	62,322	SH		DFND	1	57,843	0	4,479
LINCOLN NATIONAL CORP	COMMON STOCK	534187109	843	16,758	SH		OTR	1	16,128	236	394
LINDBLAD EXPEDITIONS HOLDING	COMMON STOCK	535219109	4	231	SH		DFND	1	0	0	231
LINDSAY CORPORATION	COMMON STOCK	535555106	13	100	SH		DFND	1	0	0	100
LINDSAY CORP	COMMON STOCK	535555106	0	0	SH		DFND	5	0	0	0
LIONS GATE ENTERTAINMENT A	COMMON STOCK	535919401	28	2,460	SH		DFND	1	2,424	0	36
LIONS GATE ENTERTAINMENT B	COMMON STOCK	535919500	0	2	SH		DFND	1	2	0	0
LITHIA MOTORS INC CL A	COMMON STOCK	536797103	595	2,032	SH		DFND	1	1,786	0	246
LITHIA MOTORS INC CL A	COMMON STOCK	536797103	303	1,034	SH		OTR	1	500	212	322
LITHIA MOTORS INC	COMMON STOCK	536797103	82	279	SH		DFND	5	279	0	0
LITTELFUSE INC	COMMON STOCK	537008104	1,277	5,012	SH		DFND	1	4,222	0	790
LITTELFUSE INC	COMMON STOCK	537008104	732	2,874	SH		OTR	1	1,794	980	100
LITTELFUSE INC	COMMON STOCK	537008104	27	106	SH		DFND	5	106	0	0
LIVE NATION ENTERTAINMENT INC	COMMON STOCK	538034109	1,275	17,345	SH		DFND	1	16,716	0	629
LIVE NATION ENTERTAINMENT INC	COMMON STOCK	538034109	179	2,435	SH		OTR	1	331	39	2,065
LIVE NATION ENTERTAINMENT	COMMON STOCK	538034109	1	13	SH		DFND	5	13	0	0
LIVE PERSON INC	COMMON STOCK	538146101	1,760	28,285	SH		DFND	1	24,766	0	3,519
LIVE PERSON INC	COMMON STOCK	538146101	41	655	SH		OTR	1	655	0	0
LIVEPERSON INC	COMMON STOCK	538146101	0	1	SH		DFND	5	1	0	0
LLOYDS BANKING GROUP PLC A D	COMMON STOCK	539439109	206	105,210	SH		DFND	1	99,427	0	5,783
LLOYDS BANKING GROUP PLC A D	COMMON STOCK	539439109	171	87,021	SH		OTR	1	64,985	22,036	0
LOCKHEED MARTIN CORP	COMMON STOCK	539830109	41,211	116,095	SH		DFND	1	111,122	0	4,973
LOCKHEED MARTIN CORP	COMMON STOCK	539830109	14,104	39,731	SH		OTR	1	37,218	997	1,516
LOCKHEED MARTIN CORP	COMMON STOCK	539830109	294	826	SH		DFND	5	826	0	0
LOEWS CORP	COMMON STOCK	540424108	640	14,212	SH		DFND	1	13,533	0	679
LOEWS CORP	COMMON STOCK	540424108	37	832	SH		OTR	1	832	0	0
LOEWS CORP	COMMON STOCK	540424108	31	695	SH		DFND	5	695	0	0
LOUISIANA PAC CORP	COMMON STOCK	546347105	1,935	52,053	SH		DFND	1	52,053	0	0
LOUISIANA PAC CORP	COMMON STOCK	546347105	132	3,559	SH		OTR	1	2,559	1,000	0
LOWES CO INC	COMMON STOCK	548661107	46,765	291,355	SH		DFND	1	273,204	0	18,151
LOWES CO INC	COMMON STOCK	548661107	66,873	416,625	SH		OTR	1	397,211	17,111	2,303
LOWES COMPANIES INC	COMMON STOCK	548661107	707	4,404	SH		DFND	5	4,365	0	39
LOWES CO INC	COMMON STOCK	548661107	24	68	SH	Put	DFND	1	68	0	0
LOWES CO INC	COMMON STOCK	548661107	41	91	SH	Put	OTR	1	0	91	0
LUBYS INC	COMMON STOCK	549282101	29	10,074	SH		DFND	1	10,074	0	0
LULULEMON ATHLETICA INC	COMMON STOCK	550021109	9,616	27,630	SH		DFND	1	26,364	0	1,266
LULULEMON ATHLETICA INC	COMMON STOCK	550021109	612	1,759	SH		OTR	1	1,502	115	142
LULULEMON ATHLETICA INC	COMMON STOCK	550021109	795	2,283	SH		DFND	5	2,259	0	24
LUNA INNOVATIONS INC	COMMON STOCK	550351100	35	3,500	SH		DFND	1	3,500	0	0
LUMINAR TECHNOLOGIES INC	COMMON STOCK	550424105	23	665	SH		DFND	1	665	0	0
LUMINAR TECHNOLOGIES INC	COMMON STOCK	550424105	34	1,000	SH		OTR	1	0	1,000	0
LYDALL INC	COMMON STOCK	550819106	1,128	37,550	SH		DFND	1	37,550	0	0
M D C HLDGS INC	COMMON STOCK	552676108	287	5,908	SH		DFND	1	962	0	4,946
M D U RES GROUP INC	COMMON STOCK	552690109	50,065	1,900,706	SH		DFND	1	1,875,696	0	25,010
M D U RES GROUP INC	COMMON STOCK	552690109	444	16,872	SH		OTR	1	14,460	601	1,811
M G I C INVT CORP	COMMON STOCK	552848103	302	24,030	SH		DFND	1	7,111	0	16,919
M G I C INVT CORP	COMMON STOCK	552848103	39	3,132	SH		OTR	1	3,132	0	0
MGM RESORTS INTERNATIONAL	COMMON STOCK	552953101	955	30,303	SH		DFND	1	26,717	0	3,586
MGM RESORTS INTERNATIONAL	COMMON STOCK	552953101	15	480	SH		OTR	1	0	115	365
MGM RESORTS INTL	COMMON STOCK	552953101	3	93	SH		DFND	5	93	0	0
MP MATLS CORP	COMMON STOCK	553368101	25	770	SH		DFND	1	770	0	0
MSA SAFETY INC	COMMON STOCK	553498106	698	4,671	SH		DFND	1	2,986	0	1,685
MSA SAFETY INC	COMMON STOCK	553498106	28	187	SH		OTR	1	187	0	0
MSC INDL DIRECT CO INC CL A	COMMON STOCK	553530106	900	10,666	SH		DFND	1	9,171	0	1,495
MSC INDL DIRECT CO INC CL A	COMMON STOCK	553530106	549	6,512	SH		OTR	1	251	991	5,270
MSC INDUSTRIAL DIRECT CO INC	COMMON STOCK	553530106	14	162	SH		DFND	5	162	0	0
MSG NETWORK INC	COMMON STOCK	553573106	14	976	SH		DFND	1	976	0	0
M T S SYS CORP	COMMON STOCK	553777103	455	7,833	SH		DFND	1	7,833	0	0
M T S SYS CORP	COMMON STOCK	553777103	162	2,791	SH		OTR	1	2,791	0	0
MTS SYSTEMS CORP	COMMON STOCK	553777103	8	143	SH		DFND	5	143	0	0
MV OIL TRUST	COMMON STOCK	553859109	1	180	SH		DFND	1	180	0	0
MACERICH CO	COMMON STOCK	554382101	106	9,942	SH		DFND	1	6,409	0	3,533
MACERICH CO	COMMON STOCK	554382101	2	174	SH		OTR	1	0	0	174
MACROGENICS INC	COMMON STOCK	556099109	5	235	SH		DFND	1	0	0	235
MADDEN STEVEN LTD	COMMON STOCK	556269108	142	4,024	SH		DFND	1	1,441	0	2,583
MADRIGAL PHARMACEUTICALS INC	COMMON STOCK	558868105	4	34	SH		DFND	1	26	0	8
MAGELLAN HEALTH, INC	COMMON STOCK	559079207	5	56	SH		DFND	1	26	0	30
MAGELLAN MIDSTREAM PARTNERS LP	COMMON STOCK	559080106	3,035	71,514	SH		DFND	1	63,614	0	7,900
MAGELLAN MIDSTREAM PARTNERS LP	COMMON STOCK	559080106	706	16,644	SH		OTR	1	6,147	10,167	330
MAGELLAN MIDSTREAM PARTNERS	COMMON STOCK	559080106	27	638	SH		DFND	5	638	0	0
MAGNA INTL INC CL A	COMMON STOCK	559222401	17,341	244,931	SH		DFND	1	240,813	0	4,118
MAGNA INTL INC CL A	COMMON STOCK	559222401	3,590	50,704	SH		OTR	1	47,864	1,400	1,440
MAGNOLIA OIL GAS CORP	COMMON STOCK	559663109	26	3,617	SH		DFND	1	766	0	2,851
MANHATTAN ASSOCS INC	COMMON STOCK	562750109	920	8,748	SH		DFND	1	4,513	0	4,235
MANHATTAN ASSOCS INC	COMMON STOCK	562750109	61	580	SH		OTR	1	90	468	22
MANITOWOC INC	COMMON STOCK	563571405	37	2,813	SH		DFND	1	2,806	0	7
MANITOWOC INC	COMMON STOCK	563571405	0	4	SH		OTR	1	0	0	4
MANTECH INTL CORP A	COMMON STOCK	564563104	495	5,564	SH		DFND	1	4,429	0	1,135
MANTECH INTL CORP A	COMMON STOCK	564563104	31	343	SH		OTR	1	343	0	0
MANTECH INTERNATIONAL CORP	COMMON STOCK	564563104	13	141	SH		DFND	5	141	0	0
MARATHON OIL CORPORATION	COMMON STOCK	565849106	336	50,387	SH		DFND	1	29,091	0	21,296
MARATHON OIL CORPORATION	COMMON STOCK	565849106	110	16,512	SH		OTR	1	15,309	0	1,203
MARCUS MILLICHAP INC	COMMON STOCK	566324109	226	6,067	SH		DFND	1	6,067	0	0
MARCUS MILLICHAP INC	COMMON STOCK	566324109	64	1,711	SH		OTR	1	1,711	0	0
MARCUS & MILLICAP CO INC	COMMON STOCK	566324109	11	287	SH		DFND	5	287	0	0
MARCUS CORP	COMMON STOCK	566330106	14	1,050	SH		DFND	1	1,050	0	0
MARINEMAX INC	COMMON STOCK	567908108	11	304	SH		DFND	1	69	0	235
MARINE PRODUCTS CORP	COMMON STOCK	568427108	1	44	SH		DFND	1	0	0	44
MARKEL CORP	COMMON STOCK	570535104	2,085	2,018	SH		DFND	1	2,010	0	8
MARKEL CORP	COMMON STOCK	570535104	1,959	1,896	SH		OTR	1	1,785	111	0
MARSH MCLENNAN COS INC	COMMON STOCK	571748102	15,723	134,385	SH		DFND	1	115,935	0	18,450
MARSH MCLENNAN COS INC	COMMON STOCK	571748102	4,589	39,222	SH		OTR	1	27,283	5,297	6,642
MARSH & MCLENNAN COMPANIES INC	COMMON STOCK	571748102	112	954	SH		DFND	5	954	0	0
MARRIOTT INTL INC	COMMON STOCK	571903202	7,195	54,544	SH		DFND	1	52,652	0	1,892
MARRIOTT INTL INC	COMMON STOCK	571903202	2,287	17,337	SH		OTR	1	6,843	306	10,188
MARTEN TRANSPORTATION LTD	COMMON STOCK	573075108	1,370	79,495	SH		DFND	1	70,903	0	8,592
MARTEN TRANSPORTATION LTD	COMMON STOCK	573075108	35	2,029	SH		OTR	1	2,029	0	0
MARTIN MARIETTA MATLS INC	COMMON STOCK	573284106	2,180	7,677	SH		DFND	1	4,329	0	3,348
MARTIN MARIETTA MATLS INC	COMMON STOCK	573284106	935	3,293	SH		OTR	1	3,089	19	185
MARTIN MARIETTA MATERIALS	COMMON STOCK	573284106	1	3	SH		DFND	5	3	0	0
MASCO CORP	COMMON STOCK	574599106	6,292	114,545	SH		DFND	1	110,062	0	4,483
MASCO CORP	COMMON STOCK	574599106	1,549	28,194	SH		OTR	1	27,655	150	389
MASCO CORP	COMMON STOCK	574599106	83	1,510	SH		DFND	5	1,462	0	48
MASIMO CORP	COMMON STOCK	574795100	916	3,414	SH		DFND	1	1,949	0	1,465
MASIMO CORP	COMMON STOCK	574795100	126	472	SH		OTR	1	345	115	12
MASONITE INTERNATIONAL CORP	COMMON STOCK	575385109	678	6,905	SH		DFND	1	5,206	0	1,699
MASONITE INTERNATIONAL CORP	COMMON STOCK	575385109	8	81	SH		OTR	1	81	0	0
MASONITE INTL CORP	COMMON STOCK	575385109	1	15	SH		DFND	5	15	0	0
MASTEC INC	COMMON STOCK	576323109	1,042	15,282	SH		DFND	1	8,698	0	6,584
MASTEC INC	COMMON STOCK	576323109	1,903	27,905	SH		OTR	1	27,905	0	0
MATADOR RESOURCES CO	COMMON STOCK	576485205	265	21,994	SH		DFND	1	20,683	0	1,311
MATADOR RESOURCES CO	COMMON STOCK	576485205	2	171	SH		OTR	1	171	0	0
MATERION CORP	COMMON STOCK	576690101	6	96	SH		DFND	1	96	0	0
MATERION CORP	COMMON STOCK	576690101	4	61	SH		DFND	5	61	0	0
MATTEL INC	COMMON STOCK	577081102	47	2,707	SH		DFND	1	2,433	0	274
MATTEL INC	COMMON STOCK	577081102	14	794	SH		OTR	1	97	697	0
MATTHEWS INTL CORP CL A	COMMON STOCK	577128101	40	1,374	SH		DFND	1	1,374	0	0
MATTHEWS INTERNATIONAL CORP	COMMON STOCK	577128101	0	8	SH		DFND	5	8	0	0
MAXIMUS INC	COMMON STOCK	577933104	1,045	14,291	SH		DFND	1	13,921	0	370
MAXIMUS INC	COMMON STOCK	577933104	70	961	SH		OTR	1	874	18	69
MAXIMUS INC	COMMON STOCK	577933104	10	141	SH		DFND	5	141	0	0
MCAFEE CORP	COMMON STOCK	579063108	9	530	SH		DFND	1	530	0	0
MCCORMICK CO NON VTG SHRS	COMMON STOCK	579780206	10,444	109,255	SH		DFND	1	101,158	0	8,097
MCCORMICK CO NON VTG SHRS	COMMON STOCK	579780206	3,888	40,662	SH		OTR	1	26,904	142	13,616
MCCORMICK & CO INC	COMMON STOCK	579780206	24	251	SH		DFND	5	229	0	22
MCDONALDS CORP	COMMON STOCK	580135101	164,583	767,000	SH		DFND	1	743,222	0	23,778
MCDONALDS CORP	COMMON STOCK	580135101	71,172	331,681	SH		OTR	1	302,254	12,986	16,441
MCDONALDS CORP	COMMON STOCK	580135101	76	353	SH		DFND	5	353	0	0
MCGRATH RENTCORP	COMMON STOCK	580589109	538	8,012	SH		DFND	1	7,995	0	17
MCGRATH RENTCORP	COMMON STOCK	580589109	104	1,557	SH		OTR	1	1,557	0	0
MELCO RESORTS AND ENT LIMITED	COMMON STOCK	585464100	221	11,874	SH		DFND	1	9,235	0	2,639
MELCO RESORTS AND ENT LIMITED	COMMON STOCK	585464100	26	1,426	SH		OTR	1	1,235	138	53
MERCANTILE BANK CORP	COMMON STOCK	587376104	150	5,518	SH		DFND	1	5,518	0	0
MERCANTILE BANK CORP	COMMON STOCK	587376104	70	2,566	SH		OTR	1	2,566	0	0
MERCANTILE BANK CORP	COMMON STOCK	587376104	3	127	SH		DFND	5	127	0	0
MERCURY SYSTEMS INC	COMMON STOCK	589378108	694	7,889	SH		DFND	1	7,872	0	17
MERCURY SYSTEMS INC	COMMON STOCK	589378108	511	5,803	SH		OTR	1	2,808	0	2,995
MERCURY COMPUTER SYSTEM	COMMON STOCK	589378108	33	378	SH		DFND	5	378	0	0
MERCURY GEN CORP	COMMON STOCK	589400100	106	2,039	SH		DFND	1	1,158	0	881
MERCURY GEN CORP	COMMON STOCK	589400100	104	1,993	SH		OTR	1	0	1,969	24
MEREDITH CORP	COMMON STOCK	589433101	56	2,938	SH		DFND	1	2,719	0	219
MEREDITH CORP	COMMON STOCK	589433101	64	3,343	SH		OTR	1	3,343	0	0
MERIDIAN BIOSCIENCE INC	COMMON STOCK	589584101	519	27,774	SH		DFND	1	27,774	0	0
MERIDIAN BIOSCIENCE INC	COMMON STOCK	589584101	7	361	SH		OTR	1	361	0	0
MERIT MED SYS INC	COMMON STOCK	589889104	522	9,395	SH		DFND	1	9,210	0	185
MERIT MED SYS INC	COMMON STOCK	589889104	193	3,473	SH		OTR	1	283	0	3,190
MESA ROYALTY TRUST	COMMON STOCK	590660106	0	100	SH		DFND	1	100	0	0
MESOBLAST LTD SPON A D R	COMMON STOCK	590717104	7	776	SH		DFND	1	303	0	473
MESOBLAST LTD SPON A D R	COMMON STOCK	590717104	9	1,100	SH		OTR	1	0	1,100	0
METHODE ELECTRONICS INC	COMMON STOCK	591520200	96	2,513	SH		DFND	1	2,500	0	13
METTLER TOLEDO INTL INC	COMMON STOCK	592688105	9,481	8,319	SH		DFND	1	8,038	0	281
METTLER TOLEDO INTL INC	COMMON STOCK	592688105	1,559	1,368	SH		OTR	1	1,344	19	5
METTLER TOLEDO INTERNATIONAL	COMMON STOCK	592688105	800	702	SH		DFND	5	700	0	2
MEXICO FD INC	OTHER	592835102	2	178	SH		OTR	1	178	0	0
MICRO FOCUS INTL SPN A D R	COMMON STOCK	594837403	780	136,547	SH		DFND	1	136,512	0	35
MICRO FOCUS INTL SPN A D R	COMMON STOCK	594837403	229	40,067	SH		OTR	1	40,067	0	0
MICRO FOCUS INTL	COMMON STOCK	594837403	2	369	SH		DFND	5	369	0	0
MICROSOFT CORP	COMMON STOCK	594918104	1,024,584	4,606,528	SH		DFND	1	4,417,914	0	188,614
MICROSOFT CORP	COMMON STOCK	594918104	314,549	1,414,208	SH		OTR	1	1,269,836	47,839	96,533
MICROSOFT CORP	COMMON STOCK	594918104	3,110	13,965	SH		DFND	5	13,220	0	745
MICROSOFT CORP	COMMON STOCK	594918104	12	20	SH	Call	DFND	1	20	0	0
MICROSOFT CORP	COMMON STOCK	594918104	22	53	SH	Put	DFND	1	53	0	0
MICROSOFT CORP	COMMON STOCK	594918104	19	36	SH	Put	OTR	1	0	36	0
MICROVISION INC	COMMON STOCK	594960304	1	200	SH		DFND	1	200	0	0
MICROSTRATEGY INC CL A	COMMON STOCK	594972408	565	1,455	SH		DFND	1	1,440	0	15
MICROSTRATEGY INC CL A	COMMON STOCK	594972408	7	17	SH		OTR	1	17	0	0
MICROCHIP TECHNOLOGY INC	COMMON STOCK	595017104	26,728	193,526	SH		DFND	1	187,840	0	5,686
MICROCHIP TECHNOLOGY INC	COMMON STOCK	595017104	9,070	65,672	SH		OTR	1	58,853	2,277	4,542
MICROCHIP TECHNOLOGY INC	COMMON STOCK	595017104	92	663	SH		DFND	5	663	0	0
MICRON TECHNOLOGY INC	COMMON STOCK	595112103	71,042	944,953	SH		DFND	1	925,247	0	19,706
MICRON TECHNOLOGY INC	COMMON STOCK	595112103	7,739	102,941	SH		OTR	1	96,009	2,002	4,930
MICRON TECHNOLOGY INC	COMMON STOCK	595112103	57	764	SH		DFND	5	764	0	0
MIDDLEBY CORP	COMMON STOCK	596278101	1,090	8,455	SH		DFND	1	8,267	0	188
MIDDLEBY CORP	COMMON STOCK	596278101	477	3,704	SH		OTR	1	3,647	21	36
MIDDLEBY CORP	COMMON STOCK	596278101	105	818	SH		DFND	5	818	0	0
MIDDLESEX WATER CO	COMMON STOCK	596680108	12	166	SH		DFND	1	144	0	22
MIDDLESEX WATER CO	COMMON STOCK	596680108	2	28	SH		OTR	1	0	28	0
MIDLAND STATES BANCORP INC	COMMON STOCK	597742105	43	2,416	SH		DFND	1	1,496	0	920
MIDWESTONE FINL GROUP INC	COMMON STOCK	598511103	18	724	SH		DFND	1	724	0	0
MILLER HERMAN INC	COMMON STOCK	600544100	28	814	SH		DFND	1	481	0	333
MILLER INDUSTRIES INC	COMMON STOCK	600551204	478	12,578	SH		DFND	1	11,464	0	1,114
MIMEDX GROUP INC	COMMON STOCK	602496101	4	451	SH		DFND	1	200	0	251
MINERALS TECHNOLOGIES INC	COMMON STOCK	603158106	566	9,116	SH		DFND	1	8,743	0	373
MINERALS TECHNOLOGIES INC	COMMON STOCK	603158106	103	1,664	SH		OTR	1	1,664	0	0
MITEK SYS INC	COMMON STOCK	606710200	231	13,019	SH		DFND	1	12,233	0	786
MITSUBISHI UFJ FINL GRP A D R	COMMON STOCK	606822104	868	195,985	SH		DFND	1	127,233	0	68,752
MITSUBISHI UFJ FINL GRP A D R	COMMON STOCK	606822104	172	39,063	SH		OTR	1	17,745	5,686	15,632
MITSUBISHI UFJ FINANCIAL	COMMON STOCK	606822104	3	780	SH		DFND	5	780	0	0
MOBILE TELESYSTEMS A D R	COMMON STOCK	607409109	14	1,612	SH		DFND	1	1,612	0	0
MOBILE TELESYSTEMS A D R	COMMON STOCK	607409109	1	140	SH		OTR	1	140	0	0
MODEL N INC	COMMON STOCK	607525102	14	394	SH		DFND	1	394	0	0
MODEL N INC	COMMON STOCK	607525102	199	5,565	SH		OTR	1	280	0	5,285
MOHAWK INDS INC	COMMON STOCK	608190104	1,734	12,306	SH		DFND	1	11,929	0	377
MOHAWK INDS INC	COMMON STOCK	608190104	412	2,923	SH		OTR	1	1,570	0	1,353
MOHAWK INDUSTRIES INC	COMMON STOCK	608190104	31	219	SH		DFND	5	219	0	0
MONARCH CASINO & RESORT INC	COMMON STOCK	609027107	185	3,014	SH		DFND	1	2,980	0	34
MONARCH CASINO & RESORT INC	COMMON STOCK	609027107	38	611	SH		OTR	1	95	516	0
MONDELEZ INTERNATIONAL W I	COMMON STOCK	609207105	30,227	516,975	SH		DFND	1	493,335	0	23,640
MONDELEZ INTERNATIONAL W I	COMMON STOCK	609207105	14,647	250,509	SH		OTR	1	223,971	17,523	9,015
MONDELEZ INTERNATIONAL INC	COMMON STOCK	609207105	130	2,212	SH		DFND	5	2,056	0	156
MONMOUTH REAL ESTATE INVT CORP	COMMON STOCK	609720107	35	2,049	SH		DFND	1	2,015	0	34
MONMOUTH REAL ESTATE INVT CORP	COMMON STOCK	609720107	1	62	SH		OTR	1	0	62	0
MONOLITHIC POWER SYSTEMS INC	COMMON STOCK	609839105	7,539	20,587	SH		DFND	1	18,714	0	1,873
MONOLITHIC POWER SYSTEMS INC	COMMON STOCK	609839105	934	2,550	SH		OTR	1	2,483	67	0
MONOLITHIC POWER SYSTEMS INC	COMMON STOCK	609839105	94	257	SH		DFND	5	244	0	13
MONRO INC	COMMON STOCK	610236101	35	656	SH		DFND	1	656	0	0
MONRO INC	COMMON STOCK	610236101	1	21	SH		OTR	1	21	0	0
MONRO MUFFLER BRAKE INC	COMMON STOCK	610236101	7	139	SH		DFND	5	139	0	0
MONTROSE ENVIRONMENTAL GROUP I	COMMON STOCK	615111101	1	33	SH		DFND	1	0	0	33
MOODYS CORP	COMMON STOCK	615369105	2,709	9,331	SH		DFND	1	7,904	0	1,427
MOODYS CORP	COMMON STOCK	615369105	1,845	6,358	SH		OTR	1	5,297	544	517
MOODYS CORPORATION	COMMON STOCK	615369105	41	139	SH		DFND	5	139	0	0
MOOG INC CLASS A	COMMON STOCK	615394202	447	5,633	SH		DFND	1	5,526	0	107
MOOG INC CLASS A	COMMON STOCK	615394202	81	1,026	SH		OTR	1	1,026	0	0
MORGAN STANLEY	COMMON STOCK	617446448	11,914	173,853	SH		DFND	1	159,163	0	14,690
MORGAN STANLEY	COMMON STOCK	617446448	4,395	64,120	SH		OTR	1	59,887	3,041	1,192
MORGAN STANLEY	COMMON STOCK	617446448	103	1,502	SH		DFND	5	1,458	0	44
MORNINGSTAR INC	COMMON STOCK	617700109	3,005	12,975	SH		DFND	1	12,778	0	197
MORNINGSTAR INC	COMMON STOCK	617700109	477	2,059	SH		OTR	1	1,607	434	18
MORNINGSTAR INC	COMMON STOCK	617700109	15	63	SH		DFND	5	63	0	0
MORPHOSYS AG A D R	COMMON STOCK	617760202	2	88	SH		DFND	1	0	0	88
MORPHOSYS AG A D R	COMMON STOCK	617760202	13	468	SH		OTR	1	0	468	0
MOTOROLA SOLUTIONS INC	COMMON STOCK	620076307	29,799	175,225	SH		DFND	1	172,002	0	3,223
MOTOROLA SOLUTIONS INC	COMMON STOCK	620076307	3,563	20,955	SH		OTR	1	20,095	197	663
MOTOROLA SOLUTIONS INC	COMMON STOCK	620076307	15	87	SH		DFND	5	49	0	38
MOVADO GROUP INC	COMMON STOCK	624580106	98	5,870	SH		DFND	1	5,870	0	0
MOVADO GROUP INC	COMMON STOCK	624580106	48	2,864	SH		OTR	1	2,864	0	0
MOVADO CORP	COMMON STOCK	624580106	6	361	SH		DFND	5	361	0	0
MUELLER INDS INC	COMMON STOCK	624756102	63	1,800	SH		DFND	1	1,800	0	0
MUELLER WTR PRODS INC	COMMON STOCK	624758108	661	53,386	SH		DFND	1	51,886	0	1,500
MUELLER WTR PRODS INC	COMMON STOCK	624758108	154	12,454	SH		OTR	1	12,454	0	0
MURPHY OIL CORP	COMMON STOCK	626717102	212	17,508	SH		DFND	1	10,342	0	7,166
MURPHY OIL CORP	COMMON STOCK	626717102	82	6,802	SH		OTR	1	6,048	0	754
MURPHY OIL CORP	COMMON STOCK	626717102	1	60	SH		DFND	5	60	0	0
MURPHY USA INC W I	COMMON STOCK	626755102	310	2,365	SH		DFND	1	617	0	1,748
MURPHY USA INC W I	COMMON STOCK	626755102	5	36	SH		OTR	1	19	5	12
MYERS INDS INC	COMMON STOCK	628464109	52	2,507	SH		DFND	1	2,507	0	0
N B T BANCORP INC	COMMON STOCK	628778102	33	1,040	SH		DFND	1	1,040	0	0
NMI HOLDINGS INC CLASS A	COMMON STOCK	629209305	40	1,782	SH		DFND	1	1,782	0	0
NRG ENERGY INC	COMMON STOCK	629377508	455	12,126	SH		DFND	1	8,678	0	3,448
NRG ENERGY INC	COMMON STOCK	629377508	26	698	SH		OTR	1	506	192	0
NRG ENERGY INC	COMMON STOCK	629377508	5	131	SH		DFND	5	131	0	0
NVE CORP	COMMON STOCK	629445206	481	8,570	SH		DFND	1	8,360	0	210
NVE CORP	COMMON STOCK	629445206	98	1,748	SH		OTR	1	1,673	0	75
NANOVIRICIDES INC	COMMON STOCK	630087302	0	43	SH		DFND	1	43	0	0
NAPCO SECURITY TECHNOLOGIES	COMMON STOCK	630402105	191	7,300	SH		DFND	5	7,300	0	0
NASDAQ INC	COMMON STOCK	631103108	1,960	14,764	SH		DFND	1	13,988	0	776
NASDAQ INC	COMMON STOCK	631103108	795	5,992	SH		OTR	1	5,433	0	559
NATERA INC	COMMON STOCK	632307104	2,073	20,826	SH		DFND	1	20,822	0	4
NATERA INC	COMMON STOCK	632307104	63	632	SH		OTR	1	632	0	0
NATIONAL BEVERAGE CORP	COMMON STOCK	635017106	1	14	SH		DFND	1	0	0	14
NATIONAL BEVERAGE CORP	COMMON STOCK	635017106	2	20	SH		OTR	1	20	0	0
NATIONAL CINEMEDIA INC	COMMON STOCK	635309107	0	67	SH		DFND	1	67	0	0
NATIONAL HEALTHCARE CORP	COMMON STOCK	635906100	283	4,273	SH		DFND	1	4,120	0	153
NATIONAL HEALTHCARE CORP	COMMON STOCK	635906100	48	724	SH		OTR	1	724	0	0
NATIONAL FUEL GAS CO NJ	COMMON STOCK	636180101	413	10,034	SH		DFND	1	8,049	0	1,985
NATIONAL FUEL GAS CO NJ	COMMON STOCK	636180101	67	1,623	SH		OTR	1	1,599	0	24
NATIONAL GENERAL HLDGS	COMMON STOCK	636220303	28	831	SH		DFND	1	282	0	549
NATIONAL GRID PLC A D R	COMMON STOCK	636274409	666	11,295	SH		DFND	1	6,065	0	5,230
NATIONAL GRID PLC A D R	COMMON STOCK	636274409	151	2,558	SH		OTR	1	1,831	638	89
NATIONAL GRID PLC	COMMON STOCK	636274409	246	4,174	SH		DFND	5	4,174	0	0
NATIONAL INSTRS CORP	COMMON STOCK	636518102	2,854	64,951	SH		DFND	1	62,926	0	2,025
NATIONAL INSTRS CORP	COMMON STOCK	636518102	85	1,952	SH		OTR	1	1,922	0	30
NATIONAL INSTRUMENTS CORP	COMMON STOCK	636518102	161	3,664	SH		DFND	5	3,664	0	0
NATIONAL OILWELL VARCO INC	COMMON STOCK	637071101	376	27,432	SH		DFND	1	24,814	0	2,618
NATIONAL OILWELL VARCO INC	COMMON STOCK	637071101	79	5,754	SH		OTR	1	5,586	0	168
NATIONAL OILWELL VARCO INC	COMMON STOCK	637071101	0	35	SH		DFND	5	35	0	0
NATIONAL PRESTO INDS INC	COMMON STOCK	637215104	84	954	SH		DFND	1	954	0	0
NATIONAL RESEARCH CORP	COMMON STOCK	637372202	2,622	61,325	SH		DFND	1	61,272	0	53
NATIONAL RESEARCH CORP	COMMON STOCK	637372202	62	1,455	SH		OTR	1	65	0	1,390
NATIONAL RETAIL PROPERTIES INC	COMMON STOCK	637417106	327	7,995	SH		DFND	1	6,571	0	1,424
NATIONAL RETAIL PROPERTIES INC	COMMON STOCK	637417106	19	459	SH		OTR	1	366	67	26
NATIONAL STORAGE AFFILIATES	COMMON STOCK	637870106	11	301	SH		DFND	1	301	0	0
NATIONAL STORAGE AFFILIATES	COMMON STOCK	637870106	53	1,475	SH		OTR	1	0	1,475	0
NATIONAL WESTERN LIFE GROU A	COMMON STOCK	638517102	17	82	SH		DFND	1	0	0	82
NATIONAL WESTERN LIFE GROU A	COMMON STOCK	638517102	1	6	SH		OTR	1	0	6	0
NATURES SUNSHINE PRODS INC	COMMON STOCK	639027101	904	60,492	SH		DFND	1	60,492	0	0
NATUS MEDICAL INC	COMMON STOCK	639050103	0	16	SH		DFND	1	16	0	0
NATWEST GROUP PLC A D R	COMMON STOCK	639057108	49	10,779	SH		DFND	1	6,439	0	4,340
NATWEST GROUP PLC A D R	COMMON STOCK	639057108	52	11,420	SH		OTR	1	11,064	0	356
NEENAH INC	COMMON STOCK	640079109	53	964	SH		DFND	1	964	0	0
NEENAH INC	COMMON STOCK	640079109	6	104	SH		OTR	1	104	0	0
NEKTAR THERAPEUTICS	COMMON STOCK	640268108	63	3,694	SH		DFND	1	3,310	0	384
NEKTAR THERAPEUTICS	COMMON STOCK	640268108	4,613	271,330	SH		OTR	1	271,330	0	0
NEOGEN CORP	COMMON STOCK	640491106	3,682	46,430	SH		DFND	1	38,960	0	7,470
NEOGEN CORP	COMMON STOCK	640491106	563	7,105	SH		OTR	1	3,797	575	2,733
NEOGEN CORP	COMMON STOCK	640491106	199	2,515	SH		DFND	5	2,515	0	0
NEW AMERICA HIGH INCOME FUND	OTHER	641876800	0	6	SH		DFND	1	6	0	0
NEW GOLD INC	COMMON STOCK	644535106	1	509	SH		DFND	1	509	0	0
NEW JERSEY RESOURCES CORP	COMMON STOCK	646025106	331	9,317	SH		DFND	1	8,518	0	799
NEW JERSEY RESOURCES CORP	COMMON STOCK	646025106	5	134	SH		OTR	1	134	0	0
NEW ORIENTAL EDUCATION SP A D	COMMON STOCK	647581107	1,021	5,495	SH		DFND	1	4,548	0	947
NEW ORIENTAL EDUCATION SP A D	COMMON STOCK	647581107	266	1,434	SH		OTR	1	1,304	124	6
NEW SENIOR INVESTMENTS GROUP	COMMON STOCK	648691103	3	518	SH		DFND	1	252	0	266
NEW SENIOR INVESTMENTS GROUP	COMMON STOCK	648691103	1	223	SH		OTR	1	0	0	223
NEW YORK COMMUNITY BANCORP INC	COMMON STOCK	649445103	312	29,652	SH		DFND	1	27,563	0	2,089
NEW YORK COMMUNITY BANCORP INC	COMMON STOCK	649445103	138	13,065	SH		OTR	1	13,065	0	0
NEW YORK MORTGAGE TRUST INC	COMMON STOCK	649604501	77	20,846	SH		DFND	1	18,656	0	2,190
NEW YORK TIMES CO	COMMON STOCK	650111107	128	2,481	SH		DFND	1	373	0	2,108
NEW YORK TIMES CO	COMMON STOCK	650111107	202	3,894	SH		OTR	1	0	0	3,894
NEW YORK TIMES CO	COMMON STOCK	650111107	2	42	SH		DFND	5	42	0	0
NEWELL BRANDS INC	COMMON STOCK	651229106	635	29,895	SH		DFND	1	26,538	0	3,357
NEWELL BRANDS INC	COMMON STOCK	651229106	551	25,936	SH		OTR	1	14,979	293	10,664
NEWMARKET CORP	COMMON STOCK	651587107	52	130	SH		DFND	1	62	0	68
NEWMONT CORPORATION	COMMON STOCK	651639106	3,327	55,562	SH		DFND	1	24,840	0	30,722
NEWMONT CORPORATION	COMMON STOCK	651639106	1,775	29,653	SH		OTR	1	15,424	8,051	6,178
NEWMONT MINING CORP	COMMON STOCK	651639106	27	446	SH		DFND	5	446	0	0
NICE LTD A D R	COMMON STOCK	653656108	19,673	69,382	SH		DFND	1	67,509	0	1,873
NICE LTD A D R	COMMON STOCK	653656108	3,658	12,901	SH		OTR	1	12,735	7	159
NICE SYSTEMS LTD	COMMON STOCK	653656108	15	53	SH		DFND	5	53	0	0
NIKE INC	COMMON STOCK	654106103	132,673	937,821	SH		DFND	1	907,428	0	30,393
NIKE INC	COMMON STOCK	654106103	49,867	352,492	SH		OTR	1	225,631	6,492	120,369
NIKE INC	COMMON STOCK	654106103	714	5,050	SH		DFND	5	4,784	0	266
NIKOLA CORP COM	COMMON STOCK	654110105	48	3,134	SH		DFND	1	3,134	0	0
NOKIA CORP SPON A D R	COMMON STOCK	654902204	8,084	2,067,461	SH		DFND	1	2,049,651	0	17,810
NOKIA CORP SPON A D R	COMMON STOCK	654902204	2,259	577,860	SH		OTR	1	574,382	818	2,660
NOKIA CORP	COMMON STOCK	654902204	16	3,967	SH		DFND	5	3,967	0	0
NORDSON CORP	COMMON STOCK	655663102	822	4,090	SH		DFND	1	2,806	0	1,284
NORDSON CORP	COMMON STOCK	655663102	392	1,949	SH		OTR	1	340	693	916
NORDSON CORP	COMMON STOCK	655663102	65	324	SH		DFND	5	324	0	0
NORDSTROM INC	COMMON STOCK	655664100	1,395	44,690	SH		DFND	1	44,630	0	60
NORDSTROM INC	COMMON STOCK	655664100	635	20,365	SH		OTR	1	19,950	0	415
NORFOLK SOUTHN CORP	COMMON STOCK	655844108	37,123	156,236	SH		DFND	1	153,544	0	2,692
NORFOLK SOUTHN CORP	COMMON STOCK	655844108	19,814	83,386	SH		OTR	1	78,235	2,014	3,137
NORFOLK SOUTHERN CORP	COMMON STOCK	655844108	26	107	SH		DFND	5	107	0	0
NORTHERN TRUST CORP	COMMON STOCK	665859104	4,262	45,761	SH		DFND	1	41,103	0	4,658
NORTHERN TRUST CORP	COMMON STOCK	665859104	1,673	17,955	SH		OTR	1	10,975	0	6,980
NORTHERN TRUST CORP	COMMON STOCK	665859104	69	739	SH		DFND	5	739	0	0
NORTHROP GRUMMAN CORPORATION	COMMON STOCK	666807102	15,667	51,415	SH		DFND	1	48,674	0	2,741
NORTHROP GRUMMAN CORPORATION	COMMON STOCK	666807102	4,479	14,698	SH		OTR	1	13,026	392	1,280
NORTHROP GRUMMAN CORP	COMMON STOCK	666807102	42	138	SH		DFND	5	118	0	20
NORTHWEST BANCSHARES INC MD	COMMON STOCK	667340103	17	1,347	SH		DFND	1	784	0	563
NORTHWEST PIPE CO	COMMON STOCK	667746101	3	108	SH		DFND	1	108	0	0
NORTHWESTERN CORP	COMMON STOCK	668074305	728	12,484	SH		DFND	1	12,231	0	253
NORTHWESTERN CORP	COMMON STOCK	668074305	112	1,918	SH		OTR	1	1,918	0	0
NORTONLIFELOCK INC	COMMON STOCK	668771108	1,790	86,160	SH		DFND	1	83,820	0	2,340
NORTONLIFELOCK INC	COMMON STOCK	668771108	827	39,799	SH		OTR	1	39,610	189	0
NORTONLIFELOCK INC	COMMON STOCK	668771108	36	1,754	SH		DFND	5	1,754	0	0
NOVAVAX INC	COMMON STOCK	670002401	706	6,329	SH		DFND	1	6,214	0	115
NOVAVAX INC	COMMON STOCK	670002401	42	377	SH		OTR	1	203	163	11
NOVO NORDISK AS A D R	COMMON STOCK	670100205	8,760	125,412	SH		DFND	1	106,584	0	18,828
NOVO NORDISK AS A D R	COMMON STOCK	670100205	1,809	25,898	SH		OTR	1	24,997	409	492
NOVO-NORDISK A/S	COMMON STOCK	670100205	21	305	SH		DFND	5	282	0	23
NUCOR CORP	COMMON STOCK	670346105	2,733	51,394	SH		DFND	1	49,108	0	2,286
NUCOR CORP	COMMON STOCK	670346105	1,321	24,830	SH		OTR	1	21,555	3,250	25
NUCOR CORPORATION	COMMON STOCK	670346105	3	65	SH		DFND	5	65	0	0
NV ENHCD AMT-FRE QLMUN INCME F	OTHER	670657105	335	22,167	SH		DFND	1	22,167	0	0
NV ENHCD AMT-FRE QLMUN INCME F	OTHER	670657105	46	3,014	SH		OTR	1	3,014	0	0
NUVEEN INSURED TAX FREE	OTHER	670657105	38	2,500	SH		DFND	5	0	0	2,500
NUVEEN MUN HIGH INCOME OPPORTU	OTHER	670682103	140	9,705	SH		DFND	1	9,705	0	0
NUVEEN NASDAQ 100 DYNAMIC OVER	OTHER	670699107	26	1,000	SH		DFND	1	1,000	0	0
NUVASIVE INC	COMMON STOCK	670704105	335	5,945	SH		DFND	1	4,493	0	1,452
NUVASIVE INC	COMMON STOCK	670704105	49	870	SH		OTR	1	87	783	0
NUVEEN MINNESOTA MUNICIPAL	OTHER	670734102	14	959	SH		DFND	1	959	0	0
O G E ENERGY CORP	COMMON STOCK	670837103	479	15,035	SH		DFND	1	14,379	0	656
O G E ENERGY CORP	COMMON STOCK	670837103	278	8,732	SH		OTR	1	8,732	0	0
OGE ENERGY CORP	COMMON STOCK	670837103	22	703	SH		DFND	5	703	0	0
NUVEEN MUN VALUE FD INC	OTHER	670928100	175	15,788	SH		DFND	1	15,788	0	0
NUVEEN MUN VALUE FD INC	OTHER	670928100	24	2,200	SH		OTR	1	2,200	0	0
NUVEEN OHIO QUALITY INCOME MUN	OTHER	670980101	348	22,250	SH		DFND	1	22,250	0	0
O S I SYS INC	COMMON STOCK	671044105	26	274	SH		DFND	1	251	0	23
OAK VALLEY BANCORP	COMMON STOCK	671807105	8	486	SH		DFND	1	486	0	0
OCCIDENTAL PETROLEUM CORPORATI	COMMON STOCK	674599105	437	25,218	SH		DFND	1	18,880	0	6,338
OCCIDENTAL PETROLEUM CORPORATI	COMMON STOCK	674599105	172	9,903	SH		OTR	1	9,549	148	206
OCCIDENTAL PETROLEUM CORP WT 8	OTHER	674599162	32	4,755	SH		DFND	1	4,079	0	676
OCCIDENTAL PETROLEUM CORP WT 8	OTHER	674599162	10	1,476	SH		OTR	1	1,270	72	134
OCCIDENTAL PETROLEUM CORP	OTHER	674599162	0	6	SH		DFND	5	0	0	6
OCEAN POWER TECHNOLOGIES INC	COMMON STOCK	674870506	2	800	SH		DFND	1	800	0	0
OCEANEERING INTERNATIONAL INC	COMMON STOCK	675232102	32	3,943	SH		DFND	1	3,943	0	0
OCEANEERING INTERNATIONAL INC	COMMON STOCK	675232102	5	656	SH		OTR	1	656	0	0
OCEANFIRST FINL CORP	COMMON STOCK	675234108	10	553	SH		DFND	1	255	0	298
ODONATE THERAPEUTICS INC	COMMON STOCK	676079106	1	62	SH		DFND	1	62	0	0
OHIO VALLEY BANC CORP	COMMON STOCK	677719106	46	1,929	SH		DFND	1	1,929	0	0
OIL DRI CORP AMER	COMMON STOCK	677864100	13	385	SH		DFND	1	385	0	0
OIL STATES INTERNATIONAL INC	COMMON STOCK	678026105	0	94	SH		DFND	1	94	0	0
OKTA INC	COMMON STOCK	679295105	5,866	23,071	SH		DFND	1	22,330	0	741
OKTA INC	COMMON STOCK	679295105	2,676	10,523	SH		OTR	1	5,180	4,324	1,019
OKTA INC	COMMON STOCK	679295105	52	203	SH		DFND	5	203	0	0
OLD DOMINION FGHT LINE INC	COMMON STOCK	679580100	3,734	19,134	SH		DFND	1	17,448	0	1,686
OLD DOMINION FGHT LINE INC	COMMON STOCK	679580100	133	683	SH		OTR	1	600	34	49
OLD DOMINION FREIGHT LINE	COMMON STOCK	679580100	3	13	SH		DFND	5	13	0	0
OLD NATL BANCORP IND	COMMON STOCK	680033107	277	16,764	SH		DFND	1	16,631	0	133
OLD NATL BANCORP IND	COMMON STOCK	680033107	77	4,640	SH		OTR	1	4,640	0	0
OLD NATIONAL BANCORP	COMMON STOCK	680033107	13	774	SH		DFND	5	774	0	0
OLD REPUBLIC INTL CORP	COMMON STOCK	680223104	9,108	462,103	SH		DFND	1	23,218	0	438,885
OLD REPUBLIC INTL CORP	COMMON STOCK	680223104	122	6,182	SH		OTR	1	6,182	0	0
OLIN CORP NEW	COMMON STOCK	680665205	229	9,342	SH		DFND	1	4,237	0	5,105
OLIN CORP NEW	COMMON STOCK	680665205	129	5,237	SH		OTR	1	128	5,109	0
OLLIES BARGAIN OUTLET HOLDI	COMMON STOCK	681116109	5,507	67,349	SH		DFND	1	65,205	0	2,144
OLLIES BARGAIN OUTLET HOLDI	COMMON STOCK	681116109	1,076	13,150	SH		OTR	1	12,527	601	22
OLLIES BARGAIN OUTLET HLDGS	COMMON STOCK	681116109	222	2,716	SH		DFND	5	2,716	0	0
OMNICOM GROUP INC	COMMON STOCK	681919106	4,562	73,152	SH		DFND	1	54,380	0	18,772
OMNICOM GROUP INC	COMMON STOCK	681919106	2,008	32,192	SH		OTR	1	30,342	0	1,850
OMNICOM GROUP	COMMON STOCK	681919106	14	231	SH		DFND	5	231	0	0
OMEGA HEALTHCARE INVESTORS	COMMON STOCK	681936100	836	23,006	SH		DFND	1	21,678	0	1,328
OMEGA HEALTHCARE INVESTORS	COMMON STOCK	681936100	106	2,924	SH		OTR	1	2,820	0	104
OMEGA HEALTHCARE INVESTORS INC	COMMON STOCK	681936100	1	18	SH		DFND	5	18	0	0
OMEGA FLEX INC	COMMON STOCK	682095104	120	821	SH		DFND	1	821	0	0
OMEGA FLEX INC	COMMON STOCK	682095104	118	805	SH		OTR	1	40	0	765
OMEROS CORPORATION	COMMON STOCK	682143102	6	436	SH		DFND	1	0	0	436
ON SEMICONDUCTOR CORPORATION	COMMON STOCK	682189105	1,857	56,752	SH		DFND	1	53,849	0	2,903
ON SEMICONDUCTOR CORPORATION	COMMON STOCK	682189105	45	1,377	SH		OTR	1	1,377	0	0
ON SEMICONDUCTOR CORP	COMMON STOCK	682189105	20	596	SH		DFND	5	596	0	0
ONE LIBERTY PROPERTIES INC	COMMON STOCK	682406103	4	217	SH		DFND	1	217	0	0
ONE LIBERTY PROPERTIES INC	COMMON STOCK	682406103	20	1,000	SH		OTR	1	1,000	0	0
ONEOK INC	COMMON STOCK	682680103	485	12,659	SH		DFND	1	9,677	0	2,982
ONEOK INC	COMMON STOCK	682680103	306	7,960	SH		OTR	1	4,560	1,061	2,339
ONEOK INC	COMMON STOCK	682680103	46	1,204	SH		DFND	5	1,204	0	0
ONTO INNOVATION INC	COMMON STOCK	683344105	230	4,829	SH		DFND	1	4,829	0	0
ONTO INNOVATION INC	COMMON STOCK	683344105	149	3,122	SH		OTR	1	1,470	1,652	0
ONTO INNOVATION INC	COMMON STOCK	683344105	10	216	SH		DFND	5	216	0	0
ONTRAK INC COM	COMMON STOCK	683373104	2	38	SH		DFND	1	0	0	38
OPEN TEXT CORPORATION	COMMON STOCK	683715106	224	4,924	SH		DFND	1	2,881	0	2,043
OPEN TEXT CORPORATION	COMMON STOCK	683715106	30	669	SH		OTR	1	595	0	74
ORANGE SPON A D R	COMMON STOCK	684060106	237	19,997	SH		DFND	1	10,040	0	9,957
ORANGE SPON A D R	COMMON STOCK	684060106	25	2,101	SH		OTR	1	1,500	471	130
ORIX CORP SPONS A D R	COMMON STOCK	686330101	1,535	19,837	SH		DFND	1	16,761	0	3,076
ORIX CORP SPONS A D R	COMMON STOCK	686330101	244	3,160	SH		OTR	1	2,772	362	26
ORIX CORPORATION	COMMON STOCK	686330101	1	7	SH		DFND	5	7	0	0
ORMAT TECHNOLOGIES INC	COMMON STOCK	686688102	196	2,168	SH		DFND	1	2,168	0	0
ORMAT TECHNOLOGIES INC	COMMON STOCK	686688102	69	764	SH		OTR	1	764	0	0
ORMAT TECHNOLOGIES INC	COMMON STOCK	686688102	1	13	SH		DFND	5	13	0	0
OSHKOSH CORPORATION	COMMON STOCK	688239201	441	5,116	SH		DFND	1	4,891	0	225
OSHKOSH CORPORATION	COMMON STOCK	688239201	28	325	SH		OTR	1	325	0	0
OTTER TAIL CORP	COMMON STOCK	689648103	692	16,248	SH		DFND	1	15,393	0	855
OTTER TAIL CORP	COMMON STOCK	689648103	348	8,169	SH		OTR	1	7,449	0	720
OVERSTOCK.COM INC	COMMON STOCK	690370101	20	410	SH		OTR	1	0	410	0
OWENS & MINOR INC	COMMON STOCK	690732102	9	328	SH		DFND	1	328	0	0
OWENS CORNING INC	COMMON STOCK	690742101	813	10,737	SH		DFND	1	9,989	0	748
OWENS CORNING INC	COMMON STOCK	690742101	73	973	SH		OTR	1	947	0	26
OWENS CORNING INC	COMMON STOCK	690742101	28	365	SH		DFND	5	365	0	0
OXFORD INDS INC	COMMON STOCK	691497309	365	5,563	SH		DFND	1	4,282	0	1,281
OXFORD INDS INC	COMMON STOCK	691497309	50	768	SH		OTR	1	768	0	0
P I C O HOLDINGS INC	COMMON STOCK	693366205	16	1,700	SH		DFND	1	1,700	0	0
P N C FINANCIAL SERVICES GROUP	COMMON STOCK	693475105	48,203	323,509	SH		DFND	1	305,648	0	17,861
P N C FINANCIAL SERVICES GROUP	COMMON STOCK	693475105	11,307	75,882	SH		OTR	1	64,406	7,091	4,385
PNC FINANCIAL SERVICES GROUP	COMMON STOCK	693475105	286	1,920	SH		DFND	5	1,920	0	0
POSCO A D R	COMMON STOCK	693483109	1,115	17,893	SH		DFND	1	9,847	0	8,046
POSCO A D R	COMMON STOCK	693483109	101	1,622	SH		OTR	1	1,343	20	259
P P G INDS INC	COMMON STOCK	693506107	22,476	155,843	SH		DFND	1	154,169	0	1,674
P P G INDS INC	COMMON STOCK	693506107	20,827	144,409	SH		OTR	1	140,412	3,400	597
PPG INDUSTRIES INC	COMMON STOCK	693506107	14	97	SH		DFND	5	97	0	0
PVH CORP	COMMON STOCK	693656100	511	5,443	SH		DFND	1	4,440	0	1,003
PVH CORP	COMMON STOCK	693656100	240	2,557	SH		OTR	1	123	98	2,336
PACCAR INC	COMMON STOCK	693718108	225,417	2,612,620	SH		DFND	1	2,391,281	0	221,339
PACCAR INC	COMMON STOCK	693718108	826	9,584	SH		OTR	1	8,675	377	532
PACCAR INC	COMMON STOCK	693718108	1	13	SH		DFND	5	13	0	0
PACIRA PHARMACEUTICALS INC	COMMON STOCK	695127100	717	11,998	SH		DFND	1	11,901	0	97
PACIRA PHARMACEUTICALS INC	COMMON STOCK	695127100	84	1,406	SH		OTR	1	642	764	0
PACIRA PHARMACEUTICALS INC	COMMON STOCK	695127100	3	53	SH		DFND	5	53	0	0
PACKAGING CORP AMERICA	COMMON STOCK	695156109	6,609	47,926	SH		DFND	1	45,604	0	2,322
PACKAGING CORP AMERICA	COMMON STOCK	695156109	1,416	10,270	SH		OTR	1	7,982	0	2,288
PACKAGING CORP OF AMERICA	COMMON STOCK	695156109	70	503	SH		DFND	5	503	0	0
PACWEST BANCORP	COMMON STOCK	695263103	455	17,929	SH		DFND	1	17,689	0	240
PACWEST BANCORP	COMMON STOCK	695263103	87	3,462	SH		OTR	1	3,347	56	59
PALATIN TECHNOLOGIES INC	COMMON STOCK	696077403	106	156,740	SH		OTR	1	156,740	0	0
PALO ALTO NETWORKS INC	COMMON STOCK	697435105	32,638	91,840	SH		DFND	1	90,296	0	1,544
PALO ALTO NETWORKS INC	COMMON STOCK	697435105	6,920	19,473	SH		OTR	1	18,631	391	451
PALO ALTO NETWORKS INC	COMMON STOCK	697435105	568	1,599	SH		DFND	5	1,599	0	0
PAN AMERICAN SILVER CORP	COMMON STOCK	697900108	56	1,613	SH		DFND	1	739	0	874
PAN AMERICAN SILVER CORP	COMMON STOCK	697900108	1	30	SH		OTR	1	0	0	30
PAPA JOHNS INTL INC	COMMON STOCK	698813102	38	444	SH		DFND	1	431	0	13
PARK HOTELS RESORTS INC WI	COMMON STOCK	700517105	272	15,873	SH		DFND	1	12,681	0	3,192
PARK HOTELS RESORTS INC WI	COMMON STOCK	700517105	33	1,869	SH		OTR	1	1,082	582	205
PARK NATIONAL CORP	COMMON STOCK	700658107	315	2,997	SH		DFND	1	2,697	0	300
PARK NATIONAL CORP	COMMON STOCK	700658107	202	1,928	SH		OTR	1	88	840	1,000
PARKER HANNIFIN CORP	COMMON STOCK	701094104	13,351	49,014	SH		DFND	1	30,050	0	18,964
PARKER HANNIFIN CORP	COMMON STOCK	701094104	1,580	5,800	SH		OTR	1	5,773	19	8
PARKER HANNIFIN CO	COMMON STOCK	701094104	88	321	SH		DFND	5	321	0	0
PARSLEY ENERGY INC CLASS A	COMMON STOCK	701877102	1,792	126,210	SH		DFND	1	107,246	0	18,964
PARSLEY ENERGY INC CLASS A	COMMON STOCK	701877102	463	32,592	SH		OTR	1	30,796	0	1,796
PARSLEY ENERGY INC	COMMON STOCK	701877102	70	4,903	SH		DFND	5	4,903	0	0
PARTY CITY HOLDCO INC	COMMON STOCK	702149105	9	1,516	SH		DFND	1	1,100	0	416
PATRICK INDUSTRIES INC	COMMON STOCK	703343103	22	319	SH		DFND	1	319	0	0
PATTERSON COMPANIES INC	COMMON STOCK	703395103	1,010	34,066	SH		DFND	1	33,580	0	486
PATTERSON COMPANIES INC	COMMON STOCK	703395103	264	8,901	SH		OTR	1	8,901	0	0
PATTERSON U T I ENERGY INC	COMMON STOCK	703481101	67	12,752	SH		DFND	1	7,363	0	5,389
PATTERSON U T I ENERGY INC	COMMON STOCK	703481101	16	3,047	SH		OTR	1	1,000	1,576	471
PAYCHEX INC	COMMON STOCK	704326107	7,484	80,320	SH		DFND	1	67,057	0	13,263
PAYCHEX INC	COMMON STOCK	704326107	6,361	68,265	SH		OTR	1	67,698	467	100
PAYCHEX INC	COMMON STOCK	704326107	218	2,342	SH		DFND	5	2,342	0	0
PEAPACK GLADSTONE FINL CORP	COMMON STOCK	704699107	8	350	SH		DFND	1	350	0	0
PEARSON P L C A D R	COMMON STOCK	705015105	243	26,485	SH		DFND	1	21,081	0	5,404
PEARSON P L C A D R	COMMON STOCK	705015105	47	5,140	SH		OTR	1	5,002	0	138
PEGASYSTEMS INC	COMMON STOCK	705573103	1,891	14,190	SH		DFND	1	14,020	0	170
PEGASYSTEMS INC	COMMON STOCK	705573103	201	1,509	SH		OTR	1	1,488	0	21
PEGASYSTEMS INC	COMMON STOCK	705573103	377	2,826	SH		DFND	5	2,826	0	0
PEMBINA PIPELINE CORP	COMMON STOCK	706327103	33	1,408	SH		DFND	1	1,408	0	0
PEMBINA PIPELINE CORP	COMMON STOCK	706327103	15	627	SH		OTR	1	627	0	0
PEMBINA PIPELINE CORP	COMMON STOCK	706327103	5	193	SH		DFND	5	193	0	0
PENN NATIONAL GAMING INC	COMMON STOCK	707569109	402	4,658	SH		DFND	1	4,529	0	129
PENN NATIONAL GAMING INC	COMMON STOCK	707569109	75	877	SH		OTR	1	678	0	199
PEOPLES BANCORP INC	COMMON STOCK	709789101	72	2,651	SH		DFND	1	2,577	0	74
PEOPLES UNITED FINANCIAL INC	COMMON STOCK	712704105	61	4,700	SH		DFND	1	2,988	0	1,712
PEOPLES UNITED FINANCIAL INC	COMMON STOCK	712704105	88	6,763	SH		OTR	1	6,700	63	0
PEOPLES UNITED FINANCIAL INC	COMMON STOCK	712704105	1	50	SH		DFND	5	50	0	0
PEPSICO INC	COMMON STOCK	713448108	218,349	1,472,352	SH		DFND	1	1,388,404	0	83,948
PEPSICO INC	COMMON STOCK	713448108	101,869	686,911	SH		OTR	1	585,168	22,620	79,123
PEPSICO INC	COMMON STOCK	713448108	499	3,361	SH		DFND	5	3,361	0	0
PERKIN ELMER INC	COMMON STOCK	714046109	310	2,164	SH		DFND	1	1,610	0	554
PERKIN ELMER INC	COMMON STOCK	714046109	4	28	SH		OTR	1	0	0	28
PERKINELMER INC	COMMON STOCK	714046109	39	274	SH		DFND	5	274	0	0
PERMA PIPE HOLDINGS INC	COMMON STOCK	714167103	5	799	SH		DFND	1	799	0	0
PERMIAN BASIN ROYALTY TRUST	COMMON STOCK	714236106	1	300	SH		DFND	1	300	0	0
PERSPECTA INC	COMMON STOCK	715347100	87	3,613	SH		DFND	1	2,789	0	824
PERSPECTA INC	COMMON STOCK	715347100	80	3,326	SH		OTR	1	3,307	0	19
P T TELEKOMUNIKASI INDONESIA A	COMMON STOCK	715684106	72	3,048	SH		DFND	1	2,465	0	583
PETMED EXPRESS COM INC	COMMON STOCK	716382106	28	882	SH		DFND	1	882	0	0
PFIZER INC	COMMON STOCK	717081103	197,581	5,367,587	SH		DFND	1	5,240,261	0	127,326
PFIZER INC	COMMON STOCK	717081103	88,881	2,414,605	SH		OTR	1	2,306,926	44,770	62,909
PFIZER INC	COMMON STOCK	717081103	389	10,572	SH		DFND	5	10,332	0	240
PFIZER INC	COMMON STOCK	717081103	7	37	SH	Put	DFND	1	37	0	0
PHILIP MORRIS INTL	COMMON STOCK	718172109	35,735	431,630	SH		DFND	1	395,611	0	36,019
PHILIP MORRIS INTL	COMMON STOCK	718172109	30,147	364,147	SH		OTR	1	350,462	10,586	3,099
PHILIP MORRIS INTERNATIONAL	COMMON STOCK	718172109	637	7,692	SH		DFND	5	7,619	0	73
PHILLIPS 66	COMMON STOCK	718546104	11,754	168,060	SH		DFND	1	158,958	0	9,102
PHILLIPS 66	COMMON STOCK	718546104	5,311	75,936	SH		OTR	1	73,064	301	2,571
PHILLIPS 66	COMMON STOCK	718546104	124	1,765	SH		DFND	5	1,765	0	0
PHILLIPS 66 PARTNERS LP	COMMON STOCK	718549207	3	116	SH		DFND	5	116	0	0
PHOTRONICS INC	COMMON STOCK	719405102	3	254	SH		DFND	1	222	0	32
PIEDMONT OFFICE REALTY TR CLAS	COMMON STOCK	720190206	316	19,481	SH		DFND	1	19,481	0	0
PIEDMONT OFFICE REALTY TR CLAS	COMMON STOCK	720190206	97	5,980	SH		OTR	1	5,980	0	0
PIERIS PHARMACEUTICALS INC	COMMON STOCK	720795103	26	10,280	SH		OTR	1	10,280	0	0
PIMCO HIGH INCOME	OTHER	722014107	93	15,445	SH		DFND	5	9,645	0	5,800
PINDUODUO INC A D R	COMMON STOCK	722304102	1,850	10,410	SH		DFND	1	10,410	0	0
PINDUODUO INC A D R	COMMON STOCK	722304102	235	1,325	SH		OTR	1	1,325	0	0
PINNACLE WEST CAP CORP	COMMON STOCK	723484101	2,834	35,445	SH		DFND	1	34,487	0	958
PINNACLE WEST CAP CORP	COMMON STOCK	723484101	1,051	13,146	SH		OTR	1	12,832	11	303
PINNACLE WEST CAPITAL	COMMON STOCK	723484101	15	182	SH		DFND	5	182	0	0
PIONEER DIVERSIFIED HIGH INC	OTHER	723653101	24	1,629	SH		DFND	1	1,629	0	0
PIONEER NATURAL RESOURCES CO	COMMON STOCK	723787107	3,363	29,533	SH		DFND	1	27,785	0	1,748
PIONEER NATURAL RESOURCES CO	COMMON STOCK	723787107	2,185	19,189	SH		OTR	1	16,778	872	1,539
PIONEER NATURAL RESOURCES CO	COMMON STOCK	723787107	31	270	SH		DFND	5	270	0	0
PIPER SANDLER CO	COMMON STOCK	724078100	1,031	10,215	SH		DFND	1	9,427	0	788
PIPER SANDLER CO	COMMON STOCK	724078100	456	4,522	SH		OTR	1	4,522	0	0
PIPER SANDLER COS	COMMON STOCK	724078100	43	429	SH		DFND	5	429	0	0
PITNEY BOWES INC	COMMON STOCK	724479100	22	3,652	SH		DFND	1	3,652	0	0
PITNEY BOWES INC	COMMON STOCK	724479100	6	957	SH		OTR	1	957	0	0
PLAINS ALL AMER PIPELINE LP	COMMON STOCK	726503105	623	75,574	SH		DFND	1	75,574	0	0
PLAINS ALL AMER PIPELINE LP	COMMON STOCK	726503105	21	2,575	SH		OTR	1	2,575	0	0
PLAINS ALL AMERICAN PIPELINE	COMMON STOCK	726503105	5	619	SH		DFND	5	619	0	0
PLANTRONICS INC	COMMON STOCK	727493108	8	292	SH		DFND	1	0	0	292
PLANTRONICS INC	COMMON STOCK	727493108	8	286	SH		OTR	1	0	286	0
PLEXUS CORP	COMMON STOCK	729132100	1,613	20,621	SH		DFND	1	19,657	0	964
PLEXUS CORP	COMMON STOCK	729132100	302	3,857	SH		OTR	1	3,857	0	0
POLARIS INC	COMMON STOCK	731068102	1,279	13,417	SH		DFND	1	13,068	0	349
POLARIS INC	COMMON STOCK	731068102	181	1,907	SH		OTR	1	1,435	472	0
POLARIS INDUSTRIES INC	COMMON STOCK	731068102	40	421	SH		DFND	5	421	0	0
POLYMET MNG CORP	COMMON STOCK	731916409	8	2,472	SH		DFND	1	2,472	0	0
POLYMET MNG CORP	COMMON STOCK	731916409	675	197,934	SH		OTR	1	197,934	0	0
POPULAR INC	COMMON STOCK	733174700	172	3,052	SH		DFND	1	2,839	0	213
POPULAR INC	COMMON STOCK	733174700	3	49	SH		OTR	1	0	0	49
PORTLAND GENERAL ELECTRIC CO	COMMON STOCK	736508847	679	15,876	SH		DFND	1	7,563	0	8,313
PORTLAND GENERAL ELECTRIC CO	COMMON STOCK	736508847	411	9,606	SH		OTR	1	9,606	0	0
PORTLAND GENERAL ELECTRIC CO	COMMON STOCK	736508847	35	827	SH		DFND	5	827	0	0
POST HOLDINGS INC	COMMON STOCK	737446104	8,618	85,319	SH		DFND	1	84,744	0	575
POST HOLDINGS INC	COMMON STOCK	737446104	1,813	17,954	SH		OTR	1	17,942	12	0
POTLATCHDELTIC CORP	COMMON STOCK	737630103	487	9,740	SH		DFND	1	9,712	0	28
POTLATCHDELTIC CORP	COMMON STOCK	737630103	239	4,764	SH		OTR	1	4,701	10	53
POWELL INDS INC	COMMON STOCK	739128106	6	201	SH		DFND	1	201	0	0
POWER INTEGRATIONS INC	COMMON STOCK	739276103	1,619	19,770	SH		DFND	1	19,762	0	8
POWER INTEGRATIONS INC	COMMON STOCK	739276103	281	3,427	SH		OTR	1	1,417	0	2,010
POWER INTEGRATIONS INC	COMMON STOCK	739276103	57	699	SH		DFND	5	699	0	0
PREFERRED BANK LOS ANGELES	COMMON STOCK	740367404	2	44	SH		DFND	1	44	0	0
PRICE SMART INC	COMMON STOCK	741511109	723	7,934	SH		DFND	1	7,902	0	32
PRICE SMART INC	COMMON STOCK	741511109	31	345	SH		OTR	1	345	0	0
PROCTER & GAMBLE CO	COMMON STOCK	742718109	427,140	3,069,856	SH		DFND	1	2,974,146	0	95,710
PROCTER & GAMBLE CO	COMMON STOCK	742718109	198,219	1,424,607	SH		OTR	1	1,320,143	64,623	39,841
PROCTER & GAMBLE CO	COMMON STOCK	742718109	276	1,982	SH		DFND	5	1,907	0	75
PROCTER & GAMBLE CO	COMMON STOCK	742718109	4	30	SH	Put	DFND	1	30	0	0
PROCTER & GAMBLE CO	COMMON STOCK	742718109	9	57	SH	Put	OTR	1	17	40	0
PROGRESS SOFTWARE CORP	COMMON STOCK	743312100	989	21,886	SH		DFND	1	20,504	0	1,382
PROGRESS SOFTWARE CORP	COMMON STOCK	743312100	121	2,670	SH		OTR	1	2,670	0	0
PROGRESSIVE CORP	COMMON STOCK	743315103	19,347	195,653	SH		DFND	1	143,584	0	52,069
PROGRESSIVE CORP	COMMON STOCK	743315103	2,283	23,093	SH		OTR	1	17,737	5,216	140
PROGRESSIVE CORP	COMMON STOCK	743315103	120	1,208	SH		DFND	5	1,196	0	12
PROOFPOINT INC	COMMON STOCK	743424103	3,440	25,221	SH		DFND	1	24,937	0	284
PROOFPOINT INC	COMMON STOCK	743424103	703	5,150	SH		OTR	1	5,136	0	14
PROSPERITY BANCSHARES INC	COMMON STOCK	743606105	327	4,707	SH		DFND	1	4,647	0	60
PROSPERITY BANCSHARES INC	COMMON STOCK	743606105	2	22	SH		OTR	1	22	0	0
PROSPERITY BANCSHARES INC	COMMON STOCK	743606105	5	65	SH		DFND	5	65	0	0
PROTO LABS INC	COMMON STOCK	743713109	2,683	17,492	SH		DFND	1	16,457	0	1,035
PROTO LABS INC	COMMON STOCK	743713109	5,294	34,509	SH		OTR	1	34,509	0	0
PROTO LABS INC	COMMON STOCK	743713109	306	1,992	SH		DFND	5	1,992	0	0
PROVIDENCE SERVICE CORP	COMMON STOCK	743815102	8	60	SH		DFND	1	0	0	60
PRUDENTIAL FINANCIAL INC	COMMON STOCK	744320102	11,657	149,319	SH		DFND	1	140,931	0	8,388
PRUDENTIAL FINANCIAL INC	COMMON STOCK	744320102	2,138	27,399	SH		OTR	1	25,702	569	1,128
PRUDENTIAL FINANCIAL	COMMON STOCK	744320102	4	51	SH		DFND	5	21	0	30
PUBLIC SVC ENTERPRISE GROUP IN	COMMON STOCK	744573106	2,218	38,042	SH		DFND	1	31,906	0	6,136
PUBLIC SVC ENTERPRISE GROUP IN	COMMON STOCK	744573106	817	14,011	SH		OTR	1	11,200	42	2,769
PUBLIC SVCS ENTERPRISE GROUP	COMMON STOCK	744573106	98	1,675	SH		DFND	5	1,675	0	0
PULTE GROUP INC	COMMON STOCK	745867101	795	18,432	SH		DFND	1	15,005	0	3,427
PULTE GROUP INC	COMMON STOCK	745867101	66	1,520	SH		OTR	1	1,496	0	24
PULTE HOMES INC	COMMON STOCK	745867101	2	36	SH		DFND	5	36	0	0
PUTNAM MANAGED MUN INCOME TR	OTHER	746823103	58	7,149	SH		DFND	1	7,149	0	0
PUTNAM PREMIER INC TR SBI	OTHER	746853100	16	3,500	SH		OTR	1	3,500	0	0
PUTNAM MUN OPPORTUNITIES TR SB	OTHER	746922103	62	4,545	SH		DFND	1	4,545	0	0
QUAKER CHEM CORP	COMMON STOCK	747316107	48	189	SH		DFND	1	189	0	0
QUAKER CHEMICAL CORP	COMMON STOCK	747316107	79	313	SH		DFND	5	313	0	0
QUALCOMM INC	COMMON STOCK	747525103	123,453	810,383	SH		DFND	1	777,364	0	33,019
QUALCOMM INC	COMMON STOCK	747525103	40,824	267,981	SH		OTR	1	249,228	13,685	5,068
QUALCOMM INC	COMMON STOCK	747525103	811	5,310	SH		DFND	5	5,124	0	186
QUALCOMM INC	COMMON STOCK	747525103	1	5	SH	Put	OTR	1	5	0	0
QUANEX BLDG PRODS CORP	COMMON STOCK	747619104	5	204	SH		DFND	1	178	0	26
QUDIAN INC SPON A D R	COMMON STOCK	747798106	2	1,162	SH		DFND	1	1,162	0	0
R1 RCM INC	COMMON STOCK	749397105	580	24,137	SH		DFND	1	23,412	0	725
R1 RCM INC	COMMON STOCK	749397105	108	4,500	SH		OTR	1	391	4,109	0
RLI CORP	COMMON STOCK	749607107	36	348	SH		DFND	1	348	0	0
RPC ENERGY SVCS INC	COMMON STOCK	749660106	1	266	SH		DFND	1	266	0	0
RPM INTERNATIONAL INC	COMMON STOCK	749685103	2,949	32,481	SH		DFND	1	25,656	0	6,825
RPM INTERNATIONAL INC	COMMON STOCK	749685103	989	10,894	SH		OTR	1	1,832	7,037	2,025
RPM INTERNATIONAL INC	COMMON STOCK	749685103	184	2,031	SH		DFND	5	2,031	0	0
RADIAN GROUP INC	COMMON STOCK	750236101	434	21,429	SH		DFND	1	20,024	0	1,405
RADIAN GROUP INC	COMMON STOCK	750236101	61	2,985	SH		OTR	1	275	2,710	0
RADIAN GROUP INC	COMMON STOCK	750236101	3	149	SH		DFND	5	149	0	0
RADIUS HEALTH INC	COMMON STOCK	750469207	20	1,141	SH		DFND	1	1,141	0	0
RADIUS HEALTH INC	COMMON STOCK	750469207	1	58	SH		OTR	1	58	0	0
RADNET INC	COMMON STOCK	750491102	47	2,403	SH		OTR	1	0	2,403	0
RAMBUS INC	COMMON STOCK	750917106	20	1,153	SH		DFND	1	1,153	0	0
RALPH LAUREN CORP	COMMON STOCK	751212101	108	1,041	SH		DFND	1	964	0	77
RALPH LAUREN CORP	COMMON STOCK	751212101	21	201	SH		OTR	1	201	0	0
RAPID7 INC	COMMON STOCK	753422104	1,419	15,736	SH		DFND	1	15,635	0	101
RAPID7 INC	COMMON STOCK	753422104	472	5,234	SH		OTR	1	5,206	13	15
RAPID7 INC	COMMON STOCK	753422104	70	773	SH		DFND	5	773	0	0
RAVEN INDS INC	COMMON STOCK	754212108	1,873	56,602	SH		DFND	1	56,579	0	23
RAYMOND JAMES FINL INC	COMMON STOCK	754730109	4,157	43,456	SH		DFND	1	42,512	0	944
RAYMOND JAMES FINL INC	COMMON STOCK	754730109	1,027	10,733	SH		OTR	1	10,729	0	4
RAYMOND JAMES FINANCIAL INC	COMMON STOCK	754730109	62	648	SH		DFND	5	628	0	20
RAYONIER INC	COMMON STOCK	754907103	122	4,171	SH		DFND	1	3,803	0	368
RAYONIER INC	COMMON STOCK	754907103	152	5,180	SH		OTR	1	5,134	0	46
REALTY INCOME CORP	COMMON STOCK	756109104	4,767	76,674	SH		DFND	1	70,287	0	6,387
REALTY INCOME CORP	COMMON STOCK	756109104	1,831	29,458	SH		OTR	1	29,442	0	16
REALTY INCOME CORP	COMMON STOCK	756109104	1	23	SH		DFND	5	23	0	0
REAVES UTILITY INCOME FUND	OTHER	756158101	219	6,667	SH		DFND	1	5,000	0	1,667
REAVES UTILITY INCOME FUND	OTHER	756158101	105	3,200	SH		OTR	1	3,200	0	0
REAVES UTILITY INCOME TR (THE)	OTHER	756158101	3	106	SH		DFND	5	0	0	106
REDHILL BIOPHARMA LTD SP A D R	COMMON STOCK	757468103	1	170	SH		DFND	1	109	0	61
REDWOOD TRUST INC	COMMON STOCK	758075402	26	2,955	SH		DFND	1	2,395	0	560
REGAL BELOIT CORP	COMMON STOCK	758750103	303	2,465	SH		DFND	1	367	0	2,098
REGAL BELOIT CORP	COMMON STOCK	758750103	65	526	SH		OTR	1	0	499	27
REGAL BELOIT CORP	COMMON STOCK	758750103	1	10	SH		DFND	5	10	0	0
REGENCY CENTERS CORPORATION	COMMON STOCK	758849103	295	6,451	SH		DFND	1	4,141	0	2,310
REGENCY CENTERS CORPORATION	COMMON STOCK	758849103	9	200	SH		OTR	1	180	20	0
REINSURANCE GROUP AMERICA	COMMON STOCK	759351604	1,210	10,440	SH		DFND	1	9,582	0	858
REINSURANCE GROUP AMERICA	COMMON STOCK	759351604	356	3,069	SH		OTR	1	2,761	0	308
REINSURANCE GROUP OF AMERICA	COMMON STOCK	759351604	24	209	SH		DFND	5	209	0	0
WTS REKOR SYSTEMS INC	COMMON STOCK	759419104	4	500	SH		DFND	1	500	0	0
RELIANCE STEEL ALUMINUM	COMMON STOCK	759509102	484	4,038	SH		DFND	1	3,221	0	817
RELIANCE STEEL ALUMINUM	COMMON STOCK	759509102	28	238	SH		OTR	1	184	33	21
RELIANCE STEEL & ALUMINUM CO	COMMON STOCK	759509102	10	87	SH		DFND	5	87	0	0
RELX PLC SPON A D R	COMMON STOCK	759530108	1,395	56,580	SH		DFND	1	48,342	0	8,238
RELX PLC SPON A D R	COMMON STOCK	759530108	56	2,260	SH		OTR	1	1,862	207	191
REPRO MEDSYSTEMS INC	COMMON STOCK	759910102	4	700	SH		DFND	1	700	0	0
REPLIGEN CORP	COMMON STOCK	759916109	741	3,865	SH		DFND	1	2,259	0	1,606
REPLIGEN CORP	COMMON STOCK	759916109	341	1,782	SH		OTR	1	146	63	1,573
REPLIGEN CORP	COMMON STOCK	759916109	35	185	SH		DFND	5	185	0	0
RENAISSANCE IPO ETF	OTHER	759937204	32	500	SH		OTR	1	0	500	0
REPUBLIC BANCORP INC CLASS A	COMMON STOCK	760281204	2	45	SH		DFND	1	45	0	0
REPUBLIC BANCORP INC CLASS A	COMMON STOCK	760281204	0	11	SH		OTR	1	0	11	0
REPUBLIC SVCS INC	COMMON STOCK	760759100	13,759	142,882	SH		DFND	1	137,425	0	5,457
REPUBLIC SVCS INC	COMMON STOCK	760759100	2,843	29,518	SH		OTR	1	29,167	131	220
REPUBLIC SERVICES INC	COMMON STOCK	760759100	45	467	SH		DFND	5	467	0	0
RESMED INC	COMMON STOCK	761152107	6,453	30,358	SH		DFND	1	27,234	0	3,124
RESMED INC	COMMON STOCK	761152107	2,572	12,099	SH		OTR	1	11,799	179	121
RESMED INC.	COMMON STOCK	761152107	404	1,898	SH		DFND	5	1,886	0	12
REVANCE THERAPEUTICS INC	COMMON STOCK	761330109	7	257	SH		DFND	1	198	0	59
REVLON INC CLASS A	COMMON STOCK	761525609	3	269	SH		DFND	1	0	0	269
REX AMERICAN RESOURCES CORP	COMMON STOCK	761624105	1	12	SH		DFND	1	0	0	12
RICEBRAN TECHNOLOGIES	COMMON STOCK	762831204	0	40	SH		DFND	1	40	0	0
RIGEL PHARMACEUTICALS INC	COMMON STOCK	766559603	3	992	SH		DFND	1	0	0	992
RIO TINTO PLC A D R	COMMON STOCK	767204100	12,345	164,108	SH		DFND	1	157,757	0	6,351
RIO TINTO PLC A D R	COMMON STOCK	767204100	2,573	34,212	SH		OTR	1	32,872	1,149	191
RIO TINTO-PLC	COMMON STOCK	767204100	41	543	SH		DFND	5	543	0	0
RITCHIE BROS AUCTIONEERS INC	COMMON STOCK	767744105	3,446	49,551	SH		DFND	1	49,158	0	393
RITCHIE BROS AUCTIONEERS INC	COMMON STOCK	767744105	354	5,090	SH		OTR	1	3,308	544	1,238
RITCHIE BROS AUCTIONEERS	COMMON STOCK	767744105	348	4,995	SH		DFND	5	4,995	0	0
RITE AID CORP	COMMON STOCK	767754872	19	1,172	SH		DFND	1	0	0	1,172
ROBERT HALF INTL INC	COMMON STOCK	770323103	767	12,270	SH		DFND	1	11,111	0	1,159
ROBERT HALF INTL INC	COMMON STOCK	770323103	427	6,838	SH		OTR	1	450	197	6,191
ROBERT HALF INTL INC	COMMON STOCK	770323103	8	136	SH		DFND	5	136	0	0
ROCKWELL AUTOMATION INC	COMMON STOCK	773903109	13,192	52,598	SH		DFND	1	49,531	0	3,067
ROCKWELL AUTOMATION INC	COMMON STOCK	773903109	6,678	26,626	SH		OTR	1	25,404	115	1,107
ROCKWELL AUTOMATION INC	COMMON STOCK	773903109	29	114	SH		DFND	5	114	0	0
RODGERS SILICON VALLEY 1 SH &	COMMON STOCK	774873202	1	100	SH		DFND	1	100	0	0
ROGERS COMMUNICATIONS INC CL B	COMMON STOCK	775109200	332	7,120	SH		DFND	1	4,799	0	2,321
ROGERS COMMUNICATIONS INC CL B	COMMON STOCK	775109200	28	595	SH		OTR	1	538	0	57
ROGERS CORP	COMMON STOCK	775133101	517	3,330	SH		DFND	1	1,768	0	1,562
ROGERS CORP	COMMON STOCK	775133101	224	1,441	SH		OTR	1	293	413	735
ROLLINS INC	COMMON STOCK	775711104	10,307	263,802	SH		DFND	1	263,089	0	713
ROLLINS INC	COMMON STOCK	775711104	3,790	97,016	SH		OTR	1	87,537	8,756	723
ROLLINS INC	COMMON STOCK	775711104	273	6,983	SH		DFND	5	6,983	0	0
ROMEO POWER INC	COMMON STOCK	776153108	182	8,077	SH		DFND	1	8,077	0	0
ROMEO POWER INC	COMMON STOCK	776153108	2	102	SH		OTR	1	102	0	0
ROPER INDS INC	COMMON STOCK	776696106	30,056	69,719	SH		DFND	1	68,728	0	991
ROPER INDS INC	COMMON STOCK	776696106	14,882	34,522	SH		OTR	1	11,417	809	22,296
ROPER INDUSTRIES INC	COMMON STOCK	776696106	422	978	SH		DFND	5	943	0	35
ROSS STORES INC	COMMON STOCK	778296103	8,483	69,076	SH		DFND	1	66,572	0	2,504
ROSS STORES INC	COMMON STOCK	778296103	4,738	38,585	SH		OTR	1	21,753	98	16,734
ROSS STORES INC	COMMON STOCK	778296103	34	275	SH		DFND	5	235	0	40
ROYAL BK CDA MONTREAL QUE	COMMON STOCK	780087102	7,399	90,160	SH		DFND	1	86,409	0	3,751
ROYAL BK CDA MONTREAL QUE	COMMON STOCK	780087102	309	3,767	SH		OTR	1	2,870	0	897
ROYAL BANK OF CANADA	COMMON STOCK	780087102	0	1	SH		DFND	5	0	0	1
ROYAL DUTCH SHELL A D R	COMMON STOCK	780259107	3,376	100,452	SH		DFND	1	98,934	0	1,518
ROYAL DUTCH SHELL A D R	COMMON STOCK	780259107	1,208	35,929	SH		OTR	1	32,008	3,066	855
ROYAL DUTCH SHELL PLC	COMMON STOCK	780259107	4	129	SH		DFND	5	129	0	0
ROYAL DUTCH SHELL PLC A D R	COMMON STOCK	780259206	3,848	109,505	SH		DFND	1	88,760	0	20,745
ROYAL DUTCH SHELL PLC A D R	COMMON STOCK	780259206	1,460	41,559	SH		OTR	1	23,494	7,061	11,004
ROYAL GOLD INC	COMMON STOCK	780287108	232	2,181	SH		DFND	1	791	0	1,390
ROYAL GOLD INC	COMMON STOCK	780287108	103	964	SH		OTR	1	41	65	858
ROYCE VALUE TRUST	OTHER	780910105	52	3,205	SH		DFND	1	0	0	3,205
ROYCE VALUE TRUST	OTHER	780910105	155	9,618	SH		OTR	1	0	9,618	0
RUSH ENTERPRISES INC	COMMON STOCK	781846209	1,002	24,197	SH		DFND	1	22,115	0	2,082
RUSH ENTERPRISES INC	COMMON STOCK	781846209	194	4,692	SH		OTR	1	3,714	978	0
RUTHS HOSPITALITY GROUP INC	COMMON STOCK	783332109	18	1,022	SH		DFND	1	500	0	522
RUTHS HOSPITALITY GROUP INC	COMMON STOCK	783332109	39	2,191	SH		OTR	1	0	2,191	0
RYANAIR HOLDINGS PLC SP A D R	COMMON STOCK	783513203	116	1,055	SH		DFND	1	1,055	0	0
RYANAIR HOLDINGS PLC SP A D R	COMMON STOCK	783513203	159	1,444	SH		OTR	1	179	0	1,265
RYANAIR HOLDINGS PLC	COMMON STOCK	783513203	2	18	SH		DFND	5	18	0	0
RYDER SYSTEM INC	COMMON STOCK	783549108	185	3,003	SH		DFND	1	1,532	0	1,471
RYDER SYSTEM INC	COMMON STOCK	783549108	90	1,450	SH		OTR	1	1,450	0	0
S & T BANCORP INC	COMMON STOCK	783859101	17	680	SH		DFND	1	50	0	630
S E I INVESTMENTS CO	COMMON STOCK	784117103	116	2,021	SH		DFND	1	1,369	0	652
S E I INVESTMENTS CO	COMMON STOCK	784117103	3	54	SH		OTR	1	0	54	0
SEI INVESTMENTS COMPANY	COMMON STOCK	784117103	35	613	SH		DFND	5	613	0	0
SJW GROUP	COMMON STOCK	784305104	13	183	SH		DFND	1	78	0	105
SJW GROUP	COMMON STOCK	784305104	14	200	SH		OTR	1	200	0	0
S P X CORP	COMMON STOCK	784635104	39	708	SH		DFND	1	708	0	0
S P X CORP	COMMON STOCK	784635104	437	8,019	SH		OTR	1	5,580	2,439	0
SSR MINING INC	COMMON STOCK	784730103	17	852	SH		DFND	1	176	0	676
SSR MINING INC	COMMON STOCK	784730103	5	245	SH		OTR	1	245	0	0
SABINE RTY TR UBI	COMMON STOCK	785688102	28	1,000	SH		OTR	1	1,000	0	0
SAGA COMMUNICATIONS INC	COMMON STOCK	786598300	2	100	SH		DFND	1	100	0	0
THE ST JOE COMPANY	COMMON STOCK	790148100	101	2,378	SH		DFND	1	2,378	0	0
SALEM MEDIA GROUP INC	COMMON STOCK	794093104	0	100	SH		DFND	1	100	0	0
SANDERSON FARMS INC	COMMON STOCK	800013104	738	5,581	SH		DFND	1	4,656	0	925
SANDERSON FARMS INC	COMMON STOCK	800013104	114	862	SH		OTR	1	862	0	0
SANDY SPRING BANCORP INC	COMMON STOCK	800363103	404	12,559	SH		DFND	1	12,392	0	167
SANDY SPRING BANCORP INC	COMMON STOCK	800363103	78	2,438	SH		OTR	1	2,438	0	0
SANFILIPPO B SANFILIPPO SON IN	COMMON STOCK	800422107	91	1,152	SH		DFND	1	1,126	0	26
SANFILIPPO B SANFILIPPO SON IN	COMMON STOCK	800422107	17	212	SH		OTR	1	0	212	0
SANGAMO THERAPEUTICS, INC.	COMMON STOCK	800677106	78	5,000	SH		DFND	1	5,000	0	0
SANMINA CORP	COMMON STOCK	801056102	15	459	SH		DFND	1	418	0	41
SAP SE SPON ADR A D R	COMMON STOCK	803054204	16,040	123,017	SH		DFND	1	118,777	0	4,240
SAP SE SPON ADR A D R	COMMON STOCK	803054204	3,719	28,521	SH		OTR	1	27,070	561	890
SAP AG	COMMON STOCK	803054204	67	514	SH		DFND	5	514	0	0
SAREPTA THERAPEUTICS INC	COMMON STOCK	803607100	426	2,496	SH		DFND	1	2,338	0	158
SAREPTA THERAPEUTICS INC	COMMON STOCK	803607100	464	2,724	SH		OTR	1	2,611	79	34
SASOL LTD SPON A D R	COMMON STOCK	803866300	779	87,881	SH		DFND	1	82,008	0	5,873
SASOL LTD SPON A D R	COMMON STOCK	803866300	161	18,162	SH		OTR	1	18,070	0	92
SASOL LTD	COMMON STOCK	803866300	3	329	SH		DFND	5	329	0	0
SAUL CTRS INC	COMMON STOCK	804395101	9	284	SH		DFND	1	231	0	53
SCHEIN HENRY INC	COMMON STOCK	806407102	2,278	34,078	SH		DFND	1	32,044	0	2,034
SCHEIN HENRY INC	COMMON STOCK	806407102	1,312	19,638	SH		OTR	1	18,371	1,217	50
SCHLUMBERGER LTD	COMMON STOCK	806857108	6,288	288,018	SH		DFND	1	274,924	0	13,094
SCHLUMBERGER LTD	COMMON STOCK	806857108	5,784	264,966	SH		OTR	1	244,998	11,562	8,406
SCHLUMBERGER LTD	COMMON STOCK	806857108	50	2,283	SH		DFND	5	2,283	0	0
SCHNITZER STEEL INDS INC A	COMMON STOCK	806882106	9	290	SH		DFND	1	256	0	34
SCHOLASTIC CORP	COMMON STOCK	807066105	24	967	SH		DFND	1	967	0	0
SCHWAB CHARLES CORP	COMMON STOCK	808513105	128,995	2,432,025	SH		DFND	1	2,353,015	0	79,010
SCHWAB CHARLES CORP	COMMON STOCK	808513105	18,017	339,701	SH		OTR	1	320,405	17,646	1,650
CHARLES SCHWAB CORPORATION	COMMON STOCK	808513105	72	1,366	SH		DFND	5	1,366	0	0
SCHWAB US BROAD MARKET ETF	OTHER	808524102	125	1,377	SH		DFND	1	1,377	0	0
SCHWAB US BROAD MARKET ETF	OTHER	808524102	27	300	SH		OTR	1	300	0	0
SCHWAB US BROAD MARKET	OTHER	808524102	1	13	SH		DFND	5	13	0	0
SCHWAB U S LARGE CAP ETF	OTHER	808524201	5,904	64,915	SH		DFND	1	58,593	0	6,322
SCHWAB US LARGE CAP	OTHER	808524201	5	51	SH		DFND	5	51	0	0
SCHWAB U S LARGE CAP GROWTH ET	OTHER	808524300	1,538	11,978	SH		DFND	1	7,788	0	4,190
SCHWAB U S LARGE CAP GROWTH ET	OTHER	808524300	51	400	SH		OTR	1	400	0	0
SCHWAB US LARGE CAP GROWTH	OTHER	808524300	1,492	11,618	SH		DFND	5	0	0	11,618
SCHWAB U S LARGE CAP VALUE ETF	OTHER	808524409	555	9,316	SH		DFND	1	764	0	8,552
SCHWAB US LARGE CAP VALUE	OTHER	808524409	8	140	SH		DFND	5	140	0	0
SCHWAB US MID CAP ETF	OTHER	808524508	330	4,836	SH		DFND	1	4,300	0	536
SCHWAB US MID CAP	OTHER	808524508	1	21	SH		DFND	5	21	0	0
SCHWAB US SMALL CAP ETF	OTHER	808524607	67	752	SH		DFND	1	348	0	404
SCHWAB US SMALL CAP	OTHER	808524607	16	182	SH		DFND	5	182	0	0
SCHWAB EMERGING MARKETS EQUITY	OTHER	808524706	30	969	SH		DFND	1	969	0	0
SCHWAB EMERGING MARKETS EQUITY	OTHER	808524706	33	1,066	SH		OTR	1	1,066	0	0
SCHWAB EMERGING MARKETS EQUITY	OTHER	808524706	2	52	SH		DFND	5	52	0	0
SCHWAB 1000 ETF	OTHER	808524722	839	22,432	SH		DFND	1	22,432	0	0
SCHWAB 1000 ETF	OTHER	808524722	21	557	SH		OTR	1	0	0	557
SCHWAB FUNDAMENTAL EMERGING ET	OTHER	808524730	76	2,685	SH		OTR	1	2,685	0	0
SCHWAB FUNDAMENTAL INTERNATION	OTHER	808524748	3	74	SH		DFND	1	74	0	0
SCHWAB FUNDAMENTAL INTERNATION	OTHER	808524755	6	208	SH		DFND	1	208	0	0
SCHWAB FUNDAMENTAL INTERNATION	OTHER	808524755	155	5,307	SH		OTR	1	5,307	0	0
SCHWAB FUNDAMENTAL INTL LARGE	OTHER	808524755	49	1,661	SH		DFND	5	0	0	1,661
SCHWAB FUNDAMENTAL U S SMALL	OTHER	808524763	116	2,694	SH		OTR	1	2,694	0	0
SCHWAB FUNDAMENTAL U S LARGE	OTHER	808524771	193	4,260	SH		OTR	1	4,260	0	0
SCHWAB U S DIVIDEND EQUITY ET	OTHER	808524797	575	8,961	SH		DFND	1	8,961	0	0
SCHWAB US DIVIDEND EQUITY	OTHER	808524797	164	2,554	SH		DFND	5	0	0	2,554
SCHWAB INTERNATIONAL EQUITY ET	OTHER	808524805	7,773	215,864	SH		DFND	1	207,004	0	8,860
SCHWAB INTL EQTY	OTHER	808524805	3	75	SH		DFND	5	75	0	0
SCHWAB U S AGGREGATE BOND ETF	OTHER	808524839	99	1,773	SH		DFND	1	1,773	0	0
SCHWAB US AGGREGATE BOND	OTHER	808524839	3	59	SH		DFND	5	59	0	0
SCHWAB US REIT ETF	OTHER	808524847	136,836	3,603,779	SH		DFND	1	3,577,812	0	25,967
SCHWAB US REIT ETF	OTHER	808524847	15,439	406,611	SH		OTR	1	399,128	3,352	4,131
SCHWAB US REIT	OTHER	808524847	74,296	1,956,622	SH		DFND	5	12,658	0	1,943,964
SCHWAB INTERMEDIATE TERM U S E	OTHER	808524854	258	4,429	SH		DFND	1	305	0	4,124
SCHWAB INTERMEDIATE TERM U S E	OTHER	808524854	148	2,534	SH		OTR	1	2,534	0	0
SCHWAB SHORT TERM US ETF	OTHER	808524862	123	2,392	SH		DFND	1	416	0	1,976
SCHWAB SHORT TERM US ETF	OTHER	808524862	77	1,500	SH		OTR	1	1,500	0	0
SCHWAB U S TIPS ETF	OTHER	808524870	145,911	2,350,382	SH		DFND	1	2,333,621	0	16,761
SCHWAB U S TIPS ETF	OTHER	808524870	19,194	309,177	SH		OTR	1	303,687	2,554	2,936
SCHWAB US TIPS	OTHER	808524870	340	5,480	SH		DFND	5	5,480	0	0
SCHWAB INTERNATIONAL SMALL CAP	OTHER	808524888	4,354	115,645	SH		DFND	1	114,860	0	785
SCHWAB INTERNATIONAL SMALL CAP	OTHER	808524888	2,192	58,216	SH		OTR	1	58,216	0	0
SCHWEITZER MAUDUIT INTL INC	COMMON STOCK	808541106	55	1,357	SH		DFND	1	903	0	454
SCIENCE APPLICATNS INTE	COMMON STOCK	808625107	34	362	SH		DFND	1	162	0	200
SCIENCE APPLICATNS INTE	COMMON STOCK	808625107	3	35	SH		OTR	1	35	0	0
SCIPLAY CORP	COMMON STOCK	809087109	0	9	SH		DFND	5	9	0	0
THE SCOTTS MIRACLE GRO COMPANY	COMMON STOCK	810186106	2,362	11,859	SH		DFND	1	11,156	0	703
THE SCOTTS MIRACLE GRO COMPANY	COMMON STOCK	810186106	702	3,522	SH		OTR	1	3,493	0	29
SCOTTS MIRACLE GRO CO	COMMON STOCK	810186106	41	205	SH		DFND	5	205	0	0
EW SCRIPPS CO CL A	COMMON STOCK	811054402	401	26,214	SH		DFND	1	26,214	0	0
SCULPTOR CAPITAL MANAGEMENT	COMMON STOCK	811246107	8	500	SH		DFND	1	500	0	0
SCYNEXIS INC	COMMON STOCK	811292200	24	3,100	SH		OTR	1	3,100	0	0
SEABOARD CORP DEL	COMMON STOCK	811543107	152	50	SH		DFND	1	35	0	15
SEABOARD CORP DEL	COMMON STOCK	811543107	9	3	SH		OTR	1	0	0	3
SEACOAST BANKING CORP FL	COMMON STOCK	811707801	1	20	SH		DFND	1	20	0	0
SEACOR HOLDINGS INC	COMMON STOCK	811904101	2	39	SH		DFND	1	39	0	0
SEABRIDGE GOLD INC	COMMON STOCK	811916105	2	76	SH		DFND	1	0	0	76
SEABRIDGE GOLD INC	COMMON STOCK	811916105	15	727	SH		OTR	1	727	0	0
SELECTIVE INS GROUP INC	COMMON STOCK	816300107	1,644	24,544	SH		DFND	1	22,780	0	1,764
SELECTIVE INS GROUP INC	COMMON STOCK	816300107	337	5,027	SH		OTR	1	5,027	0	0
SELECTIVE INSURANCE GROUP	COMMON STOCK	816300107	15	229	SH		DFND	5	229	0	0
SELECTQUOTE INC COM	COMMON STOCK	816307300	658	31,724	SH		DFND	1	31,724	0	0
SEMTECH CORP	COMMON STOCK	816850101	1,142	15,836	SH		DFND	1	15,768	0	68
SEMTECH CORP	COMMON STOCK	816850101	230	3,189	SH		OTR	1	1,327	88	1,774
SEMPRA ENERGY	COMMON STOCK	816851109	5,336	41,885	SH		DFND	1	34,786	0	7,099
SEMPRA ENERGY	COMMON STOCK	816851109	2,969	23,302	SH		OTR	1	20,876	1,203	1,223
SEMPRA ENERGY	COMMON STOCK	816851109	4	31	SH		DFND	5	16	0	15
SEQUANS COMMUNICATIONS S A A D	COMMON STOCK	817323207	3	500	SH		DFND	1	500	0	0
SERVICE CORP INTL	COMMON STOCK	817565104	118	2,409	SH		DFND	1	1,832	0	577
SERVICE CORP INTL	COMMON STOCK	817565104	2	50	SH		OTR	1	50	0	0
SERVICE CORP INTERNATIONAL	COMMON STOCK	817565104	2	33	SH		DFND	5	33	0	0
SHAKE SHACK INC CLASS A	COMMON STOCK	819047101	723	8,529	SH		DFND	1	8,518	0	11
SHARPS COMPLIANCE CORP	COMMON STOCK	820017101	5	500	SH		DFND	1	500	0	0
SHELL MIDSTREAM PARTNERS L P	COMMON STOCK	822634101	525	52,067	SH		DFND	1	52,067	0	0
SHELL MIDSTREAM PARTNERS LP	COMMON STOCK	822634101	1	78	SH		DFND	5	78	0	0
SHERWIN WILLIAMS CO	COMMON STOCK	824348106	34,241	46,592	SH		DFND	1	45,886	0	706
SHERWIN WILLIAMS CO	COMMON STOCK	824348106	7,282	9,909	SH		OTR	1	9,650	132	127
SHERWIN-WILLIAMS CO	COMMON STOCK	824348106	981	1,332	SH		DFND	5	1,314	0	18
SHINHAN FINANCIAL GROUP A D R	COMMON STOCK	824596100	635	21,334	SH		DFND	1	11,603	0	9,731
SHINHAN FINANCIAL GROUP A D R	COMMON STOCK	824596100	18	584	SH		OTR	1	0	30	554
SHOE CARNIVAL INC	COMMON STOCK	824889109	24	624	SH		OTR	1	0	624	0
SHOE CARNIVAL INC	COMMON STOCK	824889109	7	182	SH		DFND	5	182	0	0
SHUTTERSTOCK INC	COMMON STOCK	825690100	12	172	SH		DFND	1	172	0	0
SHUTTERSTOCK INC	COMMON STOCK	825690100	6	81	SH		DFND	5	81	0	0
SI BONE INC	COMMON STOCK	825704109	10	340	SH		DFND	1	272	0	68
SIERRA WIRELESS INC	COMMON STOCK	826516106	6	381	SH		DFND	1	321	0	60
SILICON LABORATORIES INC	COMMON STOCK	826919102	1,797	14,110	SH		DFND	1	14,085	0	25
SILICON LABORATORIES INC	COMMON STOCK	826919102	57	454	SH		OTR	1	435	19	0
SILICON LABORATORIES	COMMON STOCK	826919102	51	402	SH		DFND	5	402	0	0
SILGAN HLDGS INC	COMMON STOCK	827048109	321	8,670	SH		DFND	1	6,734	0	1,936
SILGAN HLDGS INC	COMMON STOCK	827048109	10	265	SH		OTR	1	265	0	0
SIMMONS 1ST NATIONAL CORP CL A	COMMON STOCK	828730200	186	8,609	SH		DFND	1	8,609	0	0
SIMMONS 1ST NATIONAL CORP CL A	COMMON STOCK	828730200	4	174	SH		OTR	1	174	0	0
SIMON PROPERTY GROUP INC	COMMON STOCK	828806109	2,506	29,393	SH		DFND	1	25,780	0	3,613
SIMON PROPERTY GROUP INC	COMMON STOCK	828806109	845	9,896	SH		OTR	1	9,273	321	302
SIMON PROPERTY GROUP INC	COMMON STOCK	828806109	14	161	SH		DFND	5	144	0	17
SIMON PROPERTY GROUP INC	COMMON STOCK	828806109	11	26	SH	Put	DFND	1	26	0	0
SIMPSON MFG CO INC	COMMON STOCK	829073105	672	7,193	SH		DFND	1	6,870	0	323
SIMPSON MFG CO INC	COMMON STOCK	829073105	247	2,640	SH		OTR	1	505	0	2,135
SIMULATIONS PLUS INC	COMMON STOCK	829214105	67	925	SH		DFND	1	635	0	290
SIMULATIONS PLUS INC	COMMON STOCK	829214105	28	392	SH		OTR	1	20	0	372
SINCLAIR BROADCAST GROUP INC A	COMMON STOCK	829226109	142	4,442	SH		DFND	1	2,880	0	1,562
SINCLAIR BROADCAST GROUP INC A	COMMON STOCK	829226109	27	862	SH		OTR	1	77	708	77
SKECHERS U S A INC	COMMON STOCK	830566105	91	2,522	SH		DFND	1	2,215	0	307
SKYLINE CHAMPION CORP	COMMON STOCK	830830105	6	192	SH		DFND	1	83	0	109
SKYWEST INC	COMMON STOCK	830879102	114	2,826	SH		DFND	1	2,060	0	766
SKYWEST INC	COMMON STOCK	830879102	5	122	SH		OTR	1	0	122	0
SMITH AND WESSON BRANDS INC	COMMON STOCK	831754106	20	1,115	SH		DFND	1	1,115	0	0
SMITH AND WESSON BRANDS INC	COMMON STOCK	831754106	4	218	SH		OTR	1	218	0	0
SMITH A O CORP	COMMON STOCK	831865209	821	14,969	SH		DFND	1	14,361	0	608
SMITH A O CORP	COMMON STOCK	831865209	1,283	23,412	SH		OTR	1	20,157	18	3,237
SMITH (A.O.) CORP	COMMON STOCK	831865209	176	3,213	SH		DFND	5	3,213	0	0
JM SMUCKER CO THE	COMMON STOCK	832696405	4,048	35,020	SH		DFND	1	33,481	0	1,539
JM SMUCKER CO THE	COMMON STOCK	832696405	2,390	20,677	SH		OTR	1	15,103	2,850	2,724
JM SMUCKER CO	COMMON STOCK	832696405	6	56	SH		DFND	5	56	0	0
SNAP ON INC	COMMON STOCK	833034101	3,764	21,993	SH		DFND	1	21,432	0	561
SNAP ON INC	COMMON STOCK	833034101	1,713	10,010	SH		OTR	1	8,276	671	1,063
SNAP-ON INC	COMMON STOCK	833034101	177	1,035	SH		DFND	5	1,035	0	0
SNOWFLAKE INC CL A	COMMON STOCK	833445109	163	579	SH		DFND	1	579	0	0
SNOWFLAKE INC	COMMON STOCK	833445109	3	10	SH		DFND	5	10	0	0
SOCIEDAD QUIMICA MINERA A D R	COMMON STOCK	833635105	646	13,154	SH		DFND	1	11,672	0	1,482
SOCIEDAD QUIMICA MINERA A D R	COMMON STOCK	833635105	44	899	SH		OTR	1	899	0	0
QUIMICA Y MINERA DE CHILE SA	COMMON STOCK	833635105	6	124	SH		DFND	5	109	0	15
SONOCO PRODS CO	COMMON STOCK	835495102	1,934	32,652	SH		DFND	1	31,777	0	875
SONOCO PRODS CO	COMMON STOCK	835495102	372	6,281	SH		OTR	1	6,281	0	0
SONOCO PRODUCTS CO	COMMON STOCK	835495102	29	491	SH		DFND	5	491	0	0
SONY CORP A D R	COMMON STOCK	835699307	7,011	69,350	SH		DFND	1	61,762	0	7,588
SONY CORP A D R	COMMON STOCK	835699307	2,278	22,538	SH		OTR	1	20,562	852	1,124
SOURCE CAPITAL INC	OTHER	836144105	29	716	SH		DFND	1	716	0	0
SOUTH JERSEY INDUSTRIES	COMMON STOCK	838518108	1,045	48,490	SH		DFND	1	47,883	0	607
SOUTH JERSEY INDUSTRIES	COMMON STOCK	838518108	30	1,410	SH		OTR	1	1,410	0	0
SOUTH STATE CORP	COMMON STOCK	840441109	867	11,993	SH		DFND	1	11,993	0	0
SOUTH STATE CORP	COMMON STOCK	840441109	224	3,095	SH		OTR	1	2,665	430	0
SOUTH STATE CORP	COMMON STOCK	840441109	11	159	SH		DFND	5	159	0	0
SOUTHERN CO THE	COMMON STOCK	842587107	13,723	223,386	SH		DFND	1	211,744	0	11,642
SOUTHERN CO THE	COMMON STOCK	842587107	7,634	124,268	SH		OTR	1	100,758	2,790	20,720
SOUTHERN CO	COMMON STOCK	842587107	327	5,319	SH		DFND	5	5,273	0	46
SOUTHERN MO BANCORP INC COM	COMMON STOCK	843380106	122	4,000	SH		DFND	1	4,000	0	0
SOUTHWEST AIRLINES CO	COMMON STOCK	844741108	6,131	131,556	SH		DFND	1	127,388	0	4,168
SOUTHWEST AIRLINES CO	COMMON STOCK	844741108	1,654	35,472	SH		OTR	1	35,051	413	8
SOUTHWEST AIRLINES CO	COMMON STOCK	844741108	23	501	SH		DFND	5	501	0	0
SOUTHWEST GAS HOLDINGS INC	COMMON STOCK	844895102	809	13,313	SH		DFND	1	12,741	0	572
SOUTHWEST GAS HOLDINGS INC	COMMON STOCK	844895102	141	2,319	SH		OTR	1	2,319	0	0
SOUTHWESTERN ENERGY CO	COMMON STOCK	845467109	39	13,122	SH		DFND	1	7,971	0	5,151
SOUTHWESTERN ENERGY CO	COMMON STOCK	845467109	6	1,952	SH		OTR	1	1,952	0	0
SPARTANNASH CO	COMMON STOCK	847215100	77	4,430	SH		DFND	1	3,186	0	1,244
SPARTANNASH CO	COMMON STOCK	847215100	8	440	SH		OTR	1	0	440	0
SPIRIT AEROSYSTEMS HOLD CL A	COMMON STOCK	848574109	192	4,915	SH		DFND	1	4,298	0	617
SPIRIT AEROSYSTEMS HOLD CL A	COMMON STOCK	848574109	5	124	SH		OTR	1	0	32	92
SPIRIT AIRLINES INC	COMMON STOCK	848577102	6	256	SH		DFND	1	256	0	0
SPIRIT AIRLINES INC	COMMON STOCK	848577102	2	80	SH		OTR	1	0	80	0
SPLUNK INC	COMMON STOCK	848637104	39,133	230,342	SH		DFND	1	227,324	0	3,018
SPLUNK INC	COMMON STOCK	848637104	7,167	42,186	SH		OTR	1	37,521	4,151	514
SPLUNK INC	COMMON STOCK	848637104	453	2,664	SH		DFND	5	2,664	0	0
SPROTT INC COM NEW	COMMON STOCK	852066208	0	14	SH		DFND	1	14	0	0
SQUARE INC A	COMMON STOCK	852234103	37,540	172,485	SH		DFND	1	169,037	0	3,448
SQUARE INC A	COMMON STOCK	852234103	8,090	37,172	SH		OTR	1	36,373	274	525
SQUARE INC	COMMON STOCK	852234103	49	225	SH		DFND	5	225	0	0
STAAR SURGICAL CO	COMMON STOCK	852312305	255	3,224	SH		DFND	1	18	0	3,206
STAMPS COM INC	COMMON STOCK	852857200	188	956	SH		DFND	1	912	0	44
STANDARD MTR PRODS INC	COMMON STOCK	853666105	196	4,842	SH		DFND	1	4,842	0	0
STANDARD MTR PRODS INC	COMMON STOCK	853666105	70	1,734	SH		OTR	1	1,297	437	0
STANDARD MOTOR PRODS INC	COMMON STOCK	853666105	9	225	SH		DFND	5	225	0	0
STANDEX INTL CORP	COMMON STOCK	854231107	129	1,661	SH		DFND	1	279	0	1,382
STANDEX INTERNATIONAL CORP	COMMON STOCK	854231107	12	153	SH		DFND	5	153	0	0
STANLEY BLACK DECKER INC	COMMON STOCK	854502101	4,763	26,675	SH		DFND	1	25,849	0	826
STANLEY BLACK DECKER INC	COMMON STOCK	854502101	2,568	14,380	SH		OTR	1	11,427	2,936	17
STANLEY BLACK & DECKER INC	COMMON STOCK	854502101	22	122	SH		DFND	5	122	0	0
STARBUCKS CORP	COMMON STOCK	855244109	88,883	830,835	SH		DFND	1	783,601	0	47,234
STARBUCKS CORP	COMMON STOCK	855244109	18,638	174,225	SH		OTR	1	143,522	7,653	23,050
STARBUCKS CORP	COMMON STOCK	855244109	450	4,205	SH		DFND	5	3,163	0	1,042
STARRETT L S CO	COMMON STOCK	855668109	2	357	SH		DFND	1	357	0	0
STATE AUTO FINL CORP	COMMON STOCK	855707105	23	1,282	SH		DFND	1	1,282	0	0
STATE STR CORP	COMMON STOCK	857477103	6,572	90,303	SH		DFND	1	65,922	0	24,381
STATE STR CORP	COMMON STOCK	857477103	3,508	48,200	SH		OTR	1	41,638	218	6,344
STATE STREET CORP	COMMON STOCK	857477103	10	134	SH		DFND	5	134	0	0
STEEL DYNAMICS INC	COMMON STOCK	858119100	543	14,702	SH		DFND	1	13,380	0	1,322
STEEL DYNAMICS INC	COMMON STOCK	858119100	55	1,483	SH		OTR	1	1,470	13	0
STEEL DYNAMICS INC	COMMON STOCK	858119100	17	461	SH		DFND	5	461	0	0
STEELCASE INC CL A	COMMON STOCK	858155203	56	4,129	SH		DFND	1	3,900	0	229
STELLUS CAP INVT CORP	COMMON STOCK	858568108	8	700	SH		OTR	1	700	0	0
STEPAN CO	COMMON STOCK	858586100	349	2,930	SH		DFND	1	2,930	0	0
STERICYCLE INC	COMMON STOCK	858912108	1,459	21,043	SH		DFND	1	20,710	0	333
STERICYCLE INC	COMMON STOCK	858912108	554	7,993	SH		OTR	1	7,793	0	200
STERICYCLE INC	COMMON STOCK	858912108	32	460	SH		DFND	5	460	0	0
STERLING CONSTRUCTION CO	COMMON STOCK	859241101	1	50	SH		DFND	1	0	0	50
STEWART INFORMATION SVCS CORP	COMMON STOCK	860372101	96	1,978	SH		DFND	1	1,647	0	331
STEWART INFORMATION SVCS CORP	COMMON STOCK	860372101	107	2,213	SH		OTR	1	1,016	1,197	0
STEWART INFORMATION SERVICES	COMMON STOCK	860372101	7	143	SH		DFND	5	143	0	0
STIFEL FINL CORP	COMMON STOCK	860630102	4,890	96,901	SH		DFND	1	95,401	0	1,500
STIFEL FINL CORP	COMMON STOCK	860630102	571	11,309	SH		OTR	1	11,309	0	0
STIFEL FINANCIAL CORP	COMMON STOCK	860630102	49	961	SH		DFND	5	961	0	0
STITCH FIX INC CLASS A	COMMON STOCK	860897107	8	143	SH		DFND	1	100	0	43
STITCH FIX INC	COMMON STOCK	860897107	0	2	SH		DFND	5	2	0	0
STMICROELECTRONICS N V	COMMON STOCK	861012102	4,262	114,804	SH		DFND	1	106,788	0	8,016
STMICROELECTRONICS N V	COMMON STOCK	861012102	1,007	27,129	SH		OTR	1	26,954	103	72
STMICROELECTRONCS NV	COMMON STOCK	861012102	5	128	SH		DFND	5	128	0	0
STOCK YARDS BANCORP INC	COMMON STOCK	861025104	91	2,240	SH		DFND	1	2,240	0	0
STONEX GROUP INC COM	COMMON STOCK	861896108	6	98	SH		DFND	1	98	0	0
STORE CAPITAL CORP	COMMON STOCK	862121100	745	21,937	SH		DFND	1	20,077	0	1,860
STORE CAPITAL CORP	COMMON STOCK	862121100	145	4,265	SH		OTR	1	1,683	2,398	184
STORE CAPITAL CORP	COMMON STOCK	862121100	4	128	SH		DFND	5	128	0	0
STRYKER CORP	COMMON STOCK	863667101	36,140	147,488	SH		DFND	1	136,005	0	11,483
STRYKER CORP	COMMON STOCK	863667101	10,403	42,451	SH		OTR	1	33,885	906	7,660
STRYKER CORP	COMMON STOCK	863667101	832	3,389	SH		DFND	5	3,389	0	0
STURM RUGER & CO INC	COMMON STOCK	864159108	199	3,061	SH		DFND	1	3,061	0	0
STURM RUGER & CO INC	COMMON STOCK	864159108	3	40	SH		OTR	1	40	0	0
SUBURBAN PROPANE PARTNERS LP	COMMON STOCK	864482104	4	300	SH		OTR	1	300	0	0
SUMMIT HOTEL PROPERTIES INC	COMMON STOCK	866082100	473	52,493	SH		DFND	1	52,147	0	346
SUMMIT HOTEL PROPERTIES INC	COMMON STOCK	866082100	30	3,301	SH		DFND	5	3,301	0	0
SUN COMMUNITIES INC	COMMON STOCK	866674104	1,839	12,103	SH		DFND	1	11,539	0	564
SUN COMMUNITIES INC	COMMON STOCK	866674104	527	3,473	SH		OTR	1	3,287	30	156
SUN COMMUNITIES INC	COMMON STOCK	866674104	86	565	SH		DFND	5	554	0	11
SUN LIFE FINANCIAL INC	COMMON STOCK	866796105	1,354	30,447	SH		DFND	1	24,889	0	5,558
SUN LIFE FINANCIAL INC	COMMON STOCK	866796105	73	1,655	SH		OTR	1	1,581	27	47
SUNCOR ENERGY INC	COMMON STOCK	867224107	696	41,474	SH		DFND	1	39,116	0	2,358
SUNCOR ENERGY INC	COMMON STOCK	867224107	277	16,512	SH		OTR	1	13,906	1,253	1,353
SUNPOWER CORP	COMMON STOCK	867652406	884	34,467	SH		DFND	1	34,414	0	53
SUNPOWER CORP	COMMON STOCK	867652406	50	1,946	SH		OTR	1	1,946	0	0
SUNSTONE HOTEL INVS INC	COMMON STOCK	867892101	16	1,452	SH		DFND	1	1,077	0	375
SUPERNUS PHARMACEUTICALS INC	COMMON STOCK	868459108	280	11,122	SH		DFND	1	11,122	0	0
SUPERNUS PHARMACEUTICALS INC	COMMON STOCK	868459108	4	160	SH		OTR	1	160	0	0
SURMODICS INC	COMMON STOCK	868873100	422	9,698	SH		DFND	1	9,698	0	0
SURMODICS INC	COMMON STOCK	868873100	8	190	SH		OTR	1	0	190	0
SWISS HELVETIA FD INC	OTHER	870875101	26	2,911	SH		DFND	1	2,911	0	0
SYKES ENTERPRISES INC	COMMON STOCK	871237103	185	4,914	SH		DFND	1	4,607	0	307
SYKES ENTERPRISES INC	COMMON STOCK	871237103	53	1,429	SH		OTR	1	1,310	0	119
SYKES ENTERPRISES INC	COMMON STOCK	871237103	10	261	SH		DFND	5	261	0	0
SYNALLOY CORP	COMMON STOCK	871565107	6	825	SH		DFND	1	825	0	0
SYNOPSYS INC	COMMON STOCK	871607107	12,501	48,223	SH		DFND	1	44,504	0	3,719
SYNOPSYS INC	COMMON STOCK	871607107	2,250	8,682	SH		OTR	1	8,202	160	320
SYNOPSYS INC	COMMON STOCK	871607107	590	2,275	SH		DFND	5	2,264	0	11
SYSCO CORP	COMMON STOCK	871829107	11,001	148,135	SH		DFND	1	137,960	0	10,175
SYSCO CORP	COMMON STOCK	871829107	10,596	142,687	SH		OTR	1	139,517	1,392	1,778
SYSCO CORP	COMMON STOCK	871829107	41	546	SH		DFND	5	546	0	0
TCG BDC INC	COMMON STOCK	872280102	1,150	112,062	SH		DFND	1	112,062	0	0
TCF FINANCIAL CORP	COMMON STOCK	872307103	1,037	28,008	SH		DFND	1	25,445	0	2,563
TCF FINANCIAL CORP	COMMON STOCK	872307103	902	24,368	SH		OTR	1	23,306	131	931
TCF FINANCIAL CORP	COMMON STOCK	872307103	43	1,154	SH		DFND	5	1,154	0	0
TCW STRATEGIC INCOME FUND INC	OTHER	872340104	31	5,525	SH		DFND	5	5,525	0	0
TJX COMPANIES INC	COMMON STOCK	872540109	20,329	297,676	SH		DFND	1	240,270	0	57,406
TJX COMPANIES INC	COMMON STOCK	872540109	8,276	121,178	SH		OTR	1	95,300	18,565	7,313
TJX COMPANIES	COMMON STOCK	872540109	108	1,577	SH		DFND	5	1,577	0	0
T MOBILE US INC	COMMON STOCK	872590104	1,606	11,908	SH		DFND	1	8,648	0	3,260
T MOBILE US INC	COMMON STOCK	872590104	853	6,331	SH		OTR	1	3,426	1,537	1,368
T-MOBILE US INC	COMMON STOCK	872590104	94	700	SH		DFND	5	700	0	0
TABULA RASA HEALTHCARE INC	COMMON STOCK	873379101	120	2,798	SH		DFND	1	0	0	2,798
TAIWAN SEMICONDUCTOR A D R	COMMON STOCK	874039100	46,189	423,601	SH		DFND	1	391,568	0	32,033
TAIWAN SEMICONDUCTOR A D R	COMMON STOCK	874039100	7,731	70,900	SH		OTR	1	66,594	1,455	2,851
TAIWAN SEMICONDUCTOR	COMMON STOCK	874039100	1,858	17,031	SH		DFND	5	17,013	0	18
TAKE-TWO INTERACTIVE SOFTWARE	COMMON STOCK	874054109	4,164	20,042	SH		DFND	1	19,161	0	881
TAKE-TWO INTERACTIVE SOFTWARE	COMMON STOCK	874054109	908	4,370	SH		OTR	1	4,190	49	131
TAKE-TWO INTERACTIVE SOFTWARE	COMMON STOCK	874054109	90	431	SH		DFND	5	402	0	29
TAKEDA PHARMACEUTICAL SP A D R	COMMON STOCK	874060205	716	39,374	SH		DFND	1	19,758	0	19,616
TAKEDA PHARMACEUTICAL SP A D R	COMMON STOCK	874060205	143	7,833	SH		OTR	1	4,866	1,836	1,131
TAL EDUCATION GROUP A D R	COMMON STOCK	874080104	1,191	16,648	SH		DFND	1	14,121	0	2,527
TAL EDUCATION GROUP A D R	COMMON STOCK	874080104	110	1,532	SH		OTR	1	1,505	0	27
TALEND SA	COMMON STOCK	874224207	2	46	SH		DFND	5	46	0	0
TANDEM DIABETES CARE INC	COMMON STOCK	875372203	74	773	SH		DFND	1	773	0	0
TANDEM DIABETES CARE INC	COMMON STOCK	875372203	1	10	SH		OTR	1	10	0	0
TANDEM DIABETES CARE INC	COMMON STOCK	875372203	1	12	SH		DFND	5	0	0	12
TANGER FACTORY OUTLET CENTER	COMMON STOCK	875465106	59	5,965	SH		DFND	1	5,460	0	505
TANGER FACTORY OUTLET CENTER	COMMON STOCK	875465106	19	1,910	SH		OTR	1	1,910	0	0
TAPESTRY INC	COMMON STOCK	876030107	431	13,859	SH		DFND	1	13,015	0	844
TAPESTRY INC	COMMON STOCK	876030107	219	7,062	SH		OTR	1	6,909	0	153
TAPESTRY INC	COMMON STOCK	876030107	27	882	SH		DFND	5	882	0	0
TASEKO MINES LTD	COMMON STOCK	876511106	13	10,000	SH		OTR	1	10,000	0	0
TATA MOTORS LTD A D R	COMMON STOCK	876568502	353	27,971	SH		DFND	1	16,264	0	11,707
TATA MOTORS LTD A D R	COMMON STOCK	876568502	2	184	SH		OTR	1	0	0	184
TEAM INC	COMMON STOCK	878155100	80	7,301	SH		DFND	1	0	0	7,301
TECK RESOURCES LIMITED	COMMON STOCK	878742204	280	15,402	SH		DFND	1	11,012	0	4,390
TECK RESOURCES LIMITED	COMMON STOCK	878742204	236	13,000	SH		OTR	1	13,000	0	0
TEJON RANCH CO	COMMON STOCK	879080109	12	828	SH		DFND	1	748	0	80
TELEDYNE TECHNOLOGIES INC	COMMON STOCK	879360105	1,983	5,060	SH		DFND	1	4,821	0	239
TELEDYNE TECHNOLOGIES INC	COMMON STOCK	879360105	635	1,619	SH		OTR	1	1,612	0	7
TELEDYNE TECHNOLOGIES INC	COMMON STOCK	879360105	27	68	SH		DFND	5	63	0	5
TELEFLEX INC	COMMON STOCK	879369106	2,681	6,512	SH		DFND	1	4,830	0	1,682
TELEFLEX INC	COMMON STOCK	879369106	1,459	3,545	SH		OTR	1	2,927	567	51
TELEFONICA SA SPON A D R	COMMON STOCK	879382208	446	110,384	SH		DFND	1	52,047	0	58,337
TELEFONICA SA SPON A D R	COMMON STOCK	879382208	19	4,753	SH		OTR	1	3,458	545	750
TELEPHONE AND DATA SYSTEMS INC	COMMON STOCK	879433829	108	5,802	SH		DFND	1	4,914	0	888
TELEPHONE AND DATA SYSTEMS INC	COMMON STOCK	879433829	6	331	SH		OTR	1	57	150	124
TENNANT CO	COMMON STOCK	880345103	2,120	30,216	SH		DFND	1	29,716	0	500
TENNANT CO	COMMON STOCK	880345103	412	5,872	SH		OTR	1	5,622	0	250
TENNECO INC	COMMON STOCK	880349105	10	899	SH		DFND	1	50	0	849
TERADYNE INC	COMMON STOCK	880770102	2,088	17,416	SH		DFND	1	14,963	0	2,453
TERADYNE INC	COMMON STOCK	880770102	916	7,639	SH		OTR	1	5,240	2,399	0
TERADYNE INC	COMMON STOCK	880770102	2	14	SH		DFND	5	14	0	0
TEREX CORP	COMMON STOCK	880779103	244	7,005	SH		DFND	1	6,057	0	948
TEREX CORP	COMMON STOCK	880779103	3	80	SH		OTR	1	0	0	80
TERNIUM SA SPONSORED A D R	COMMON STOCK	880890108	14	491	SH		DFND	1	491	0	0
TEVA PHARMACEUTICAL INDS LTD A	COMMON STOCK	881624209	75	7,731	SH		DFND	1	6,372	0	1,359
TEVA PHARMACEUTICAL INDS LTD A	COMMON STOCK	881624209	80	8,285	SH		OTR	1	5,819	1,766	700
TEXAS INSTRUMENTS INC	COMMON STOCK	882508104	107,509	655,029	SH		DFND	1	633,862	0	21,167
TEXAS INSTRUMENTS INC	COMMON STOCK	882508104	40,616	247,462	SH		OTR	1	232,637	5,197	9,628
TEXAS INSTRUMENTS INC	COMMON STOCK	882508104	824	5,013	SH		DFND	5	5,013	0	0
TEXAS INSTRUMENTS INC	COMMON STOCK	882508104	5	16	SH	Put	DFND	1	16	0	0
TEXAS INSTRUMENTS INC	COMMON STOCK	882508104	3	46	SH	Put	OTR	1	46	0	0
TEXAS PAC LD TR	COMMON STOCK	882610108	7,998	11,001	SH		DFND	1	10,976	0	25
TEXAS PACIFIC LAND TRUST	COMMON STOCK	882610108	19	26	SH		DFND	5	26	0	0
TEXAS ROADHOUSE INC	COMMON STOCK	882681109	1,390	17,788	SH		DFND	1	14,520	0	3,268
TEXAS ROADHOUSE INC	COMMON STOCK	882681109	93	1,194	SH		OTR	1	612	582	0
TEXAS ROADHOUSE INC	COMMON STOCK	882681109	53	675	SH		DFND	5	675	0	0
TEXTRON INC	COMMON STOCK	883203101	570	11,789	SH		DFND	1	9,985	0	1,804
TEXTRON INC	COMMON STOCK	883203101	865	17,889	SH		OTR	1	17,384	273	232
TEXTRON INC	COMMON STOCK	883203101	33	686	SH		DFND	5	686	0	0
THERMO FISHER SCIENTIFIC INC	COMMON STOCK	883556102	58,446	125,478	SH		DFND	1	119,723	0	5,755
THERMO FISHER SCIENTIFIC INC	COMMON STOCK	883556102	38,130	81,862	SH		OTR	1	76,500	3,695	1,667
THERMO FISHER SCIENTIFIC INC	COMMON STOCK	883556102	1,532	3,287	SH		DFND	5	3,255	0	32
THOMSON REUTERS CORP	COMMON STOCK	884903709	1,588	19,390	SH		DFND	1	16,343	0	3,047
THOMSON REUTERS CORP	COMMON STOCK	884903709	258	3,156	SH		OTR	1	3,124	0	32
THOMSON REUTERS CORP	COMMON STOCK	884903709	13	160	SH		DFND	5	138	0	22
THOR INDUSTRIES INC	COMMON STOCK	885160101	433	4,657	SH		DFND	1	1,057	0	3,600
THOR INDUSTRIES INC	COMMON STOCK	885160101	9	100	SH		OTR	1	0	50	50
TIFFANY & CO	COMMON STOCK	886547108	391	2,975	SH		DFND	1	2,107	0	868
TIFFANY & CO	COMMON STOCK	886547108	93	705	SH		OTR	1	669	0	36
TIMKEN CO	COMMON STOCK	887389104	789	10,201	SH		DFND	1	9,291	0	910
TIMKEN CO	COMMON STOCK	887389104	397	5,127	SH		OTR	1	194	0	4,933
TOLL BROS INC	COMMON STOCK	889478103	310	7,122	SH		DFND	1	5,417	0	1,705
TOLL BROS INC	COMMON STOCK	889478103	13	306	SH		OTR	1	192	57	57
TOMPKINS FINANCIAL CORPORATION	COMMON STOCK	890110109	1	10	SH		DFND	1	0	0	10
TOMPKINS FINANCIAL CORPORATION	COMMON STOCK	890110109	131	1,858	SH		OTR	1	1,858	0	0
TONIX PHARMACEUTICALS	COMMON STOCK	890260706	0	2	SH		DFND	1	2	0	0
TOOTSIE ROLL INDS	COMMON STOCK	890516107	362	12,194	SH		DFND	1	11,141	0	1,053
TOOTSIE ROLL INDS	COMMON STOCK	890516107	99	3,296	SH		OTR	1	1,453	1	1,842
TORO CO	COMMON STOCK	891092108	89,148	939,984	SH		DFND	1	91,754	0	848,230
TORO CO	COMMON STOCK	891092108	4,067	42,877	SH		OTR	1	42,677	0	200
TORO CO (THE)	COMMON STOCK	891092108	69	725	SH		DFND	5	725	0	0
TORONTO DOMINION BANK	COMMON STOCK	891160509	1,076	19,070	SH		DFND	1	15,063	0	4,007
TORONTO DOMINION BANK	COMMON STOCK	891160509	60	1,072	SH		OTR	1	780	0	292
TOWNSQUARE MEDIA INC	COMMON STOCK	892231101	1	100	SH		DFND	1	100	0	0
TOYOTA MTR CORP A D R	COMMON STOCK	892331307	4,818	31,175	SH		DFND	1	25,795	0	5,380
TOYOTA MTR CORP A D R	COMMON STOCK	892331307	998	6,455	SH		OTR	1	5,273	289	893
TRACTOR SUPPLY COMPANY	COMMON STOCK	892356106	6,534	46,481	SH		DFND	1	42,196	0	4,285
TRACTOR SUPPLY COMPANY	COMMON STOCK	892356106	958	6,811	SH		OTR	1	3,771	1,209	1,831
TRACTOR SUPPLY COMPANY	COMMON STOCK	892356106	71	507	SH		DFND	5	507	0	0
TRADEWEB MARKETS INC CLASS A	COMMON STOCK	892672106	1,342	21,488	SH		DFND	1	21,256	0	232
TRADEWEB MARKETS INC CLASS A	COMMON STOCK	892672106	20	321	SH		OTR	1	321	0	0
TRADEWEB MARKETS INC CL A	COMMON STOCK	892672106	3	44	SH		DFND	5	44	0	0
TRANSDIGM GROUP INC	COMMON STOCK	893641100	67,665	109,340	SH		DFND	1	108,205	0	1,135
TRANSDIGM GROUP INC	COMMON STOCK	893641100	10,062	16,260	SH		OTR	1	15,698	312	250
TREDEGAR CORP	COMMON STOCK	894650100	30	1,800	SH		DFND	1	1,800	0	0
TRI CONTINENTAL CORP	OTHER	895436103	87	2,964	SH		DFND	1	2,964	0	0
TRICO BANCSHARES	COMMON STOCK	896095106	1	25	SH		DFND	1	25	0	0
TRICO BANCSHARES	COMMON STOCK	896095106	55	1,555	SH		OTR	1	0	1,555	0
TRIMAS CORP	COMMON STOCK	896215209	440	13,878	SH		DFND	1	13,878	0	0
TRIMAS CORP	COMMON STOCK	896215209	72	2,284	SH		OTR	1	2,284	0	0
TRIMBLE NAV LTD	COMMON STOCK	896239100	4,420	66,202	SH		DFND	1	65,118	0	1,084
TRIMBLE NAV LTD	COMMON STOCK	896239100	612	9,163	SH		OTR	1	9,101	0	62
TRIMBLE NAVIGATION LTD	COMMON STOCK	896239100	2	36	SH		DFND	5	36	0	0
TRINITY BIOTECH PLC SPON A D R	COMMON STOCK	896438306	4	1,119	SH		DFND	1	1,119	0	0
TRINITY INDS INC	COMMON STOCK	896522109	357	13,514	SH		DFND	1	12,585	0	929
TRINITY INDS INC	COMMON STOCK	896522109	8	300	SH		OTR	1	300	0	0
TRIUMPH GROUP INC	COMMON STOCK	896818101	7	549	SH		DFND	1	276	0	273
TRIUMPH GROUP INC	COMMON STOCK	896818101	1	111	SH		OTR	1	0	0	111
TRIPADVISOR INC	COMMON STOCK	896945201	57	2,012	SH		DFND	1	1,744	0	268
TRIPADVISOR INC	COMMON STOCK	896945201	0	6	SH		OTR	1	0	0	6
TRUPANION INC	COMMON STOCK	898202106	13	112	SH		DFND	1	54	0	58
TRUSTCO BK CORP NY	COMMON STOCK	898349105	9	1,301	SH		DFND	1	1,000	0	301
TRUSTMARK CORPORATION	COMMON STOCK	898402102	23	846	SH		DFND	1	816	0	30
TUPPERWARE BRANDS CORP	COMMON STOCK	899896104	3	106	SH		DFND	1	106	0	0
TUPPERWARE BRANDS CORP	COMMON STOCK	899896104	96	2,958	SH		OTR	1	2,958	0	0
TURKCELL ILETISIM HIZMET A D R	COMMON STOCK	900111204	61	11,421	SH		DFND	1	8,020	0	3,401
TURKCELL ILETISIM HIZMET A D R	COMMON STOCK	900111204	2	256	SH		OTR	1	0	135	121
TURTLE BEACH CORP	COMMON STOCK	900450206	11	500	SH		DFND	1	500	0	0
TUTOR PERINI CORP	COMMON STOCK	901109108	27	2,056	SH		DFND	1	1,148	0	908
II-VI INC	COMMON STOCK	902104108	1,046	13,767	SH		DFND	1	12,170	0	1,597
II-VI INC	COMMON STOCK	902104108	103	1,355	SH		OTR	1	142	1,213	0
II-VI INC	COMMON STOCK	902104108	2	21	SH		DFND	5	21	0	0
TYLER TECHNOLOGIES INC	COMMON STOCK	902252105	32,533	74,530	SH		DFND	1	72,783	0	1,747
TYLER TECHNOLOGIES INC	COMMON STOCK	902252105	4,422	10,130	SH		OTR	1	7,923	1,892	315
TYLER TECHNOLOGIES INC	COMMON STOCK	902252105	470	1,076	SH		DFND	5	1,076	0	0
TYSON FOODS INC CL A	COMMON STOCK	902494103	3,827	59,383	SH		DFND	1	56,846	0	2,537
TYSON FOODS INC CL A	COMMON STOCK	902494103	1,428	22,164	SH		OTR	1	21,968	196	0
TYSON FOODS INC	COMMON STOCK	902494103	2	24	SH		DFND	5	24	0	0
UDR INC	COMMON STOCK	902653104	131	3,402	SH		DFND	1	1,757	0	1,645
UDR INC	COMMON STOCK	902653104	5	128	SH		OTR	1	59	11	58
UDR INC	COMMON STOCK	902653104	6	146	SH		DFND	5	146	0	0
UFP TECHNOLOGIES INC	COMMON STOCK	902673102	186	4,000	SH		DFND	1	4,000	0	0
UFP TECHNOLOGIES INC	COMMON STOCK	902673102	60	1,290	SH		OTR	1	0	1,290	0
U G I CORP	COMMON STOCK	902681105	443	12,659	SH		DFND	1	7,558	0	5,101
U G I CORP	COMMON STOCK	902681105	1	35	SH		OTR	1	35	0	0
UGI CORP	COMMON STOCK	902681105	35	1,007	SH		DFND	5	999	0	8
UMB FINL CORP	COMMON STOCK	902788108	1,225	17,757	SH		DFND	1	17,166	0	591
UMB FINL CORP	COMMON STOCK	902788108	427	6,190	SH		OTR	1	6,190	0	0
UMB FINANCIAL CORP	COMMON STOCK	902788108	7	107	SH		DFND	5	107	0	0
USA TRUCK INC	COMMON STOCK	902925106	1,029	115,235	SH		DFND	1	115,235	0	0
US BANCORP	COMMON STOCK	902973304	319,516	6,858,037	SH		DFND	1	1,337,710	4,336	5,515,991
US BANCORP	COMMON STOCK	902973304	179,147	3,845,188	SH		OTR	1	344,574	928,069	2,572,545
US BANCORP	COMMON STOCK	902973304	688	14,755	SH		DFND	5	1,146	0	13,609
UMH PROPERTIES INC	COMMON STOCK	903002103	11	764	SH		DFND	1	764	0	0
UMH PROPERTIES INC	COMMON STOCK	903002103	1	35	SH		OTR	1	0	35	0
UMH PROPERTIES INC	COMMON STOCK	903002103	0	4	SH		DFND	5	4	0	0
UMPQUA HOLDINGS CORP	COMMON STOCK	904214103	1,020	67,367	SH		DFND	1	52,068	0	15,299
UMPQUA HOLDINGS CORP	COMMON STOCK	904214103	87	5,748	SH		OTR	1	5,558	0	190
UNDER ARMOUR INC CL A	COMMON STOCK	904311107	18	1,053	SH		DFND	1	650	0	403
UNDER ARMOUR INC	COMMON STOCK	904311107	26	1,497	SH		DFND	5	1,497	0	0
UNDER ARMOUR INC	COMMON STOCK	904311206	5	351	SH		DFND	1	351	0	0
UNIFIRST CORP MASS	COMMON STOCK	904708104	821	3,880	SH		DFND	1	3,880	0	0
UNIFIRST CORP MASS	COMMON STOCK	904708104	105	495	SH		OTR	1	495	0	0
UNIFIRST CORP	COMMON STOCK	904708104	12	59	SH		DFND	5	59	0	0
UNILEVER PLC SPONSORED A D R	COMMON STOCK	904767704	10,585	175,370	SH		DFND	1	132,361	0	43,009
UNILEVER PLC SPONSORED A D R	COMMON STOCK	904767704	3,441	57,003	SH		OTR	1	48,068	3,719	5,216
UNILEVER PLC	COMMON STOCK	904767704	331	5,475	SH		DFND	5	5,475	0	0
UNION PACIFIC CORP	COMMON STOCK	907818108	126,172	605,955	SH		DFND	1	578,290	0	27,665
UNION PACIFIC CORP	COMMON STOCK	907818108	54,099	259,820	SH		OTR	1	238,941	4,402	16,477
UNION PACIFIC CORP	COMMON STOCK	907818108	383	1,836	SH		DFND	5	1,811	0	25
UNION PACIFIC CORP	COMMON STOCK	907818108	4	26	SH	Put	OTR	1	26	0	0
UNITED BANKSHARES INC W VA	COMMON STOCK	909907107	576	17,800	SH		DFND	1	17,232	0	568
UNITED BANKSHARES INC W VA	COMMON STOCK	909907107	177	5,489	SH		OTR	1	5,382	0	107
UNITED BANKSHARES INC	COMMON STOCK	909907107	13	386	SH		DFND	5	386	0	0
UNITED AIRLINES HOLDINGS INC	COMMON STOCK	910047109	446	10,309	SH		DFND	1	8,163	0	2,146
UNITED AIRLINES HOLDINGS INC	COMMON STOCK	910047109	5	127	SH		OTR	1	46	0	81
UNITED FIRE GROUP INC	COMMON STOCK	910340108	331	13,175	SH		DFND	1	13,175	0	0
UNITED GUARDIAN INC	COMMON STOCK	910571108	43	2,959	SH		OTR	1	0	2,959	0
UNITED MICROELECTRON SP A D R	COMMON STOCK	910873405	769	91,141	SH		DFND	1	45,624	0	45,517
UNITED MICROELECTRON SP A D R	COMMON STOCK	910873405	240	28,497	SH		OTR	1	1,000	280	27,217
UNITED MICROELECTRONICS CORP	COMMON STOCK	910873405	3	403	SH		DFND	5	403	0	0
UNITED NAT FOODS INC	COMMON STOCK	911163103	33	2,037	SH		DFND	1	1,160	0	877
UNITED PARCEL SERVICE CL B	COMMON STOCK	911312106	51,259	304,389	SH		DFND	1	293,972	0	10,417
UNITED PARCEL SERVICE CL B	COMMON STOCK	911312106	54,149	321,550	SH		OTR	1	308,374	4,110	9,066
UNITED PARCEL SERVICE	COMMON STOCK	911312106	853	5,055	SH		DFND	5	4,705	0	350
UNITED RENTALS INC	COMMON STOCK	911363109	6,343	27,347	SH		DFND	1	26,424	0	923
UNITED RENTALS INC	COMMON STOCK	911363109	1,143	4,930	SH		OTR	1	4,788	4	138
UNITED RENTALS INC	COMMON STOCK	911363109	22	97	SH		DFND	5	97	0	0
UNITED STATES ANTIMONY COM	COMMON STOCK	911549103	7	14,000	SH		OTR	1	14,000	0	0
US CELLULAR CORP	COMMON STOCK	911684108	64	2,070	SH		DFND	1	1,613	0	457
US CELLULAR CORP	COMMON STOCK	911684108	87	2,873	SH		OTR	1	0	80	2,793
US FOODS HOLDING CORP	COMMON STOCK	912008109	24	719	SH		DFND	1	306	0	413
UNITED STATES STEEL CORP	COMMON STOCK	912909108	110	6,557	SH		DFND	1	4,967	0	1,590
UNITED STATES STEEL CORP	COMMON STOCK	912909108	3	187	SH		OTR	1	8	0	179
UNITIL CORP	COMMON STOCK	913259107	104	2,339	SH		DFND	1	2,274	0	65
UNITIL CORP	COMMON STOCK	913259107	36	807	SH		OTR	1	788	19	0
UNITIL CORP	COMMON STOCK	913259107	5	122	SH		DFND	5	122	0	0
UNIVERSAL CORP VA	COMMON STOCK	913456109	23	481	SH		DFND	1	153	0	328
UNIVERSAL ELECTRS INC	COMMON STOCK	913483103	229	4,349	SH		DFND	1	4,349	0	0
UNIVERSAL ELECTRS INC	COMMON STOCK	913483103	53	1,025	SH		OTR	1	7	1,018	0
UNIVERSAL HLTH SVCS INC	COMMON STOCK	913903100	1,078	7,845	SH		DFND	1	7,292	0	553
UNIVERSAL HLTH SVCS INC	COMMON STOCK	913903100	516	3,753	SH		OTR	1	2,183	1,325	245
UNIVERSAL HEALTH SERVICES	COMMON STOCK	913903100	38	277	SH		DFND	5	277	0	0
UNIVERSAL TECHNICAL INSTITUT	COMMON STOCK	913915104	145	22,506	SH		DFND	1	22,506	0	0
UNIVESTE FINANCIAL CORPORATION	COMMON STOCK	915271100	97	4,728	SH		DFND	1	216	0	4,512
URANIUM ENERGY CORP	COMMON STOCK	916896103	1	595	SH		DFND	1	595	0	0
URBAN OUTFITTERS INC	COMMON STOCK	917047102	7	269	SH		DFND	1	0	0	269
URBAN OUTFITTERS INC	COMMON STOCK	917047102	2	64	SH		OTR	1	0	0	64
URSTADT BIDDLE CL A	COMMON STOCK	917286205	2	153	SH		DFND	1	0	0	153
UTAH MED PRODS INC	COMMON STOCK	917488108	173	2,057	SH		DFND	1	2,057	0	0
UTAH MED PRODS INC	COMMON STOCK	917488108	4	48	SH		OTR	1	48	0	0
V F CORP	COMMON STOCK	918204108	24,898	291,514	SH		DFND	1	284,530	0	6,984
V F CORP	COMMON STOCK	918204108	8,157	95,513	SH		OTR	1	87,163	4,664	3,686
VF CORP	COMMON STOCK	918204108	22	261	SH		DFND	5	232	0	29
VSE CORP	COMMON STOCK	918284100	6	168	SH		DFND	1	0	0	168
VALLEY NATIONAL BANCORP	COMMON STOCK	919794107	11	1,159	SH		DFND	1	837	0	322
VALLEY NATIONAL BANCORP	COMMON STOCK	919794107	1	147	SH		OTR	1	0	0	147
VALMONT INDS INC	COMMON STOCK	920253101	339	1,938	SH		DFND	1	1,752	0	186
VALMONT INDS INC	COMMON STOCK	920253101	2	9	SH		OTR	1	9	0	0
VALUE LINE INC	COMMON STOCK	920437100	4	128	SH		DFND	1	128	0	0
VANDA PHARMACEUTICALS INC	COMMON STOCK	921659108	6	446	SH		DFND	1	446	0	0
VANGUARD DIVIDEND APPRECIATION	OTHER	921908844	12,811	90,746	SH		DFND	1	90,021	0	725
VANGUARD DIVIDEND APPRECIATION	OTHER	921908844	5,089	36,048	SH		OTR	1	35,112	190	746
VANGUARD DIVIDND APPRECIATION	OTHER	921908844	6,464	45,785	SH		DFND	5	12,428	0	33,357
VANGUARD TOTAL INTERNATIONAL E	OTHER	921909768	203	3,371	SH		DFND	1	3,371	0	0
VANGUARD TOTAL INTERNATIONAL E	OTHER	921909768	75	1,245	SH		OTR	1	200	0	1,045
VANGUARD TOTAL INTL STOCK INDX	OTHER	921909768	1	15	SH		DFND	5	15	0	0
VANGUARD MEGA CAP GROWTH ETF E	OTHER	921910816	897	4,404	SH		DFND	1	4,404	0	0
VANGUARD MEGA CAP GROWTH ETF E	OTHER	921910816	258	1,266	SH		OTR	1	1,266	0	0
VANGUARD MEGA CAP 300 GROWTH	OTHER	921910816	7,314	35,898	SH		DFND	5	3	0	35,895
VANGUARD MEGA CAP VALUE INDEX	OTHER	921910840	16	181	SH		DFND	1	181	0	0
VANGUARD MEGA CAP VALUE INDEX	OTHER	921910840	114	1,315	SH		OTR	1	1,315	0	0
VANGUARD MEGA CAP 300 VALUE	OTHER	921910840	5,411	62,323	SH		DFND	5	0	0	62,323
VANGUARD MEGA CAP INDEX FUND E	OTHER	921910873	1,019	7,634	SH		DFND	1	7,634	0	0
VANGUARD S P 500 GROWTH INDEX	OTHER	921932505	3,037	13,200	SH		DFND	1	13,150	0	50
VANGUARD S&P 500 GROWTH	OTHER	921932505	1	3	SH		DFND	5	3	0	0
VANGUARD S P 500 VALUE INDEX E	OTHER	921932703	807	6,523	SH		DFND	1	6,523	0	0
VANGUARD S&P SMALL CAP 600	OTHER	921932794	15	75	SH		DFND	5	75	0	0
VANGUARD S P SMALL CAP 600 ETF	OTHER	921932828	109	651	SH		DFND	1	651	0	0
VANGUARD S P MID CAP 400 INDEX	OTHER	921932885	150	967	SH		DFND	1	967	0	0
VANGUARD LONG TERM BOND INDEX	OTHER	921937793	4,107	37,461	SH		DFND	1	37,372	0	89
VANGUARD LONG TERM BOND INDEX	OTHER	921937793	348	3,178	SH		OTR	1	3,178	0	0
VANGUARD LONG TERM BOND	OTHER	921937793	1,013	9,240	SH		DFND	5	8,312	0	928
VANGUARD INTERMEDIATE TERM BON	OTHER	921937819	125,928	1,356,252	SH		DFND	1	1,346,972	0	9,280
VANGUARD INTERMEDIATE TERM BON	OTHER	921937819	12,845	138,338	SH		OTR	1	124,763	1,650	11,925
VANGUARD INTERMED TERM BOND	OTHER	921937819	6,220	66,972	SH		DFND	5	34,783	0	32,189
VANGUARD SHORT TERM BOND INDEX	OTHER	921937827	48,071	579,865	SH		DFND	1	578,311	0	1,554
VANGUARD SHORT TERM BOND INDEX	OTHER	921937827	5,101	61,528	SH		OTR	1	57,503	2,562	1,463
VANGUARD SHORT TERM BOND	OTHER	921937827	4,827	58,227	SH		DFND	5	20,184	0	38,043
VANGUARD TOTAL BOND MARKET ETF	OTHER	921937835	16,963	192,345	SH		DFND	1	192,045	0	300
VANGUARD TOTAL BOND MARKET ETF	OTHER	921937835	3,241	36,746	SH		OTR	1	33,811	1,200	1,735
VANGUARD TOTAL BOND MARKET	OTHER	921937835	1,460	16,550	SH		DFND	5	4,038	0	12,512
VANGUARD FTSE DEVELOPED ETF	OTHER	921943858	120,765	2,558,020	SH		DFND	1	2,467,832	0	90,188
VANGUARD FTSE DEVELOPED ETF	OTHER	921943858	22,942	485,957	SH		OTR	1	479,032	630	6,295
VANGUARD FTSE DEVELOPED MKTS	OTHER	921943858	4,144	87,780	SH		DFND	5	65,836	0	21,944
VANGUARD HIGH DIVIDEND YIELD E	OTHER	921946406	4,564	49,877	SH		DFND	1	49,877	0	0
VANGUARD HIGH DIVIDEND YIELD E	OTHER	921946406	432	4,720	SH		OTR	1	4,720	0	0
VANGUARD HIGH DVD YIELD	OTHER	921946406	4,127	45,099	SH		DFND	5	113	0	44,986
VANGUARD INTERNATIONAL DIVIDEN	OTHER	921946810	132	1,623	SH		DFND	1	1,623	0	0
VANGUARD EMERGING MARKETS ETF	OTHER	921946885	12	142	SH		DFND	1	142	0	0
VANGUARD SHORT TERM INFLATION	OTHER	922020805	28,083	549,572	SH		DFND	1	546,851	0	2,721
VANGUARD SHORT TERM INFLATION	OTHER	922020805	5,489	107,422	SH		OTR	1	103,993	193	3,236
VANGUARD SHORT TERM INFLATION	OTHER	922020805	20	392	SH		DFND	5	136	0	256
VANGUARD GLOBAL EX U S REAL ET	OTHER	922042676	22,964	422,821	SH		DFND	1	416,120	0	6,701
VANGUARD GLOBAL EX U S REAL ET	OTHER	922042676	9,171	168,865	SH		OTR	1	166,839	794	1,232
VANGUARD GLOBAL EX US REAL	OTHER	922042676	291	5,352	SH		DFND	5	5,352	0	0
VANGUARD FTSE ALL WORLD EX US	OTHER	922042718	50	415	SH		DFND	1	365	0	50
VANGUARD FTSE ALL WORLD EX US	OTHER	922042718	84	693	SH		OTR	1	693	0	0
VANGUARD TOTAL WORLD STOCK IND	OTHER	922042742	204	2,208	SH		DFND	1	2,208	0	0
VANGUARD FTSE ALL WORLD EX US	OTHER	922042775	29,642	507,923	SH		DFND	1	492,330	0	15,593
VANGUARD FTSE ALL WORLD EX US	OTHER	922042775	4,132	70,799	SH		OTR	1	65,214	5,250	335
VANGUARD FTSE ALL WORLD EX US	OTHER	922042775	1,117	19,138	SH		DFND	5	19,138	0	0
VANGUARD FTSE EMERGING MARKETS	OTHER	922042858	141,498	2,823,736	SH		DFND	1	2,483,322	0	340,414
VANGUARD FTSE EMERGING MARKETS	OTHER	922042858	17,923	357,647	SH		OTR	1	345,643	135	11,869
VANGUARD EMERGING MARKETS	OTHER	922042858	7,513	149,922	SH		DFND	5	25,220	0	124,702
VANGUARD FTSE PACIFIC INDEX FU	OTHER	922042866	22	282	SH		DFND	1	282	0	0
VANGUARD FTSE PACIFIC INDEX FU	OTHER	922042866	3	35	SH		OTR	1	35	0	0
VANGUARD FTSE EUROPE INDEX FUN	OTHER	922042874	153	2,548	SH		DFND	1	2,548	0	0
VANGUARD FTSE EUROPE INDEX FUN	OTHER	922042874	21	345	SH		OTR	1	345	0	0
VAPOTHERM INC	COMMON STOCK	922107107	15	546	SH		DFND	1	459	0	87
VARONIS SYSTEMS INC	COMMON STOCK	922280102	1,525	9,323	SH		DFND	1	9,301	0	22
VARONIS SYSTEMS INC	COMMON STOCK	922280102	264	1,616	SH		OTR	1	1,616	0	0
VEEVA SYSTEMS INC CLASS A	COMMON STOCK	922475108	5,895	21,652	SH		DFND	1	21,046	0	606
VEEVA SYSTEMS INC CLASS A	COMMON STOCK	922475108	1,587	5,829	SH		OTR	1	1,661	4,150	18
VEEVA SYSTEMS INC	COMMON STOCK	922475108	77	282	SH		DFND	5	278	0	4
VANGUARD TAX EXEMPT BOND ETF	OTHER	922907746	45,639	826,938	SH		DFND	1	826,938	0	0
VANGUARD TAX EXEMPT BOND ETF	OTHER	922907746	6,049	109,603	SH		OTR	1	107,918	0	1,685
VANGUARD TAX EXPEMPT BOND INDX	OTHER	922907746	1,537	27,846	SH		DFND	5	1,981	0	25,865
VANGUARD S P 500 ETF	OTHER	922908363	25,975	75,576	SH		DFND	1	73,886	0	1,690
VANGUARD S P 500 ETF	OTHER	922908363	9,169	26,678	SH		OTR	1	24,178	2,125	375
VANGUARD S&P 500 INDEX	OTHER	922908363	11,228	32,666	SH		DFND	5	18,905	0	13,761
VANGUARD MID CAP VALUE INDEX E	OTHER	922908512	15,994	134,445	SH		DFND	1	133,378	0	1,067
VANGUARD MID CAP VALUE INDEX E	OTHER	922908512	3,415	28,704	SH		OTR	1	28,704	0	0
VANGUARD MID CAP VALUE INDEX	OTHER	922908512	2,722	22,880	SH		DFND	5	218	0	22,662
VANGUARD MID CAP GROWTH INDEX	OTHER	922908538	2,420	11,410	SH		DFND	1	10,651	0	759
VANGUARD MID CAP GROWTH INDEX	OTHER	922908538	468	2,207	SH		OTR	1	975	0	1,232
VANGUARD MID CAP GROWTH INDEX	OTHER	922908538	3,617	17,054	SH		DFND	5	0	0	17,054
VANGUARD REAL ESTATE ETF	OTHER	922908553	349,620	4,116,559	SH		DFND	1	3,856,581	0	259,978
VANGUARD REAL ESTATE ETF	OTHER	922908553	53,226	626,703	SH		OTR	1	612,254	8,577	5,872
VANGUARD REIT	OTHER	922908553	3,771	44,402	SH		DFND	5	38,480	0	5,922
VANGUARD REAL ESTATE ETF	OTHER	922908553	63	37	SH	Put	DFND	1	37	0	0
VANGUARD SMALL CAP GROWTH INDE	OTHER	922908595	7,042	26,309	SH		DFND	1	24,643	0	1,666
VANGUARD SMALL CAP GROWTH INDE	OTHER	922908595	186	694	SH		OTR	1	694	0	0
VANGUARD SMALL CAP GROWTH	OTHER	922908595	6,450	24,096	SH		DFND	5	0	0	24,096
VANGUARD SMALL CAP VALUE INDEX	OTHER	922908611	1,057	7,433	SH		DFND	1	5,631	0	1,802
VANGUARD SMALL CAP VALUE INDEX	OTHER	922908611	221	1,555	SH		OTR	1	1,555	0	0
VANGUARD SMALL CAP VALUE	OTHER	922908611	2,480	17,437	SH		DFND	5	17	0	17,420
VANGUARD MID CAP INDEX FUND ET	OTHER	922908629	8,819	42,651	SH		DFND	1	42,272	0	379
VANGUARD MID CAP INDEX FUND ET	OTHER	922908629	13,673	66,127	SH		OTR	1	62,627	0	3,500
VANGUARD MID CAP	OTHER	922908629	2,463	11,908	SH		DFND	5	4,525	0	7,383
VANGUARD LARGE CAP INDEX FUND	OTHER	922908637	1,436	8,172	SH		DFND	1	8,172	0	0
VANGUARD LARGE CAP INDEX FUND	OTHER	922908637	932	5,306	SH		OTR	1	4,856	0	450
VANGUARD LARGE CAP ETF	OTHER	922908637	240	1,363	SH		DFND	5	490	0	873
VANGUARD EXTENDED MARKET INDEX	OTHER	922908652	85,611	519,864	SH		DFND	1	516,219	0	3,645
VANGUARD EXTENDED MARKET INDEX	OTHER	922908652	13,429	81,544	SH		OTR	1	81,284	0	260
VANGUARD EXTENDED MARKET	OTHER	922908652	15,897	96,530	SH		DFND	5	96,041	0	489
VANGUARD GROWTH INDEX FUND ETF	OTHER	922908736	45,725	180,489	SH		DFND	1	179,320	0	1,169
VANGUARD GROWTH INDEX FUND ETF	OTHER	922908736	10,663	42,089	SH		OTR	1	41,192	0	897
VANGUARD GROWTH	OTHER	922908736	6,620	26,129	SH		DFND	5	22,309	0	3,820
VANGUARD VALUE INDEX FUND ETF	OTHER	922908744	26,711	224,541	SH		DFND	1	222,498	0	2,043
VANGUARD VALUE INDEX FUND ETF	OTHER	922908744	4,307	36,205	SH		OTR	1	36,060	0	145
VANGUARD VALUE	OTHER	922908744	6,293	52,897	SH		DFND	5	22,894	0	30,003
VANGUARD SMALL CAP INDEX FUND	OTHER	922908751	7,587	38,970	SH		DFND	1	36,182	0	2,788
VANGUARD SMALL CAP INDEX FUND	OTHER	922908751	12,540	64,414	SH		OTR	1	63,813	0	601
VANGUARD SMALL CAP	OTHER	922908751	4,141	21,269	SH		DFND	5	4,305	0	16,964
VANGUARD TOTAL STOCK MARKET ET	OTHER	922908769	373,984	1,921,411	SH		DFND	1	1,908,079	0	13,332
VANGUARD TOTAL STOCK MARKET ET	OTHER	922908769	91,669	470,965	SH		OTR	1	449,498	16,942	4,525
VANGUARD TOTAL STOCK MARKET	OTHER	922908769	1,299	6,673	SH		DFND	5	28	0	6,645
VERITEX HOLDINGS INC	COMMON STOCK	923451108	43	1,661	SH		OTR	1	0	1,661	0
VERITIV CORP	COMMON STOCK	923454102	0	2	SH		DFND	1	2	0	0
VERITIV CORP	COMMON STOCK	923454102	0	4	SH		OTR	1	0	4	0
VERMILION ENERGY INC	COMMON STOCK	923725105	1	300	SH		DFND	1	300	0	0
VERMILION ENERGY INC	COMMON STOCK	923725105	1	300	SH		OTR	1	300	0	0
VERONA PHARMA PLC A D R	COMMON STOCK	925050106	27	3,840	SH		OTR	1	3,840	0	0
VIAVI SOLUTIONS INC	COMMON STOCK	925550105	166	11,082	SH		DFND	1	1,374	0	9,708
VIAVI SOLUTIONS INC	COMMON STOCK	925550105	34	2,255	SH		OTR	1	0	2,255	0
VICI PPTYS INC	COMMON STOCK	925652109	697	27,338	SH		DFND	1	25,560	0	1,778
VICI PPTYS INC	COMMON STOCK	925652109	91	3,568	SH		OTR	1	1,035	36	2,497
VICI PROPERTIES INC	COMMON STOCK	925652109	9	340	SH		DFND	5	340	0	0
VISTA GOLD CORP	COMMON STOCK	927926303	0	100	SH		DFND	1	100	0	0
VIRTU FINANCIAL INC CLASS A	COMMON STOCK	928254101	65	2,566	SH		DFND	1	2,104	0	462
VIRTU FINANCIAL INC CLASS A	COMMON STOCK	928254101	69	2,718	SH		OTR	1	0	2,574	144
VISHAY INTERTECHNOLOGY INC	COMMON STOCK	928298108	47	2,257	SH		DFND	1	1,280	0	977
VISTA OUTDOOR INC WI	COMMON STOCK	928377100	1,130	47,572	SH		DFND	1	3,628	0	43,944
VISTA OUTDOOR INC WI	COMMON STOCK	928377100	5	216	SH		OTR	1	216	0	0
VMWARE INC CL A	COMMON STOCK	928563402	1,003	7,150	SH		DFND	1	6,632	0	518
VMWARE INC CL A	COMMON STOCK	928563402	59	421	SH		OTR	1	326	86	9
VMWARE INC	COMMON STOCK	928563402	298	2,114	SH		DFND	5	2,114	0	0
VONTIER CORPORATION COM	COMMON STOCK	928881101	572	17,138	SH		DFND	1	16,252	0	886
VONTIER CORPORATION COM	COMMON STOCK	928881101	24	741	SH		OTR	1	716	11	14
VORNADO REALTY TRUST	COMMON STOCK	929042109	322	8,605	SH		DFND	1	4,561	0	4,044
VORNADO REALTY TRUST	COMMON STOCK	929042109	52	1,405	SH		OTR	1	1,345	25	35
VORNADO REALTY TRUST	COMMON STOCK	929042109	1	19	SH		DFND	5	19	0	0
VOYA FINANCIAL INC	COMMON STOCK	929089100	565	9,608	SH		DFND	1	7,729	0	1,879
VOYA FINANCIAL INC	COMMON STOCK	929089100	5	78	SH		OTR	1	78	0	0
VOYA FINANCIAL INC	COMMON STOCK	929089100	54	922	SH		DFND	5	922	0	0
VULCAN MATERIALS CO	COMMON STOCK	929160109	1,681	11,329	SH		DFND	1	9,300	0	2,029
VULCAN MATERIALS CO	COMMON STOCK	929160109	262	1,767	SH		OTR	1	1,019	25	723
W D 40 CO	COMMON STOCK	929236107	388	1,460	SH		DFND	1	839	0	621
W D 40 CO	COMMON STOCK	929236107	230	865	SH		OTR	1	470	0	395
WSFS FINL CORP	COMMON STOCK	929328102	1,955	43,568	SH		DFND	1	42,869	0	699
WSFS FINL CORP	COMMON STOCK	929328102	53	1,170	SH		OTR	1	1,170	0	0
WABASH NATL CORP	COMMON STOCK	929566107	258	14,963	SH		DFND	1	1,255	0	13,708
WABTEC CORP	COMMON STOCK	929740108	351	4,796	SH		DFND	1	3,453	0	1,343
WABTEC CORP	COMMON STOCK	929740108	224	3,060	SH		OTR	1	2,907	82	71
WABTEC CORP	COMMON STOCK	929740108	1	19	SH		DFND	5	19	0	0
WADDELL & REED FINANCIAL INC	COMMON STOCK	930059100	173	6,773	SH		DFND	1	6,307	0	466
WADDELL & REED FINANCIAL INC	COMMON STOCK	930059100	63	2,454	SH		OTR	1	2,054	0	400
WALMART INC	COMMON STOCK	931142103	260,316	1,805,870	SH		DFND	1	1,781,750	0	24,120
WALMART INC	COMMON STOCK	931142103	90,302	626,447	SH		OTR	1	586,715	24,718	15,014
WAL-MART STORES INC	COMMON STOCK	931142103	116	800	SH		DFND	5	800	0	0
WALGREENS BOOTS ALLIANCE INC	COMMON STOCK	931427108	21,531	539,889	SH		DFND	1	455,947	0	83,942
WALGREENS BOOTS ALLIANCE INC	COMMON STOCK	931427108	3,683	92,328	SH		OTR	1	83,317	2,515	6,496
WALGREENS BOOTS ALLIANCE INC	COMMON STOCK	931427108	24	603	SH		DFND	5	603	0	0
WARNER MUSIC GROUP CORP CL A	COMMON STOCK	934550203	292	7,679	SH		DFND	1	7,679	0	0
WARNER MUSIC GROUP CORP CL A	COMMON STOCK	934550203	8	206	SH		OTR	1	206	0	0
WASHINGTON FEDERAL INC	COMMON STOCK	938824109	24	929	SH		DFND	1	929	0	0
WASHINGTON FEDERAL INC	COMMON STOCK	938824109	155	6,020	SH		OTR	1	419	5,532	69
WASHINGTON REAL ESTATE INVESTM	COMMON STOCK	939653101	165	7,654	SH		DFND	1	7,654	0	0
WASHINGTON REAL ESTATE INVESTM	COMMON STOCK	939653101	4	196	SH		OTR	1	196	0	0
WASHINGTON TRUST BANCORP INC	COMMON STOCK	940610108	10	215	SH		DFND	1	215	0	0
WATERS CORP	COMMON STOCK	941848103	446	1,806	SH		DFND	1	1,350	0	456
WATERS CORP	COMMON STOCK	941848103	38	154	SH		OTR	1	127	17	10
WATSCO INC	COMMON STOCK	942622200	2,054	9,067	SH		DFND	1	8,819	0	248
WATSCO INC	COMMON STOCK	942622200	21	92	SH		OTR	1	74	8	10
WATSCO INC	COMMON STOCK	942622200	124	547	SH		DFND	5	547	0	0
WATTS WATER TECHNOLOGIES CL A	COMMON STOCK	942749102	82	670	SH		DFND	1	670	0	0
WEBSTER FINANCIAL CORP	COMMON STOCK	947890109	62	1,458	SH		DFND	1	796	0	662
WEBSTER FINANCIAL CORP	COMMON STOCK	947890109	4	84	SH		OTR	1	0	45	39
WEIBO CORP SPON A D R	COMMON STOCK	948596101	364	8,872	SH		DFND	1	7,888	0	984
WEIBO CORP SPON A D R	COMMON STOCK	948596101	41	994	SH		OTR	1	994	0	0
WEINGARTEN RLTY INVTS	COMMON STOCK	948741103	224	10,347	SH		DFND	1	9,752	0	595
WEINGARTEN RLTY INVTS	COMMON STOCK	948741103	92	4,233	SH		OTR	1	4,197	0	36
WEIS MKTS INC	COMMON STOCK	948849104	124	2,590	SH		DFND	1	1,293	0	1,297
WELBILT INC	COMMON STOCK	949090104	155	11,761	SH		DFND	1	11,323	0	438
WELBILT INC	COMMON STOCK	949090104	0	19	SH		OTR	1	0	0	19
WELLS FARGO CO	COMMON STOCK	949746101	56,688	1,878,345	SH		DFND	1	1,822,392	0	55,953
WELLS FARGO CO	COMMON STOCK	949746101	24,453	810,259	SH		OTR	1	759,926	12,421	37,912
WELLS FARGO & COMPANY	COMMON STOCK	949746101	31	1,016	SH		DFND	5	998	0	18
WERNER ENTERPRISES INC	COMMON STOCK	950755108	828	21,121	SH		DFND	1	18,660	0	2,461
WERNER ENTERPRISES INC	COMMON STOCK	950755108	157	4,017	SH		OTR	1	3,803	21	193
WERNER ENTERPRISES INC	COMMON STOCK	950755108	2	52	SH		DFND	5	0	0	52
WESBANCO INC	COMMON STOCK	950810101	37	1,242	SH		DFND	1	653	0	589
WESBANCO INC	COMMON STOCK	950810101	1,939	64,707	SH		OTR	1	64,707	0	0
WEST PHARMACEUTICAL SVCS INC	COMMON STOCK	955306105	8,320	29,368	SH		DFND	1	27,587	0	1,781
WEST PHARMACEUTICAL SVCS INC	COMMON STOCK	955306105	935	3,302	SH		OTR	1	3,212	74	16
WEST PHARMACEUTICALS SVCS	COMMON STOCK	955306105	1,107	3,907	SH		DFND	5	3,899	0	8
WESTAMERICA BANCORPORATION	COMMON STOCK	957090103	6	105	SH		DFND	1	105	0	0
WESTAMERICA BANCORPORATION	COMMON STOCK	957090103	217	3,918	SH		OTR	1	3,918	0	0
WESTERN ALLIANCE BANCORPORATIO	COMMON STOCK	957638109	942	15,714	SH		DFND	1	10,761	0	4,953
WESTERN ALLIANCE BANCORPORATIO	COMMON STOCK	957638109	160	2,674	SH		OTR	1	166	96	2,412
WESTERN ALLIANCE BANCORP	COMMON STOCK	957638109	67	1,121	SH		DFND	5	1,121	0	0
WESTERN DIGITAL CORP	COMMON STOCK	958102105	35,993	649,806	SH		DFND	1	646,969	0	2,837
WESTERN DIGITAL CORP	COMMON STOCK	958102105	4,183	75,519	SH		OTR	1	65,040	1,505	8,974
WESTERN DIGITAL CORP	COMMON STOCK	958102105	11	203	SH		DFND	5	203	0	0
WESTERN ASSET INTM MUNI FD ICO	OTHER	958435109	5	526	SH		DFND	1	526	0	0
WESTERN MIDSTREAM PARTNERS L	COMMON STOCK	958669103	220	15,925	SH		DFND	1	925	0	15,000
WESTERN MIDSTREAM PARTNERS L	COMMON STOCK	958669103	48	3,508	SH		OTR	1	0	0	3,508
WESTERN UNION CO	COMMON STOCK	959802109	640	29,152	SH		DFND	1	25,424	0	3,728
WESTERN UNION CO	COMMON STOCK	959802109	144	6,548	SH		OTR	1	6,465	0	83
WESTERN UNION CO	COMMON STOCK	959802109	2	69	SH		DFND	5	69	0	0
WESTLAKE CHEMICAL CORP	COMMON STOCK	960413102	184	2,251	SH		DFND	1	1,409	0	842
WESTLAKE CHEMICAL CORP	COMMON STOCK	960413102	10	112	SH		OTR	1	28	52	32
WESTPORT FUEL SYSTEMS INC	COMMON STOCK	960908309	1,334	250,200	SH		DFND	1	250,200	0	0
WESTPAC BANKING CORP SP A D R	COMMON STOCK	961214301	31	2,088	SH		DFND	1	2,088	0	0
WESTPAC BANKING CORP SP A D R	COMMON STOCK	961214301	17	1,130	SH		OTR	1	1,130	0	0
WESTWATER RESOURCES INC	COMMON STOCK	961684206	3	600	SH		DFND	1	600	0	0
WEYCO GROUP INC	COMMON STOCK	962149100	6	388	SH		DFND	1	0	0	388
WEYERHAEUSER CO	COMMON STOCK	962166104	11,818	352,476	SH		DFND	1	343,358	0	9,118
WEYERHAEUSER CO	COMMON STOCK	962166104	8,116	242,057	SH		OTR	1	230,030	9,291	2,736
WEYERHAEUSER CO	COMMON STOCK	962166104	48	1,432	SH		DFND	5	1,432	0	0
WHEATON PRECIOUS METAL CORP	COMMON STOCK	962879102	361	8,666	SH		DFND	1	5,349	0	3,317
WHEATON PRECIOUS METAL CORP	COMMON STOCK	962879102	33	799	SH		OTR	1	774	0	25
WHIRLPOOL CORP	COMMON STOCK	963320106	1,538	8,523	SH		DFND	1	8,156	0	367
WHIRLPOOL CORP	COMMON STOCK	963320106	2,829	15,672	SH		OTR	1	15,216	256	200
WHIRLPOOL CORP	COMMON STOCK	963320106	1	8	SH		DFND	5	8	0	0
WHITESTONE REIT B	COMMON STOCK	966084204	4	443	SH		DFND	1	0	0	443
WILEY JOHN SONS INC	COMMON STOCK	968223206	54	1,186	SH		DFND	1	1,079	0	107
WILEY JOHN SONS INC	COMMON STOCK	968223206	14	300	SH		OTR	1	0	0	300
WILLIAMS COS INC	COMMON STOCK	969457100	2,020	100,726	SH		DFND	1	92,607	0	8,119
WILLIAMS COS INC	COMMON STOCK	969457100	1,106	55,130	SH		OTR	1	53,653	659	818
WILLIAMS COMPANIES INC	COMMON STOCK	969457100	20	996	SH		DFND	5	996	0	0
WILLIAMS SONOMA INC	COMMON STOCK	969904101	3,289	32,297	SH		DFND	1	31,551	0	746
WILLIAMS SONOMA INC	COMMON STOCK	969904101	1,125	11,041	SH		OTR	1	11,003	38	0
WILLIAMS-SONOMA INC	COMMON STOCK	969904101	43	421	SH		DFND	5	394	0	27
WILLSCOT MOBIL MINI HLDNG CORP	COMMON STOCK	971378104	656	28,304	SH		DFND	1	28,241	0	63
WILLSCOT MOBIL MINI HLDNG CORP	COMMON STOCK	971378104	100	4,339	SH		OTR	1	4,248	43	48
WILLSCOT MOBILE MINI HOLDINGS	COMMON STOCK	971378104	0	11	SH		DFND	5	11	0	0
WINGSTOP INC	COMMON STOCK	974155103	2	14	SH		DFND	1	10	0	4
WINGSTOP INC	COMMON STOCK	974155103	12	92	SH		OTR	1	0	92	0
WINGSTOP INC	COMMON STOCK	974155103	3	26	SH		DFND	5	26	0	0
WINMARK CORP	COMMON STOCK	974250102	630	3,390	SH		DFND	1	3,300	0	90
WINNEBAGO INDS INC	COMMON STOCK	974637100	206	3,440	SH		DFND	1	3,040	0	400
WOLVERINE WORLD WIDE INC	COMMON STOCK	978097103	1,601	51,235	SH		DFND	1	51,216	0	19
WOLVERINE WORLD WIDE INC	COMMON STOCK	978097103	171	5,476	SH		OTR	1	3,855	1,621	0
WOODWARD INC	COMMON STOCK	980745103	1,331	10,948	SH		DFND	1	10,737	0	211
WOODWARD INC	COMMON STOCK	980745103	1,297	10,673	SH		OTR	1	3,766	56	6,851
WOODWARD GOVERNOR CO	COMMON STOCK	980745103	34	283	SH		DFND	5	283	0	0
WOORI FINANCIAL SPON ADR A D R	COMMON STOCK	981064108	766	27,990	SH		DFND	1	14,794	0	13,196
WOORI FINANCIAL SPON ADR A D R	COMMON STOCK	981064108	39	1,414	SH		OTR	1	0	43	1,371
WORLD ACCEP CORP	COMMON STOCK	981419104	2	20	SH		DFND	1	20	0	0
WORLD FUEL SVCS CORP	COMMON STOCK	981475106	65	2,083	SH		DFND	1	1,918	0	165
WORLD FUEL SVCS CORP	COMMON STOCK	981475106	124	3,996	SH		OTR	1	0	3,996	0
WORTHINGTON INDUSTRIES INC	COMMON STOCK	981811102	23	445	SH		DFND	1	124	0	321
WORTHINGTON INDUSTRIES INC	COMMON STOCK	981811102	8	152	SH		OTR	1	0	0	152
WYNN RESORTS LTD	COMMON STOCK	983134107	453	4,015	SH		DFND	1	3,007	0	1,008
WYNN RESORTS LTD	COMMON STOCK	983134107	15	143	SH		OTR	1	18	74	51
WYNN RESORTS LTD	COMMON STOCK	983134107	5	44	SH		DFND	5	44	0	0
XPO LOGISTICS INC	COMMON STOCK	983793100	1,679	14,086	SH		DFND	1	12,511	0	1,575
XPO LOGISTICS INC	COMMON STOCK	983793100	22	180	SH		OTR	1	157	0	23
XILINX INC	COMMON STOCK	983919101	9,164	64,642	SH		DFND	1	61,656	0	2,986
XILINX INC	COMMON STOCK	983919101	3,701	26,102	SH		OTR	1	16,701	848	8,553
XILINX INC	COMMON STOCK	983919101	79	557	SH		DFND	5	557	0	0
XENIA HOTELS RESORTS INC	COMMON STOCK	984017103	5	326	SH		DFND	1	326	0	0
Y MABS THERAPEUTICS INC	COMMON STOCK	984241109	23	465	SH		DFND	1	384	0	81
YRC WORLDWIDE INC	COMMON STOCK	984249607	4	1,000	SH		DFND	1	1,000	0	0
YELP INC	COMMON STOCK	985817105	31	951	SH		DFND	1	912	0	39
YELP INC	COMMON STOCK	985817105	1	45	SH		OTR	1	0	0	45
YELP INC	COMMON STOCK	985817105	0	1	SH		DFND	5	1	0	0
YORK WATER CO	COMMON STOCK	987184108	1	24	SH		DFND	1	24	0	0
YUM BRANDS INC	COMMON STOCK	988498101	11,083	102,087	SH		DFND	1	97,158	0	4,929
YUM BRANDS INC	COMMON STOCK	988498101	4,825	44,442	SH		OTR	1	39,828	4,008	606
YUM BRANDS INC	COMMON STOCK	988498101	7	65	SH		DFND	5	65	0	0
ZEBRA TECHNOLOGIES CORP CL A	COMMON STOCK	989207105	9,602	24,982	SH		DFND	1	23,791	0	1,191
ZEBRA TECHNOLOGIES CORP CL A	COMMON STOCK	989207105	1,460	3,798	SH		OTR	1	3,165	633	0
ZEBRA TECHNOLOGIES CORP	COMMON STOCK	989207105	178	462	SH		DFND	5	450	0	12
ZIONS BANCORP NA	COMMON STOCK	989701107	1,052	24,230	SH		DFND	1	20,826	0	3,404
ZIONS BANCORP NA	COMMON STOCK	989701107	136	3,145	SH		OTR	1	3,109	0	36
ZUMIEZ INC	COMMON STOCK	989817101	6	176	SH		DFND	1	176	0	0
KIRKLAND LAKE GOLD LTD	COMMON STOCK	49741E100	83	2,022	SH		DFND	1	1,922	0	100
KIRKLAND LAKE GOLD LTD	COMMON STOCK	49741E100	13	314	SH		OTR	1	214	0	100
PETRO CHINA COMPANY LTD A D R	COMMON STOCK	71646E100	39	1,271	SH		DFND	1	1,181	0	90
PETRO CHINA COMPANY LTD A D R	COMMON STOCK	71646E100	23	755	SH		OTR	1	737	0	18
TERMINIX GLOBAL HLDGS INC	COMMON STOCK	88087E100	520	10,203	SH		DFND	1	10,033	0	170
TERMINIX GLOBAL HLDGS INC	COMMON STOCK	88087E100	71	1,385	SH		OTR	1	1,385	0	0
BLACKROCK CORE BOND TRUST	OTHER	09249E101	6	380	SH		DFND	1	380	0	0
CDK GLOBAL INC	COMMON STOCK	12508E101	626	12,078	SH		DFND	1	10,977	0	1,101
CDK GLOBAL INC	COMMON STOCK	12508E101	479	9,230	SH		OTR	1	6,800	1,536	894
CONFORMIS INC	COMMON STOCK	20717E101	1	2,000	SH		DFND	1	2,000	0	0
EQUITABLE HLDGS INC	COMMON STOCK	29452E101	203	7,952	SH		DFND	1	4,353	0	3,599
EQUITABLE HLDGS INC	COMMON STOCK	29452E101	7	266	SH		OTR	1	0	37	229
RAYTHEON TECHNOLOGIES CORP COM	COMMON STOCK	75513E101	60,070	840,026	SH		DFND	1	812,017	0	28,009
RAYTHEON TECHNOLOGIES CORP COM	COMMON STOCK	75513E101	27,718	387,615	SH		OTR	1	355,378	16,375	15,862
RAYTHEON TECHNOLOGIES CORP	COMMON STOCK	75513E101	415	5,801	SH		DFND	5	5,708	0	93
GLOBE LIFE INC	COMMON STOCK	37959E102	2,372	24,979	SH		DFND	1	24,033	0	946
GLOBE LIFE INC	COMMON STOCK	37959E102	1,523	16,040	SH		OTR	1	16,040	0	0
GLOBE LIFE INC	COMMON STOCK	37959E102	3	29	SH		DFND	5	29	0	0
LUMINEX CORP	COMMON STOCK	55027E102	159	6,878	SH		DFND	1	6,806	0	72
LUMINEX CORP	COMMON STOCK	55027E102	32	1,363	SH		OTR	1	74	1,289	0
LUMINEX CORPORATION	COMMON STOCK	55027E102	7	312	SH		DFND	5	312	0	0
NICOLET BANKSHARES INC	COMMON STOCK	65406E102	4	65	SH		DFND	1	65	0	0
PROTAGONIST THERAPEUTICS INC	COMMON STOCK	74366E102	9	428	SH		DFND	1	61	0	367
QUANTA SVCS INC	COMMON STOCK	74762E102	483	6,714	SH		DFND	1	4,832	0	1,882
QUANTA SVCS INC	COMMON STOCK	74762E102	57	794	SH		OTR	1	786	0	8
QUANTA SERVICES INC	COMMON STOCK	74762E102	3	48	SH		DFND	5	48	0	0
VERISIGN INC	COMMON STOCK	92343E102	1,351	6,244	SH		DFND	1	5,253	0	991
VERISIGN INC	COMMON STOCK	92343E102	23	106	SH		OTR	1	0	94	12
VERISIGN INC	COMMON STOCK	92343E102	25	114	SH		DFND	5	114	0	0
BLACKROCK MUNIYIELD QUALITY FU	OTHER	09254E103	103	7,150	SH		DFND	1	7,150	0	0
CNO FINANCIAL GROUP INC	COMMON STOCK	12621E103	54	2,423	SH		DFND	1	987	0	1,436
CNO FINANCIAL GROUP INC	COMMON STOCK	12621E103	68	3,082	SH		OTR	1	0	3,031	51
HCI GROUP INC	COMMON STOCK	40416E103	6	112	SH		DFND	1	112	0	0
INVESCO QQQ TRUST ETF	OTHER	46090E103	529,093	1,686,406	SH		DFND	1	1,673,601	0	12,805
INVESCO QQQ TRUST ETF	OTHER	46090E103	95,606	304,729	SH		OTR	1	294,635	1,961	8,133
INVESCO QQQ TRUST SERIES 1	OTHER	46090E103	14,672	46,764	SH		DFND	5	1,549	0	45,215
INVESCO QQQ TRUST ETF	OTHER	46090E103	7	25	SH	Put	DFND	1	25	0	0
INVESCO ADVANTAGE MUNI INC TR	OTHER	46132E103	19	1,694	SH		DFND	1	1,694	0	0
INVESCO ADVANTAGE MUNI INC TR	OTHER	46132E103	4	353	SH		OTR	1	353	0	0
PROGYNY INC	COMMON STOCK	74340E103	0	6	SH		DFND	1	6	0	0
PROGYNY INC	COMMON STOCK	74340E103	30	717	SH		OTR	1	0	717	0
US SILICA HOLDINGS INC	COMMON STOCK	90346E103	3	386	SH		DFND	1	0	0	386
EVO PAYMENTS INC CLASS A	COMMON STOCK	26927E104	137	5,057	SH		DFND	1	0	0	5,057
EVO PAYMENTS INC CLASS A	COMMON STOCK	26927E104	80	2,957	SH		OTR	1	0	2,957	0
AMERISOURCEBERGEN CORP	COMMON STOCK	03073E105	4,494	45,976	SH		DFND	1	44,568	0	1,408
AMERISOURCEBERGEN CORP	COMMON STOCK	03073E105	995	10,172	SH		OTR	1	9,820	166	186
AMERISOURCEBERGEN CORP	COMMON STOCK	03073E105	281	2,875	SH		DFND	5	2,875	0	0
FIRST TRUST CAPITAL STRENGTH	OTHER	33733E104	5,054	74,874	SH		DFND	5	22,907	0	51,967
APOLLO GLOBAL MANAGEMENT INC	COMMON STOCK	03768E105	15	300	SH		DFND	1	300	0	0
APOLLO GLOBAL MANAGEMENT INC	COMMON STOCK	03768E105	62	1,271	SH		OTR	1	1,271	0	0
BLACK DIAMOND THERAPEUTICS I	COMMON STOCK	09203E105	22	700	SH		DFND	1	700	0	0
CARS COM INC W I	COMMON STOCK	14575E105	2	162	SH		DFND	1	162	0	0
CARS COM INC W I	COMMON STOCK	14575E105	0	33	SH		OTR	1	33	0	0
AMERICAN EAGLE OUTFITTERS	COMMON STOCK	02553E106	95	4,711	SH		DFND	1	198	0	4,513
ITAU CORPBANCA A D R	COMMON STOCK	45033E105	3	669	SH		DFND	1	574	0	95
UNIVERSAL HLTH RLTY INCOME TR	COMMON STOCK	91359E105	1	20	SH		DFND	1	0	0	20
VALE SA SP A D R	COMMON STOCK	91912E105	269	16,065	SH		DFND	1	8,000	0	8,065
VALE SA SP A D R	COMMON STOCK	91912E105	18	1,075	SH		OTR	1	1,000	0	75
FLAHERTY & CRUMRINE	OTHER	33848E106	433	33,014	SH		DFND	5	14,400	0	18,614
TARGET CORP	COMMON STOCK	87612E106	241,785	1,369,652	SH		DFND	1	1,343,017	0	26,635
TARGET CORP	COMMON STOCK	87612E106	82,644	468,160	SH		OTR	1	446,358	9,564	12,238
TARGET CORPORATION	COMMON STOCK	87612E106	174	986	SH		DFND	5	986	0	0
AMERESCO INC CL A	COMMON STOCK	02361E108	1	10	SH		DFND	1	0	0	10
AMERESCO INC CL A	COMMON STOCK	02361E108	6	114	SH		OTR	1	114	0	0
ESSENTIAL PROPERTIES REALTY	COMMON STOCK	29670E107	25	1,179	SH		DFND	1	459	0	720
RENASANT CORP	COMMON STOCK	75970E107	488	14,496	SH		DFND	1	14,479	0	17
RENASANT CORP	COMMON STOCK	75970E107	54	1,598	SH		OTR	1	1,598	0	0
BLACKROCK TCP CAPITAL CORP	COMMON STOCK	09259E108	15	1,315	SH		DFND	1	1,315	0	0
EDWARDS LIFESCIENCES CORP	COMMON STOCK	28176E108	51,407	563,486	SH		DFND	1	552,105	0	11,381
EDWARDS LIFESCIENCES CORP	COMMON STOCK	28176E108	14,197	155,621	SH		OTR	1	145,675	8,335	1,611
EDWARDS LIFESCIENCE CORP	COMMON STOCK	28176E108	183	2,008	SH		DFND	5	1,882	0	126
FIRST TRUST PREFERRED SEC INC	OTHER	33739E108	399	19,771	SH		DFND	1	19,771	0	0
FIRST TRUST PREFERRED SEC INC	OTHER	33739E108	15	727	SH		OTR	1	727	0	0
FT PREFERRED SECURITIES & INC	OTHER	33739E108	2,155	106,714	SH		DFND	5	21,149	0	85,565
ARGAN INC	COMMON STOCK	04010E109	369	8,300	SH		DFND	1	8,300	0	0
NCR CORPORATION	COMMON STOCK	62886E108	1,586	42,219	SH		DFND	1	42,123	0	96
NCR CORPORATION	COMMON STOCK	62886E108	360	9,567	SH		OTR	1	6,954	2,532	81
NCR CORPORATION	COMMON STOCK	62886E108	23	618	SH		DFND	5	618	0	0
NAVISTAR INTERNATIONAL	COMMON STOCK	63934E108	47	1,072	SH		DFND	1	1,072	0	0
NEXTCURE INC	COMMON STOCK	65343E108	1	107	SH		DFND	1	0	0	107
BEYOND MEAT INC	COMMON STOCK	08862E109	647	5,170	SH		DFND	1	3,835	0	1,335
BEYOND MEAT INC	COMMON STOCK	08862E109	563	4,507	SH		OTR	1	406	3,974	127
BEYOND MEAT INC	COMMON STOCK	08862E109	9	74	SH		DFND	5	74	0	0
EVENTBRITE INC CLASS A	COMMON STOCK	29975E109	2	133	SH		DFND	1	133	0	0
FIRST TR HIGH INCOME LONG SHOR	OTHER	33738E109	6	432	SH		DFND	1	432	0	0
FORTINET INC	COMMON STOCK	34959E109	6,899	46,444	SH		DFND	1	43,550	0	2,894
FORTINET INC	COMMON STOCK	34959E109	1,932	13,010	SH		OTR	1	9,630	249	3,131
FORTINET INC	COMMON STOCK	34959E109	71	481	SH		DFND	5	467	0	14
IMAX CORP	COMMON STOCK	45245E109	39	2,163	SH		OTR	1	0	2,163	0
PENNANT GROUP INC THE	COMMON STOCK	70805E109	343	5,894	SH		DFND	1	5,867	0	27
PENNANT GROUP INC THE	COMMON STOCK	70805E109	49	839	SH		OTR	1	839	0	0
TOTAL SE A D R	COMMON STOCK	89151E109	1,667	39,767	SH		DFND	1	31,815	0	7,952
TOTAL SE A D R	COMMON STOCK	89151E109	1,380	32,939	SH		OTR	1	27,321	1,683	3,935
TRAVELERS COS INC	COMMON STOCK	89417E109	65,736	468,305	SH		DFND	1	465,567	0	2,738
TRAVELERS COS INC	COMMON STOCK	89417E109	32,990	235,020	SH		OTR	1	227,060	4,372	3,588
TRAVELERS COMPANY INC (THE)	COMMON STOCK	89417E109	15	110	SH		DFND	5	110	0	0
TREVENA INC	COMMON STOCK	89532E109	11	5,000	SH		DFND	1	5,000	0	0
INVESCO S P SMALLCAP INFORMATI	OTHER	46138E115	173	1,413	SH		DFND	1	1,413	0	0
INVESCO S P SMALLCAP INDUSTRIA	OTHER	46138E123	159	2,000	SH		DFND	1	2,000	0	0
INVESCO S P SMALLCAP HEALTH CA	OTHER	46138E149	15	91	SH		DFND	1	91	0	0
INVESCO S P SMALLCAP FINANCIAL	OTHER	46138E156	1	20	SH		DFND	1	20	0	0
INVESCO S P SMALLCAP CONSUMER	OTHER	46138E172	35	400	SH		DFND	1	400	0	0
INVESCO POWERSHARES S&P MIDCAP	OTHER	46138E198	211	4,376	SH		DFND	5	352	0	4,024
BRAINSTORM CELL THERAPEUTICS	COMMON STOCK	10501E201	14	3,000	SH		OTR	1	3,000	0	0
FIRSTSERVICE CORP	COMMON STOCK	33767E202	89	653	SH		DFND	1	536	0	117
FIRSTSERVICE CORP	COMMON STOCK	33767E202	276	2,019	SH		OTR	1	100	0	1,919
GRUPO AEROPORTUARIO SURE SPON	COMMON STOCK	40051E202	75	452	SH		DFND	1	52	0	400
GRUPO AEROPORTUARIO SURE SPON	COMMON STOCK	40051E202	1	4	SH		OTR	1	0	0	4
FIRST TRUST NYSE ARCA BIOTEC E	OTHER	33733E203	24	145	SH		DFND	1	145	0	0
FIRST TRUST AMEX BIO INDEX	OTHER	33733E203	217	1,292	SH		DFND	5	682	0	610
BARCLAYS PLC A D R	COMMON STOCK	06738E204	706	88,314	SH		DFND	1	60,787	0	27,527
BARCLAYS PLC A D R	COMMON STOCK	06738E204	102	12,762	SH		OTR	1	9,644	2,655	463
SPDR GOLD MINISHARES ETF	OTHER	98149E204	10	525	SH		DFND	1	525	0	0
ISHARES IBONDS DEC 2029 TERM C	OTHER	46436E205	3,110	113,146	SH		DFND	1	111,751	0	1,395
INVESCO CALIFORNIA AMT FREE MU	OTHER	46138E206	110	3,944	SH		DFND	1	3,944	0	0
NOVAGOLD RESOURCES INC	COMMON STOCK	66987E206	18	1,911	SH		DFND	1	1,340	0	571
B H P BILLITON PLC SPON A D R	COMMON STOCK	05545E209	304	5,725	SH		DFND	1	5,580	0	145
INVESCO POWERSHARES S&P INTL	OTHER	46138E230	20	673	SH		DFND	5	0	0	673
INVESCO S P GLOBAL WATER INDEX	OTHER	46138E263	1,870	39,952	SH		DFND	1	39,952	0	0
INVESCO S P GLOBAL WATER INDEX	OTHER	46138E263	104	2,224	SH		OTR	1	2,224	0	0
TRIUMPH BANCORP INC	COMMON STOCK	89679E300	1,398	28,788	SH		DFND	1	28,788	0	0
TRIUMPH BANCORP INC	COMMON STOCK	89679E300	25	513	SH		OTR	1	513	0	0
FIRST TRUST DOW JONES INTERNET	OTHER	33733E302	658	3,099	SH		DFND	1	1,650	0	1,449
FIRST TRUST DOW JONES INTERNET	OTHER	33733E302	145	682	SH		OTR	1	182	0	500
FIRST TRUST DOW JONES INTERNET	OTHER	33733E302	6,187	29,153	SH		DFND	5	15,445	0	13,708
GREENPOWER MTR CO INC	COMMON STOCK	39540E302	29	1,000	SH		DFND	1	1,000	0	0
ACCO BRANDS CORP	COMMON STOCK	00081T108	25	3,014	SH		DFND	1	1,442	0	1,572
ACCO BRANDS CORP	COMMON STOCK	00081T108	1	125	SH		OTR	1	0	125	0
ADT INC	COMMON STOCK	00090Q103	16	2,068	SH		DFND	1	0	0	2,068
AGNC INVESTMENT CORP	COMMON STOCK	00123Q104	540	34,611	SH		DFND	1	28,672	0	5,939
AGNC INVESTMENT CORP	COMMON STOCK	00123Q104	5	352	SH		DFND	5	352	0	0
A E S CORP	COMMON STOCK	00130H105	633	26,946	SH		DFND	1	23,754	0	3,192
A E S CORP	COMMON STOCK	00130H105	16	696	SH		OTR	1	0	296	400
AES CORPORATION	COMMON STOCK	00130H105	10	427	SH		DFND	5	427	0	0
ALERIAN MLP ETF	OTHER	00162Q452	19	758	SH		DFND	1	758	0	0
ALERIAN MLP ETF	OTHER	00162Q452	14	560	SH		OTR	1	160	0	400
ALPS CLEAN ENERGY ALPS ETF	OTHER	00162Q460	126	1,569	SH		DFND	1	1,569	0	0
ALPS CLEAN ENERGY ALPS ETF	OTHER	00162Q460	699	8,726	SH		OTR	1	8,726	0	0
ALPS MEDICAL BREAKTHROUGHS ETF	OTHER	00162Q593	33	645	SH		DFND	1	645	0	0
ALPS MEDICAL BREAKTHROUGHS ETF	OTHER	00162Q593	18	355	SH		OTR	1	355	0	0
AMC NETWORKS INC A W I	COMMON STOCK	00164V103	39	1,104	SH		DFND	1	816	0	288
ASGN INC	COMMON STOCK	00191U102	1,701	20,368	SH		DFND	1	15,550	0	4,818
ASGN INC	COMMON STOCK	00191U102	125	1,497	SH		OTR	1	1,473	0	24
AT T INC	COMMON STOCK	00206R102	86,356	3,002,640	SH		DFND	1	2,894,259	0	108,381
AT T INC	COMMON STOCK	00206R102	36,242	1,260,128	SH		OTR	1	1,180,451	32,978	46,699
AT&T INC	COMMON STOCK	00206R102	456	15,859	SH		DFND	5	15,577	0	282
AT T INC	COMMON STOCK	00206R102	2	25	SH	Put	DFND	1	25	0	0
ARK INNOVATION ETF	OTHER	00214Q104	704	5,658	SH		DFND	1	5,658	0	0
ARK GENOMIC REVOLUTION ETF	OTHER	00214Q302	211	2,264	SH		DFND	1	2,264	0	0
ARK FINTECH INNOVATION ETF	OTHER	00214Q708	11	215	SH		DFND	1	215	0	0
ATN INTERNATIONAL INC	COMMON STOCK	00215F107	1	21	SH		DFND	1	0	0	21
ASE TECHNOLOGY HOLDING CO LTD	COMMON STOCK	00215W100	397	67,932	SH		DFND	1	63,929	0	4,003
ASE TECHNOLOGY HOLDING CO LTD	COMMON STOCK	00215W100	65	11,122	SH		OTR	1	10,940	0	182
ASE TECHNOLOGY HOLDING CO LTD	COMMON STOCK	00215W100	3	507	SH		DFND	5	507	0	0
AARONS CO INC	COMMON STOCK	00258W108	45	2,417	SH		DFND	1	2,242	0	175
AARONS CO INC	COMMON STOCK	00258W108	5	231	SH		OTR	1	92	113	26
AARONS COMPANY INC (THE)	COMMON STOCK	00258W108	4	237	SH		DFND	5	237	0	0
ABBVIE INC	COMMON STOCK	00287Y109	199,176	1,858,848	SH		DFND	1	1,812,541	0	46,307
ABBVIE INC	COMMON STOCK	00287Y109	88,331	824,375	SH		OTR	1	777,709	29,596	17,070
ABBVIE INC	COMMON STOCK	00287Y109	920	8,584	SH		DFND	5	8,511	0	73
ABBVIE INC	COMMON STOCK	00287Y109	0	5	SH	Put	DFND	1	5	0	0
ABBVIE INC	COMMON STOCK	00287Y109	1	14	SH	Put	OTR	1	14	0	0
ABERDEEN EMERGING MARKETS EQUI	OTHER	00301W105	28	3,395	SH		DFND	1	3,395	0	0
ABERDEEN TOTAL DYNAMIC DIVID	OTHER	00326L100	34	3,838	SH		DFND	1	3,838	0	0
ACACIA COMMUNICATIONS INC	COMMON STOCK	00401C108	87	1,199	SH		DFND	1	248	0	951
ACADIA HEALTHCARE CO INC	COMMON STOCK	00404A109	687	13,682	SH		DFND	1	12,414	0	1,268
ACADIA HEALTHCARE CO INC	COMMON STOCK	00404A109	69	1,377	SH		OTR	1	182	1,095	100
ACADIA HEALTHCARE COMPANY INC	COMMON STOCK	00404A109	59	1,178	SH		DFND	5	1,178	0	0
ACCELERATE DIAGNOSTICS INC	COMMON STOCK	00430H102	1	94	SH		DFND	1	0	0	94
ACCELERON PHARMA INC	COMMON STOCK	00434H108	15	120	SH		DFND	1	74	0	46
ACCELERON PHARMA INC	COMMON STOCK	00434H108	203	1,587	SH		OTR	1	0	59	1,528
ACTIVISION BLIZZARD INC	COMMON STOCK	00507V109	6,367	68,568	SH		DFND	1	60,937	0	7,631
ACTIVISION BLIZZARD INC	COMMON STOCK	00507V109	1,394	15,010	SH		OTR	1	12,152	680	2,178
ACTIVISION BLIZZARD INC	COMMON STOCK	00507V109	125	1,345	SH		DFND	5	1,345	0	0
ACUITY BRANDS INC	COMMON STOCK	00508Y102	253	2,089	SH		DFND	1	1,934	0	155
ACUITY BRANDS INC	COMMON STOCK	00508Y102	480	3,964	SH		OTR	1	12	12	3,940
ADAMS NATURAL RESOURCES	OTHER	00548F105	71	6,250	SH		DFND	1	6,250	0	0
ADAPTIVE BIOTECHNOLOGIES	COMMON STOCK	00650F109	1,442	24,380	SH		DFND	1	24,380	0	0
ADAPTIVE BIOTECHNOLOGIES	COMMON STOCK	00650F109	13	228	SH		DFND	5	228	0	0
ADAPTIMMUNE THERAPEUTICS A D R	COMMON STOCK	00653A107	1	276	SH		DFND	1	276	0	0
ADAPTIMMUNE THERAPEUTICS A D R	COMMON STOCK	00653A107	6	1,160	SH		OTR	1	0	1,160	0
ADAPTHEALTH CORP	COMMON STOCK	00653Q102	68	1,820	SH		OTR	1	0	1,820	0
ADOBE INC	COMMON STOCK	00724F101	26,062	52,111	SH		DFND	1	37,601	0	14,510
ADOBE INC	COMMON STOCK	00724F101	7,931	15,858	SH		OTR	1	8,525	2,729	4,604
ADOBE INC	COMMON STOCK	00724F101	1,025	2,050	SH		DFND	5	2,028	0	22
ADTALEM GLOBAL EDUCATION INC	COMMON STOCK	00737L103	21	626	SH		DFND	1	397	0	229
ADTALEM GLOBAL EDUCATION INC	COMMON STOCK	00737L103	19	562	SH		OTR	1	562	0	0
ADTRAN INC	COMMON STOCK	00738A106	11	742	SH		DFND	1	742	0	0
ADVANCE AUTO PARTS INC	COMMON STOCK	00751Y106	217	1,378	SH		DFND	1	880	0	498
ADVANCE AUTO PARTS INC	COMMON STOCK	00751Y106	97	616	SH		OTR	1	548	68	0
ADVANCE AUTO PARTS	COMMON STOCK	00751Y106	314	1,993	SH		DFND	5	1,993	0	0
ADVENT CONVERTIBLE AND INCOME	OTHER	00764C109	11	687	SH		OTR	1	687	0	0
AECOM	COMMON STOCK	00766T100	89	1,789	SH		DFND	1	1,228	0	561
AECOM	COMMON STOCK	00766T100	7	133	SH		OTR	1	133	0	0
AEGION CORP	COMMON STOCK	00770F104	2	117	SH		DFND	1	117	0	0
AERIE PHARMACEUTICALS INC	COMMON STOCK	00771V108	29	2,119	SH		DFND	1	1,500	0	619
AERIE PHARMACEUTICALS INC	COMMON STOCK	00771V108	1	65	SH		OTR	1	65	0	0
ADVANSIX INC	COMMON STOCK	00773T101	189	9,458	SH		DFND	1	9,456	0	2
ADVANSIX INC	COMMON STOCK	00773T101	16	770	SH		OTR	1	731	9	30
ADVERUM BIOTECHNOLOGIES INC	COMMON STOCK	00773U108	2	193	SH		DFND	1	193	0	0
ADVANCED DRAINAGE SYSTEMS IN	COMMON STOCK	00790R104	31	373	SH		DFND	1	373	0	0
ADVANCED DRAINAGE SYSTEMS IN	COMMON STOCK	00790R104	50	598	SH		OTR	1	0	598	0
ADVANCED DRAINAGE SYSTEMS INC	COMMON STOCK	00790R104	1	9	SH		DFND	5	9	0	0
AERPIO PHARMACEUTICALS INC	COMMON STOCK	00810B105	49	50,000	SH		DFND	1	50,000	0	0
AGILENT TECHNOLOGIES INC	COMMON STOCK	00846U101	4,525	38,190	SH		DFND	1	34,971	0	3,219
AGILENT TECHNOLOGIES INC	COMMON STOCK	00846U101	4,131	34,865	SH		OTR	1	21,696	9,122	4,047
AGILENT TECHNOLOGIES INC	COMMON STOCK	00846U101	43	365	SH		DFND	5	342	0	23
AGIOS PHARMACEUTICALS INC	COMMON STOCK	00847X104	67	1,541	SH		DFND	1	997	0	544
AGIOS PHARMACEUTICALS INC	COMMON STOCK	00847X104	140	3,232	SH		OTR	1	3,047	185	0
AILERON THERAPEUTICS INC	COMMON STOCK	00887A105	26	25,000	SH		DFND	1	25,000	0	0
AIR LEASE CORP	COMMON STOCK	00912X302	864	19,442	SH		DFND	1	18,097	0	1,345
AIR LEASE CORP	COMMON STOCK	00912X302	411	9,255	SH		OTR	1	8,964	134	157
AIR LEASE CORP	COMMON STOCK	00912X302	21	464	SH		DFND	5	464	0	0
AIR TRANSPORT SERVICES GROUP	COMMON STOCK	00922R105	2	79	SH		DFND	1	79	0	0
AIR TRANSPORT SERVICES GROUP	COMMON STOCK	00922R105	10	328	SH		DFND	5	328	0	0
AIRGAIN INC	COMMON STOCK	00938A104	1,160	65,241	SH		DFND	1	65,241	0	0
AKAMAI TECHNOLOGIES INC	COMMON STOCK	00971T101	5,370	51,141	SH		DFND	1	47,975	0	3,166
AKAMAI TECHNOLOGIES INC	COMMON STOCK	00971T101	2,133	20,318	SH		OTR	1	19,848	367	103
AKAMAI TECHNOLOGIES INC	COMMON STOCK	00971T101	460	4,377	SH		DFND	5	4,363	0	14
AKEBIA THERAPEUTICS INC	COMMON STOCK	00972D105	6	2,000	SH		OTR	1	2,000	0	0
AKOUSTIS TECHNOLOGIES INC	COMMON STOCK	00973N102	49	4,000	SH		DFND	1	4,000	0	0
ALBIREO PHARMA INC	COMMON STOCK	01345P106	10	265	SH		DFND	1	38	0	227
ALBIREO PHARMA INC	COMMON STOCK	01345P106	70	1,867	SH		OTR	1	1,867	0	0
ALERUS FINANCIAL CORP	COMMON STOCK	01446U103	396	14,455	SH		DFND	1	14,455	0	0
ALEXCO RESOURCE CORP	COMMON STOCK	01535P106	1	160	SH		OTR	1	160	0	0
ALIBABA GROUP HOLDING LTD A D	COMMON STOCK	01609W102	17,353	74,563	SH		DFND	1	68,306	0	6,257
ALIBABA GROUP HOLDING LTD A D	COMMON STOCK	01609W102	5,873	25,234	SH		OTR	1	15,319	3,354	6,561
ALIBABA GROUP HOLDING LTD	COMMON STOCK	01609W102	489	2,100	SH		DFND	5	2,077	0	23
ALLAKOS INC	COMMON STOCK	01671P100	9	63	SH		DFND	1	31	0	32
ALLEGHENY TECHNOLOGIES INC	COMMON STOCK	01741R102	141	8,387	SH		DFND	1	8,252	0	135
ALLEGHENY TECHNOLOGIES INC	COMMON STOCK	01741R102	4	262	SH		OTR	1	262	0	0
ALLEGIANT TRAVEL CO	COMMON STOCK	01748X102	197	1,039	SH		DFND	1	76	0	963
ALLIANCEBERNSTEIN NATIONAL MUN	OTHER	01864U106	184	12,787	SH		OTR	1	12,787	0	0
ALLIANCE RESOURCE PARTNERS L P	COMMON STOCK	01877R108	15	3,300	SH		DFND	1	3,300	0	0
ALLIANCEBERNSTEIN GL HIGH INCO	OTHER	01879R106	48	4,100	SH		DFND	1	4,100	0	0
ALLIANCEBERNSTEIN GL HIGH INCO	OTHER	01879R106	105	8,933	SH		OTR	1	5,921	0	3,012
ALLIANCEBERNSTEIN HLDG LP	COMMON STOCK	01881G106	19	550	SH		DFND	1	550	0	0
ALLIANCEBERNSTEIN HLDG LP	COMMON STOCK	01881G106	123	3,650	SH		OTR	1	1,400	0	2,250
ALLIANZGI NFJ DIV INTEREST PRE	OTHER	01883A107	8	570	SH		DFND	1	570	0	0
ALLIANZGI NFJ DIV INTEREST PRE	OTHER	01883A107	37	2,775	SH		OTR	1	2,775	0	0
ALLISON TRANSMISSION HOLDINGS	COMMON STOCK	01973R101	296	6,850	SH		DFND	1	5,851	0	999
ALLISON TRANSMISSION HOLDINGS	COMMON STOCK	01973R101	633	14,670	SH		OTR	1	1,655	1,535	11,480
ALLISON TRANSMISSION HLGDS INC	COMMON STOCK	01973R101	35	814	SH		DFND	5	814	0	0
ALLY FINANCIAL INC	COMMON STOCK	02005N100	610	17,099	SH		DFND	1	7,751	0	9,348
ALLY FINANCIAL INC	COMMON STOCK	02005N100	14	404	SH		OTR	1	0	101	303
ALLY FINANCIAL INC	COMMON STOCK	02005N100	4	111	SH		DFND	5	111	0	0
ALNYLAM PHARMACEUTICALS INC	COMMON STOCK	02043Q107	236	1,816	SH		DFND	1	1,499	0	317
ALNYLAM PHARMACEUTICALS INC	COMMON STOCK	02043Q107	32	242	SH		OTR	1	14	119	109
ALPHABET INC CL C	COMMON STOCK	02079K107	158,969	90,742	SH		DFND	1	85,843	0	4,899
ALPHABET INC CL C	COMMON STOCK	02079K107	50,062	28,576	SH		OTR	1	24,180	1,255	3,141
ALPHABET INC	COMMON STOCK	02079K107	445	254	SH		DFND	5	245	0	9
ALPHABET INC CL A	COMMON STOCK	02079K305	399,291	227,823	SH		DFND	1	220,512	0	7,311
ALPHABET INC CL A	COMMON STOCK	02079K305	87,940	50,176	SH		OTR	1	43,252	2,514	4,410
ALPHABET INC	COMMON STOCK	02079K305	712	406	SH		DFND	5	404	0	2
ALPHABET INC CL A	COMMON STOCK	02079K305	0	1	SH	Put	DFND	1	1	0	0
ALTERYX INC CLASS A	COMMON STOCK	02156B103	1,897	15,573	SH		DFND	1	15,476	0	97
ALTERYX INC CLASS A	COMMON STOCK	02156B103	81	669	SH		OTR	1	609	51	9
ALTERYX LLC	COMMON STOCK	02156B103	17	139	SH		DFND	5	139	0	0
ALTICE USA INC A	COMMON STOCK	02156K103	84	2,216	SH		DFND	1	1,341	0	875
ALTICE USA INC A	COMMON STOCK	02156K103	6	161	SH		OTR	1	0	0	161
ALTICE USA INC	COMMON STOCK	02156K103	2	51	SH		DFND	5	0	0	51
ALTRA INDUSTRIAL MOTION CORP	COMMON STOCK	02208R106	1,987	35,841	SH		DFND	1	32,830	0	3,011
ALTRA INDUSTRIAL MOTION CORP	COMMON STOCK	02208R106	93	1,681	SH		OTR	1	962	719	0
ALTRIA GROUP INC	COMMON STOCK	02209S103	17,623	429,832	SH		DFND	1	412,449	0	17,383
ALTRIA GROUP INC	COMMON STOCK	02209S103	16,306	397,684	SH		OTR	1	378,482	12,286	6,916
ALTRIA GROUP INC	COMMON STOCK	02209S103	157	3,829	SH		DFND	5	3,829	0	0
AMBEV SA SPN A D R	COMMON STOCK	02319V103	3,143	1,026,846	SH		DFND	1	1,009,312	0	17,534
AMBEV SA SPN A D R	COMMON STOCK	02319V103	585	191,169	SH		OTR	1	177,392	6,639	7,138
AMBEV SA	COMMON STOCK	02319V103	4	1,369	SH		DFND	5	1,369	0	0
AMERICA FIRST MULTIFAMILY IN L	COMMON STOCK	02364V107	558	131,213	SH		DFND	1	131,213	0	0
AMERICA MOVIL A D R	COMMON STOCK	02364W105	307	21,139	SH		DFND	1	20,619	0	520
AMERICA MOVIL A D R	COMMON STOCK	02364W105	1,264	86,949	SH		OTR	1	83,952	2,997	0
AMERICAN AIRLINES GROUP INC	COMMON STOCK	02376R102	614	38,960	SH		DFND	1	13,431	0	25,529
AMERICAN AIRLINES GROUP INC	COMMON STOCK	02376R102	124	7,837	SH		OTR	1	752	7,085	0
AMERICAN FINANCE TRUST INC	COMMON STOCK	02607T109	3	337	SH		DFND	1	337	0	0
AMERICAN HOMES 4 RENT A	COMMON STOCK	02665T306	245	8,178	SH		DFND	1	8,044	0	134
AMERICAN HOMES 4 RENT A	COMMON STOCK	02665T306	42	1,400	SH		OTR	1	1,327	0	73
AMERICAN HOMES 4 RENT	COMMON STOCK	02665T306	1	34	SH		DFND	5	34	0	0
AMERICAN NATIONAL GROUP INC CO	COMMON STOCK	02772A109	10	100	SH		DFND	1	100	0	0
AMERICAN OUTDOOR BRANDS INC	COMMON STOCK	02875D109	3	184	SH		DFND	1	184	0	0
AMERICAN PUBLIC EDUCATION	COMMON STOCK	02913V103	3	106	SH		DFND	1	88	0	18
AMERICAN TOWER CORP	COMMON STOCK	03027X100	204,331	910,321	SH		DFND	1	898,086	0	12,235
AMERICAN TOWER CORP	COMMON STOCK	03027X100	40,643	181,074	SH		OTR	1	161,575	7,593	11,906
AMERICAN TOWER CORP CL A	COMMON STOCK	03027X100	180	800	SH		DFND	5	800	0	0
AMERICAN WELL CORP	COMMON STOCK	03044L105	1	48	SH		DFND	1	48	0	0
AMERICAS SILVER CORP	COMMON STOCK	03062D100	46	14,286	SH		DFND	1	14,286	0	0
AMERICAS CAR MART INC	COMMON STOCK	03062T105	2	18	SH		DFND	1	18	0	0
AMERICOLD REALTY TRUST	COMMON STOCK	03064D108	1,819	48,740	SH		DFND	1	48,073	0	667
AMERICOLD REALTY TRUST	COMMON STOCK	03064D108	652	17,473	SH		OTR	1	15,294	81	2,098
AMERICOLD REALTY TRUST	COMMON STOCK	03064D108	48	1,281	SH		DFND	5	1,281	0	0
AMERISAFE INC	COMMON STOCK	03071H100	1	9	SH		DFND	1	9	0	0
AMERISAFE INC	COMMON STOCK	03071H100	7	117	SH		DFND	5	117	0	0
AMERIPRISE FINL INC	COMMON STOCK	03076C106	9,209	47,389	SH		DFND	1	45,656	0	1,733
AMERIPRISE FINL INC	COMMON STOCK	03076C106	4,870	25,059	SH		OTR	1	24,219	504	336
AMERIPRISE FINANCIAL INC	COMMON STOCK	03076C106	153	784	SH		DFND	5	784	0	0
AMERIS BANCORP	COMMON STOCK	03076K108	239	6,311	SH		DFND	1	5,887	0	424
AMERIS BANCORP	COMMON STOCK	03076K108	5	126	SH		OTR	1	126	0	0
AMICUS THERAPEUTICS INC	COMMON STOCK	03152W109	7	320	SH		DFND	1	111	0	209
AMNEAL PHARMACEUTICALS INC	COMMON STOCK	03168L105	4	822	SH		DFND	1	822	0	0
AMPHASTAR PHARMACEUTICALS IN	COMMON STOCK	03209R103	1	31	SH		DFND	1	0	0	31
AMPHASTAR PHARMACEUTICALS IN	COMMON STOCK	03209R103	23	1,161	SH		OTR	1	0	1,161	0
ANAPLAN INC	COMMON STOCK	03272L108	33	465	SH		DFND	1	457	0	8
ANAPLAN INC	COMMON STOCK	03272L108	108	1,496	SH		OTR	1	1,400	61	35
ANGEL OAK FINANCIAL STRATS INC	OTHER	03464A100	42	2,500	SH		DFND	1	2,500	0	0
ANGIODYNAMICS INC	COMMON STOCK	03475V101	46	2,992	SH		DFND	1	2,930	0	62
ANHEUSER BUSCH INBEV NV A D R	COMMON STOCK	03524A108	923	13,215	SH		DFND	1	9,290	0	3,925
ANHEUSER BUSCH INBEV NV A D R	COMMON STOCK	03524A108	963	13,774	SH		OTR	1	13,267	175	332
ANHEUSER BUSCH INBEV	COMMON STOCK	03524A108	276	3,940	SH		DFND	5	3,940	0	0
ANSYS INC	COMMON STOCK	03662Q105	11,598	31,880	SH		DFND	1	29,342	0	2,538
ANSYS INC	COMMON STOCK	03662Q105	1,567	4,309	SH		OTR	1	4,077	138	94
ANSYS INC	COMMON STOCK	03662Q105	922	2,534	SH		DFND	5	2,534	0	0
ANTERO RESOURCES CORP	COMMON STOCK	03674X106	4	740	SH		DFND	1	740	0	0
ANTERO MIDSTREAM CORP	COMMON STOCK	03676B102	0	64	SH		DFND	5	64	0	0
ANTERIX INC	COMMON STOCK	03676C100	7	183	SH		DFND	1	0	0	183
APARTMENT INVT MGMT CO	COMMON STOCK	03748R747	58	11,037	SH		DFND	1	10,877	0	160
APARTMENT INVT MGMT CO	COMMON STOCK	03748R747	21	4,097	SH		OTR	1	4,040	24	33
APARTMENT INVESTEMENT & MGMT	COMMON STOCK	03748R747	0	7	SH		DFND	5	7	0	0
APARTMENT INCOME REIT CORP	COMMON STOCK	03750L109	2,171	56,520	SH		DFND	1	56,360	0	160
APARTMENT INCOME REIT CORP	COMMON STOCK	03750L109	157	4,097	SH		OTR	1	4,040	24	33
APARTMENT INCOME REIT CORP	COMMON STOCK	03750L109	1	33	SH		DFND	5	33	0	0
APELLIS PHARMACEUTICALS INC	COMMON STOCK	03753U106	3	58	SH		DFND	1	58	0	0
APELLIS PHARMACEUTICALS INC	COMMON STOCK	03753U106	3	49	SH		OTR	1	0	23	26
APOLLO INVESTMENT CORP	COMMON STOCK	03761U502	224	21,026	SH		DFND	1	20,556	0	470
APOLLO COMMERCIAL REAL ESTAT	COMMON STOCK	03762U105	16	1,436	SH		DFND	1	1,175	0	261
APHRIA INC	COMMON STOCK	03765K104	6	900	SH		OTR	1	900	0	0
APPIAN CORP	COMMON STOCK	03782L101	209	1,290	SH		DFND	1	1,273	0	17
APPIAN CORP	COMMON STOCK	03782L101	15	91	SH		OTR	1	80	0	11
APPFOLIO INC A	COMMON STOCK	03783C100	35	193	SH		DFND	1	7	0	186
APPFOLIO INC A	COMMON STOCK	03783C100	18	100	SH		OTR	1	100	0	0
APPLE HOSPITALITY REIT INC	COMMON STOCK	03784Y200	53	4,098	SH		DFND	1	449	0	3,649
APPLE HOSPITALITY REIT INC	COMMON STOCK	03784Y200	62	4,835	SH		OTR	1	0	4,739	96
APPLIED INDL TECHNOLOGIES INC	COMMON STOCK	03820C105	4,116	52,778	SH		DFND	1	52,739	0	39
APTEVO THERAPEUTICS INC COM	COMMON STOCK	03835L207	0	1	SH		DFND	1	1	0	0
APTOSE BIOSCIENCES INC	COMMON STOCK	03835T200	133	30,300	SH		OTR	1	30,300	0	0
ARAMARK	COMMON STOCK	03852U106	83	2,160	SH		DFND	1	1,142	0	1,018
ARAMARK HOLDINGS CORP	COMMON STOCK	03852U106	26	687	SH		DFND	5	687	0	0
ARCBEST CORP	COMMON STOCK	03937C105	33	766	SH		DFND	1	414	0	352
ARCELORMITTAL CLASS A N Y REGI	COMMON STOCK	03938L203	246	10,760	SH		DFND	1	6,808	0	3,952
ARCELORMITTAL CLASS A N Y REGI	COMMON STOCK	03938L203	24	1,070	SH		OTR	1	423	629	18
ARCH RESOURCES INC	COMMON STOCK	03940R107	16	367	SH		DFND	1	179	0	188
ARCH RESOURCES INC	COMMON STOCK	03940R107	3	75	SH		OTR	1	0	75	0
ARCHROCK INC	COMMON STOCK	03957W106	43	4,997	SH		DFND	1	2,021	0	2,976
ARCONIC CORPORATION	COMMON STOCK	03966V107	154	5,151	SH		DFND	1	2,500	0	2,651
ARCONIC CORPORATION	COMMON STOCK	03966V107	14	482	SH		OTR	1	457	25	0
ARCONIC CORP	COMMON STOCK	03966V107	7	226	SH		DFND	5	226	0	0
ARCUS BIOSCIENCES INC	COMMON STOCK	03969F109	3	99	SH		DFND	1	0	0	99
ARCTURUS THERAPEUTICS HOLDIN	COMMON STOCK	03969T109	20	470	SH		OTR	1	470	0	0
ARES MANAGEMENT CORP A	COMMON STOCK	03990B101	331	7,038	SH		DFND	1	0	0	7,038
ARES MANAGEMENT CORP A	COMMON STOCK	03990B101	908	19,299	SH		OTR	1	0	19,299	0
ARES MANAGEMENT CORP	COMMON STOCK	03990B101	1	27	SH		DFND	5	27	0	0
ARES CAPITAL CORP	COMMON STOCK	04010L103	735	43,510	SH		DFND	1	41,895	0	1,615
ARES CAPITAL CORP	COMMON STOCK	04010L103	191	11,280	SH		OTR	1	11,280	0	0
ARES COMMERCIAL REAL ESTATE	COMMON STOCK	04013V108	4	338	SH		DFND	1	0	0	338
ARGENX SE A D R	COMMON STOCK	04016X101	150	508	SH		DFND	1	290	0	218
ARGENX SE A D R	COMMON STOCK	04016X101	149	508	SH		OTR	1	26	67	415
ARLO TECHNOLOGIES INC	COMMON STOCK	04206A101	4	533	SH		DFND	1	533	0	0
ARMADA HOFFLER PROPERTIES INC	COMMON STOCK	04208T108	11	969	SH		DFND	1	466	0	503
ARMSTRONG WORLD INDS INC	COMMON STOCK	04247X102	13	177	SH		DFND	1	129	0	48
ARRAY TECHNOLOGIES INC	COMMON STOCK	04271T100	413	9,585	SH		DFND	1	9,585	0	0
ARRAY TECHNOLOGIES INC	COMMON STOCK	04271T100	6	141	SH		OTR	1	141	0	0
ARROWHEAD PHARMACEUTICALS INC	COMMON STOCK	04280A100	13	164	SH		DFND	1	116	0	48
ARROWHEAD PHARMACEUTICALS INC	COMMON STOCK	04280A100	333	4,344	SH		OTR	1	0	0	4,344
ARTISAN PARTNERS ASSET MANAG	COMMON STOCK	04316A108	46	923	SH		DFND	1	169	0	754
ARTISAN PARTNERS ASST MGMT INC	COMMON STOCK	04316A108	28	546	SH		DFND	5	546	0	0
ARVINAS INC	COMMON STOCK	04335A105	12	147	SH		DFND	1	0	0	147
ASCENDIS PHARMA A S A D R	COMMON STOCK	04351P101	96	574	SH		DFND	1	495	0	79
ASCENDIS PHARMA A S A D R	COMMON STOCK	04351P101	15	89	SH		OTR	1	0	89	0
ASSURANT INC	COMMON STOCK	04621X108	162	1,193	SH		DFND	1	765	0	428
ASSURANT INC	COMMON STOCK	04621X108	1	5	SH		OTR	1	0	0	5
AT HOME GROUP INC	COMMON STOCK	04650Y100	22	1,438	SH		DFND	1	1,000	0	438
ATLANTIC UNION BANKSHARES CO	COMMON STOCK	04911A107	321	9,758	SH		DFND	1	9,740	0	18
ATLANTIC UNION BANKSHARES CO	COMMON STOCK	04911A107	73	2,204	SH		OTR	1	2,204	0	0
ATLANTIC UNION BANKSHARES CO	COMMON STOCK	04911A107	40	1,210	SH		DFND	5	1,210	0	0
ATRICURE INC	COMMON STOCK	04963C209	49	872	SH		DFND	1	872	0	0
ATRICURE INC	COMMON STOCK	04963C209	214	3,850	SH		OTR	1	34	1,064	2,752
ATRECA INC A	COMMON STOCK	04965G109	15	900	SH		DFND	1	900	0	0
AURINIA PHARMACEUTICALS INC	COMMON STOCK	05156V102	122	8,820	SH		OTR	1	8,820	0	0
AURORA CANNABIS INC COM	COMMON STOCK	05156X884	4	456	SH		DFND	1	456	0	0
AURORA CANNABIS INC COM	COMMON STOCK	05156X884	46	5,562	SH		OTR	1	5,562	0	0
AUTOHOME INC A D R	COMMON STOCK	05278C107	4,128	41,436	SH		DFND	1	40,870	0	566
AUTOHOME INC A D R	COMMON STOCK	05278C107	525	5,268	SH		OTR	1	5,259	9	0
AUTOLUS THERAPEUTICS PLC A D R	COMMON STOCK	05280R100	2	228	SH		DFND	1	121	0	107
AUTOLUS THERAPEUTICS PLC A D R	COMMON STOCK	05280R100	3	311	SH		OTR	1	0	311	0
AUTONATION INC	COMMON STOCK	05329W102	905	12,973	SH		DFND	1	11,930	0	1,043
AUTONATION INC	COMMON STOCK	05329W102	16	227	SH		OTR	1	0	227	0
AVALARA INC	COMMON STOCK	05338G106	86,289	523,315	SH		DFND	1	523,272	0	43
AVALARA INC	COMMON STOCK	05338G106	1,536	9,314	SH		OTR	1	5,063	4,228	23
AVALARA INC	COMMON STOCK	05338G106	195	1,185	SH		DFND	5	1,185	0	0
AVANOS MEDICAL INC	COMMON STOCK	05350V106	55	1,189	SH		DFND	1	788	0	401
AVANOS MEDICAL INC	COMMON STOCK	05350V106	145	3,180	SH		OTR	1	2,397	782	1
AVANGRID INC	COMMON STOCK	05351W103	137	3,010	SH		DFND	1	1,732	0	1,278
AVANGRID INC	COMMON STOCK	05351W103	143	3,149	SH		OTR	1	1,078	156	1,915
AVAYA HLDGS CORP	COMMON STOCK	05351X101	5	277	SH		DFND	1	0	0	277
AVANTOR INC	COMMON STOCK	05352A100	584	20,765	SH		DFND	1	7,390	0	13,375
AVANTOR INC	COMMON STOCK	05352A100	339	12,048	SH		OTR	1	8,113	3,255	680
AVANTOR INC	COMMON STOCK	05352A100	21	736	SH		DFND	5	736	0	0
AVIENT CORPORATION	COMMON STOCK	05368V106	95	2,351	SH		DFND	1	1,784	0	567
AVIENT CORPORATION	COMMON STOCK	05368V106	13	333	SH		OTR	1	230	103	0
AVISTA CORP	COMMON STOCK	05379B107	791	19,695	SH		DFND	1	19,638	0	57
AVISTA CORP	COMMON STOCK	05379B107	85	2,120	SH		OTR	1	2,101	19	0
AVISTA CORP	COMMON STOCK	05379B107	36	889	SH		DFND	5	889	0	0
AVROBIO INC	COMMON STOCK	05455M100	3	188	SH		OTR	1	0	188	0
AXON ENTERPRISE INC	COMMON STOCK	05464C101	7,718	62,988	SH		DFND	1	62,981	0	7
AXON ENTERPRISE INC	COMMON STOCK	05464C101	235	1,918	SH		OTR	1	95	0	1,823
AXSOME THERAPEUTICS INC	COMMON STOCK	05464T104	725	8,900	SH		DFND	1	8,878	0	22
AXSOME THERAPEUTICS INC	COMMON STOCK	05464T104	13	161	SH		OTR	1	161	0	0
AXOS FINANCIAL INC	COMMON STOCK	05465C100	125	3,321	SH		DFND	1	800	0	2,521
AXONICS MODULATION TECHNOLOG	COMMON STOCK	05465P101	4	82	SH		DFND	1	52	0	30
AXONICS MODULATION TECHNOLOG	COMMON STOCK	05465P101	33	653	SH		OTR	1	0	631	22
AZEK CO INC CL A	COMMON STOCK	05478C105	48	1,245	SH		DFND	1	1,245	0	0
AZIYO BIOLOGICS INC	COMMON STOCK	05479K106	9	625	SH		DFND	1	625	0	0
AZUL SA A D R	COMMON STOCK	05501U106	18	770	SH		DFND	1	1	0	769
AZUL SA A D R	COMMON STOCK	05501U106	1	57	SH		OTR	1	0	57	0
AZURRX BIOPHARMA INC COM	COMMON STOCK	05502L105	49	51,020	SH		OTR	1	51,020	0	0
B G FOODS INC NEW	COMMON STOCK	05508R106	172	6,191	SH		DFND	1	5,870	0	321
B G FOODS INC NEW	COMMON STOCK	05508R106	1,847	66,600	SH		OTR	1	66,500	0	100
BCE INC	COMMON STOCK	05534B760	226	5,267	SH		DFND	1	3,513	0	1,754
BCE INC	COMMON STOCK	05534B760	9	204	SH		OTR	1	203	0	1
BCE INC	COMMON STOCK	05534B760	302	7,056	SH		DFND	5	7,056	0	0
BGC PARTNERS INC CL A	COMMON STOCK	05541T101	28	7,038	SH		DFND	1	7,038	0	0
BJS WHOLESALE CLUB HOLDINGS IN	COMMON STOCK	05550J101	1,527	40,976	SH		DFND	1	39,430	0	1,546
BJS WHOLESALE CLUB HOLDINGS IN	COMMON STOCK	05550J101	59	1,573	SH		OTR	1	1,433	0	140
BJ'S WHOLESALE CLUB HOLDINGS	COMMON STOCK	05550J101	57	1,532	SH		DFND	5	1,532	0	0
B O K FINANCIAL CORP COM NEW	COMMON STOCK	05561Q201	113	1,656	SH		DFND	1	803	0	853
B O K FINANCIAL CORP COM NEW	COMMON STOCK	05561Q201	5	68	SH		OTR	1	24	44	0
BOK FINANCIAL CORPORATION	COMMON STOCK	05561Q201	40	586	SH		DFND	5	586	0	0
BP MIDSTREAM PARTNERS L P	COMMON STOCK	0556EL109	247	23,300	SH		DFND	1	23,300	0	0
BNY MELLON STRATEGIC MUNICIPAL	OTHER	05588W108	8	1,000	SH		DFND	1	1,000	0	0
BRP GROUP INC A	COMMON STOCK	05589G102	63	2,116	SH		OTR	1	0	2,116	0
BNY MELLON MUNICIPAL INCOME	OTHER	05589T104	9	1,000	SH		DFND	1	1,000	0	0
BMC STOCK HOLDINGS INC	COMMON STOCK	05591B109	20	372	SH		DFND	1	0	0	372
BWX TECHNOLOGIES INC	COMMON STOCK	05605H100	455	7,553	SH		DFND	1	3,146	0	4,407
BWX TECHNOLOGIES INC	COMMON STOCK	05605H100	34	572	SH		OTR	1	266	101	205
BWX TECHNOLOGIES INC	COMMON STOCK	05605H100	12	203	SH		DFND	5	203	0	0
BAKER HUGHES COMPANY	COMMON STOCK	05722G100	358	17,212	SH		DFND	1	13,215	0	3,997
BAKER HUGHES COMPANY	COMMON STOCK	05722G100	252	12,100	SH		OTR	1	11,910	93	97
BAKER HUGHES	COMMON STOCK	05722G100	38	1,841	SH		DFND	5	1,841	0	0
BALLY'S CORP	COMMON STOCK	05875B106	10	191	SH		DFND	5	191	0	0
BANCFIRST CORP	COMMON STOCK	05945F103	9	155	SH		DFND	1	29	0	126
BANCO BILBAO VIZCAYA ARGEN A D	COMMON STOCK	05946K101	4,376	886,064	SH		DFND	1	862,158	0	23,906
BANCO BILBAO VIZCAYA ARGEN A D	COMMON STOCK	05946K101	872	176,527	SH		OTR	1	166,477	9,241	809
BANCO BILBAO VIZCAYA	COMMON STOCK	05946K101	10	1,993	SH		DFND	5	1,993	0	0
BANCO SANTANDER SA A D R	COMMON STOCK	05964H105	428	140,549	SH		DFND	1	128,605	0	11,944
BANCO SANTANDER SA A D R	COMMON STOCK	05964H105	201	65,964	SH		OTR	1	50,838	14,054	1,072
BANCO SANTANDER CHILE A D R	COMMON STOCK	05965X109	19	993	SH		DFND	1	844	0	149
BANCO SANTANDER BRASIL SA A D	COMMON STOCK	05967A107	134	15,534	SH		DFND	1	8,111	0	7,423
BANCO SANTANDER BRASIL SA A D	COMMON STOCK	05967A107	1	155	SH		OTR	1	0	0	155
BANCOLOMBIA SA A D R	COMMON STOCK	05968L102	285	7,091	SH		DFND	1	2,554	0	4,537
BANCOLOMBIA SA A D R	COMMON STOCK	05968L102	1	30	SH		OTR	1	0	30	0
BANCO SANTANDER MEXICO A D R	COMMON STOCK	05969B103	1	200	SH		DFND	1	200	0	0
BANCORPSOUTH BANK	COMMON STOCK	05971J102	133	4,868	SH		DFND	1	4,417	0	451
BANCORPSOUTH BANK	COMMON STOCK	05971J102	42	1,535	SH		OTR	1	0	1,522	13
BANDWIDTH INC CLASS A	COMMON STOCK	05988J103	1,265	8,233	SH		DFND	1	8,169	0	64
BANDWIDTH INC CLASS A	COMMON STOCK	05988J103	26	166	SH		OTR	1	166	0	0
BANC OF CALIFORNIA INC	COMMON STOCK	05990K106	9	617	SH		DFND	1	502	0	115
BANK FIRST CORP	COMMON STOCK	06211J100	721	11,126	SH		DFND	1	11,096	0	30
BANK OZK	COMMON STOCK	06417N103	70	2,241	SH		DFND	1	1,885	0	356
BANK OZK	COMMON STOCK	06417N103	62	1,988	SH		OTR	1	71	1,791	126
BANK OZK	COMMON STOCK	06417N103	13	401	SH		DFND	5	401	0	0
BANK OF COMMERCE HOLDINGS	COMMON STOCK	06424J103	3	300	SH		DFND	1	300	0	0
BANKUNITED INC	COMMON STOCK	06652K103	243	6,973	SH		DFND	1	82	0	6,891
BANKUNITED INC	COMMON STOCK	06652K103	3	88	SH		OTR	1	0	0	88
BANNER CORPORATION	COMMON STOCK	06652V208	17	357	SH		DFND	1	357	0	0
BANKWELL FINANCIAL GROUP INC	COMMON STOCK	06654A103	25	1,269	SH		DFND	1	1,269	0	0
BAOZUN INC SPON A D R	COMMON STOCK	06684L103	76	2,209	SH		DFND	1	1,673	0	536
IPATH DOW JONES UBS COMMODITY	OTHER	06738C778	1,170	53,432	SH		DFND	1	53,332	0	100
IPATH DOW JONES UBS COMMODITY	OTHER	06738C778	3,338	152,437	SH		OTR	1	152,437	0	0
IPATH S&P GSCI TOTAL RETURN ET	OTHER	06738C794	5,464	496,106	SH		DFND	1	491,388	0	4,718
IPATH S&P GSCI TOTAL RETURN ET	OTHER	06738C794	85	7,674	SH		OTR	1	7,674	0	0
IPATH SELECT MLP ETN	OTHER	06742C723	175	15,000	SH		OTR	1	15,000	0	0
IPATH B BLOOMBERG SUGAR TTL RE	OTHER	06746P449	2	49	SH		DFND	1	49	0	0
IPATH B BLOOMBERG GRAINS TTL R	OTHER	06746P522	5	100	SH		DFND	1	100	0	0
IPATH SERIES B BLOOMBERG ETN	OTHER	06746P563	4	109	SH		DFND	1	109	0	0
BARINGS BDC INC	COMMON STOCK	06759L103	2	249	SH		DFND	1	120	0	129
BARINGS BDC INC	COMMON STOCK	06759L103	1	86	SH		OTR	1	0	86	0
BAYCOM CORP	COMMON STOCK	07272M107	6	415	SH		DFND	1	415	0	0
BEAM THERAPEUTICS INC	COMMON STOCK	07373V105	82	1,000	SH		DFND	1	1,000	0	0
BEAZER HOMES USA INC	COMMON STOCK	07556Q881	8	558	SH		DFND	1	96	0	462
BEIGENE LTD A D R	COMMON STOCK	07725L102	386	1,494	SH		DFND	1	1,277	0	217
BEIGENE LTD A D R	COMMON STOCK	07725L102	217	841	SH		OTR	1	157	50	634
BENCHMARK ELECTRONICS INC	COMMON STOCK	08160H101	186	6,883	SH		DFND	1	6,289	0	594
BENCHMARK ELECTRONICS INC	COMMON STOCK	08160H101	94	3,488	SH		OTR	1	3,488	0	0
BENCHMARK ELECTRONICS INC	COMMON STOCK	08160H101	10	370	SH		DFND	5	370	0	0
BENEFITFOCUS INC	COMMON STOCK	08180D106	1	60	SH		DFND	1	0	0	60
BERRY GLOBAL GROUP INC	COMMON STOCK	08579W103	94	1,666	SH		DFND	1	1,166	0	500
BERRY GLOBAL GROUP INC	COMMON STOCK	08579W103	408	7,270	SH		OTR	1	2,775	157	4,338
BEST INC A D R	COMMON STOCK	08653C106	26	12,757	SH		DFND	1	3,340	0	9,417
BIOCRYST PHARMACEUTICALS INC	COMMON STOCK	09058V103	15	2,000	SH		DFND	1	2,000	0	0
BIODELIVERY SCIENCES INTL INC	COMMON STOCK	09060J106	1	200	SH		DFND	1	200	0	0
BIOMARIN PHARMACEUTICAL INC	COMMON STOCK	09061G101	856	9,762	SH		DFND	1	9,080	0	682
BIOMARIN PHARMACEUTICAL INC	COMMON STOCK	09061G101	3,708	42,284	SH		OTR	1	38,055	174	4,055
BIOMARIN PHARMACEUTICAL INC	COMMON STOCK	09061G101	221	2,522	SH		DFND	5	2,522	0	0
BIOLIFE SOLUTIONS INC	COMMON STOCK	09062W204	2	45	SH		OTR	1	45	0	0
BIOGEN, INC	COMMON STOCK	09062X103	11,266	46,013	SH		DFND	1	44,418	0	1,595
BIOGEN, INC	COMMON STOCK	09062X103	3,479	14,208	SH		OTR	1	11,614	145	2,449
BIOGEN INC	COMMON STOCK	09062X103	4	15	SH		DFND	5	15	0	0
BIOCEPT INC	COMMON STOCK	09072V501	2	500	SH		DFND	1	500	0	0
BIOCEPT INC	COMMON STOCK	09072V501	0	83	SH		OTR	1	83	0	0
BIO TECHNE CORP	COMMON STOCK	09073M104	8,761	27,592	SH		DFND	1	24,566	0	3,026
BIO TECHNE CORP	COMMON STOCK	09073M104	5,594	17,619	SH		OTR	1	16,689	32	898
BIO TECHNE CORP	COMMON STOCK	09073M104	306	963	SH		DFND	5	963	0	0
BIOSIG TECHNOLOGIES INC	COMMON STOCK	09073N201	9	2,404	SH		DFND	1	2,404	0	0
BIOXCEL THERAPEUTICS INC	COMMON STOCK	09075P105	236	5,110	SH		DFND	1	5,092	0	18
BIOXCEL THERAPEUTICS INC	COMMON STOCK	09075P105	2	51	SH		OTR	1	51	0	0
BIONTECH SE A D R	COMMON STOCK	09075V102	46	567	SH		DFND	1	567	0	0
BJS RESTAURANTS INC	COMMON STOCK	09180C106	5	135	SH		DFND	1	67	0	68
BJS RESTAURANTS INC	COMMON STOCK	09180C106	19	500	SH		OTR	1	500	0	0
BLACK KNIGHT INC	COMMON STOCK	09215C105	4,180	47,308	SH		DFND	1	46,840	0	468
BLACK KNIGHT INC	COMMON STOCK	09215C105	768	8,687	SH		OTR	1	5,608	2,854	225
BLACK KNIGHT INC	COMMON STOCK	09215C105	77	866	SH		DFND	5	866	0	0
BLACK STONE MINERALS L P	COMMON STOCK	09225M101	80	11,950	SH		DFND	1	0	0	11,950
BLACK STONE MINERALS L P	COMMON STOCK	09225M101	7	1,000	SH		OTR	1	1,000	0	0
BLACKBAUD INC	COMMON STOCK	09227Q100	108	1,882	SH		DFND	1	445	0	1,437
BLACKBAUD INC	COMMON STOCK	09227Q100	2	42	SH		OTR	1	0	0	42
BLACKBAUD INC	COMMON STOCK	09227Q100	1	11	SH		DFND	5	11	0	0
BLACKBERRY LTD	COMMON STOCK	09228F103	103	15,630	SH		DFND	1	12,875	0	2,755
BLACKBERRY LTD	COMMON STOCK	09228F103	16	2,506	SH		OTR	1	2,169	0	337
BLACKLINE INC	COMMON STOCK	09239B109	7,124	53,414	SH		DFND	1	51,155	0	2,259
BLACKLINE INC	COMMON STOCK	09239B109	1,229	9,214	SH		OTR	1	5,767	208	3,239
BLACKLINE INC	COMMON STOCK	09239B109	334	2,505	SH		DFND	5	2,505	0	0
BLACKROCK INVT QUALITY MUNI	OTHER	09247D105	170	9,900	SH		DFND	1	9,900	0	0
BLACKROCK INC	COMMON STOCK	09247X101	87,163	120,800	SH		DFND	1	115,015	0	5,785
BLACKROCK INC	COMMON STOCK	09247X101	16,991	23,548	SH		OTR	1	21,517	1,649	382
BLACKROCK INC	COMMON STOCK	09247X101	261	361	SH		DFND	5	361	0	0
BLACKROCK UTILITY INFRASTRUCTU	OTHER	09248D104	119	4,750	SH		DFND	1	4,750	0	0
BLACKROCK MUNICIPAL INCOME TRU	OTHER	09248F109	19	1,250	SH		DFND	1	1,250	0	0
BLACKROCK TAXABLE MUNI BD TRUS	OTHER	09248X100	24	900	SH		DFND	1	900	0	0
BLACKROCK MUN IN TRUST II	OTHER	09249N101	56	3,600	SH		DFND	1	3,600	0	0
BLACKROCK LTD DURATION INC	OTHER	09249W101	33	2,100	SH		DFND	1	2,100	0	0
BLACKROCK ENHANCED EQUITY DIVI	OTHER	09251A104	112	13,203	SH		DFND	1	7,723	0	5,480
BLACKROCK MUNIHOLDINGS FUND	OTHER	09253N104	77	4,780	SH		DFND	1	4,780	0	0
BLACKROCK MUNI INTERM DURATION	OTHER	09253X102	662	44,508	SH		DFND	1	44,508	0	0
BLACKROCK MUNIYIELD QUALITY FD	OTHER	09254F100	134	8,123	SH		DFND	1	8,123	0	0
BLACKROCK MUNIYIELD QUALITY FD	OTHER	09254G108	7	500	SH		DFND	1	500	0	0
BLACKROCK MUNIASSETS FD INC	OTHER	09254J102	46	3,101	SH		OTR	1	3,101	0	0
BLACKROCK MUNIHOLDNGS CA QUALI	OTHER	09254L107	188	12,545	SH		DFND	1	12,545	0	0
BLACKROCK MUNIYIELD CA	OTHER	09254M105	3	200	SH		DFND	1	200	0	0
BLACKROCK CORP HIGH YIELD VI C	OTHER	09255P107	119	10,375	SH		DFND	1	10,375	0	0
BLACKROCK CORP HIGH YIELD VI C	OTHER	09255P107	186	16,246	SH		OTR	1	2,400	1,000	12,846
BLACKROCK ENHANCED CAP INCOME	OTHER	09256A109	33	1,915	SH		DFND	1	1,915	0	0
BLACKROCK MUNI 2030 TGT TM TR	OTHER	09257P105	550	21,413	SH		DFND	1	21,413	0	0
BLACKSTONE MORTGAGE TRU CL A	COMMON STOCK	09257W100	398	14,472	SH		DFND	1	9,458	0	5,014
BLACKSTONE MORTGAGE TRU CL A	COMMON STOCK	09257W100	88	3,197	SH		OTR	1	3,120	36	41
BLACKSTONE GROUP INC CLASS A	COMMON STOCK	09260D107	3,255	50,231	SH		DFND	1	49,431	0	800
BLACKSTONE GROUP INC CLASS A	COMMON STOCK	09260D107	3,096	47,771	SH		OTR	1	30,140	5,823	11,808
BLACKROCK SCIENCE TECHNOLOGY T	OTHER	09260K101	92	2,520	SH		DFND	1	2,520	0	0
BLUEBIRD BIO INC	COMMON STOCK	09609G100	53	1,231	SH		DFND	1	706	0	525
BLUEBIRD BIO INC	COMMON STOCK	09609G100	2	47	SH		OTR	1	0	0	47
BLUEPRINT MEDICINES CORP	COMMON STOCK	09627Y109	28	250	SH		DFND	1	230	0	20
BLUEPRINT MEDICINES CORP	COMMON STOCK	09627Y109	1	10	SH		OTR	1	10	0	0
BOINGO WIRELESS INC	COMMON STOCK	09739C102	1	115	SH		DFND	1	115	0	0
BOISE CASCADE CO	COMMON STOCK	09739D100	380	7,955	SH		DFND	1	7,583	0	372
BOISE CASCADE CO	COMMON STOCK	09739D100	5	106	SH		OTR	1	106	0	0
BOOKING HOLDINGS INC	COMMON STOCK	09857L108	50,256	22,564	SH		DFND	1	22,145	0	419
BOOKING HOLDINGS INC	COMMON STOCK	09857L108	9,521	4,275	SH		OTR	1	3,644	203	428
BOOKING HOLDINGS INC	COMMON STOCK	09857L108	376	169	SH		DFND	5	169	0	0
BOX INC CLASS A	COMMON STOCK	10316T104	17	963	SH		DFND	1	918	0	45
BOX INC CLASS A	COMMON STOCK	10316T104	9	500	SH		OTR	1	500	0	0
BRAEMAR HOTELS RESORTS INC	COMMON STOCK	10482B101	7	1,556	SH		DFND	1	1,556	0	0
BRASIL SA A D R	COMMON STOCK	10552T107	90	21,317	SH		DFND	1	16,212	0	5,105
BRASIL SA A D R	COMMON STOCK	10552T107	1	176	SH		OTR	1	0	0	176
BRICKELL BIOTECH INC	COMMON STOCK	10802T105	0	14	SH		DFND	1	14	0	0
BRIGHAM MINERALS INC CL A	COMMON STOCK	10918L103	3	239	SH		DFND	1	126	0	113
BRIGHTHOUSE FINL INC	COMMON STOCK	10922N103	1,628	44,955	SH		DFND	1	43,544	0	1,411
BRIGHTHOUSE FINL INC	COMMON STOCK	10922N103	25	710	SH		OTR	1	102	112	496
BRIGHTSPHERE INVESTMENT GROU	COMMON STOCK	10948W103	31	1,614	SH		DFND	1	321	0	1,293
BRISTOW GROUP INC COM	COMMON STOCK	11040G103	4	150	SH		DFND	1	0	0	150
BRIXMOR PROPERTY GROUP INC	COMMON STOCK	11120U105	65	3,932	SH		DFND	1	2,211	0	1,721
BRIXMOR PROPERTY GROUP INC	COMMON STOCK	11120U105	10	642	SH		OTR	1	0	242	400
BROADRIDGE FINANCIAL SOLUTIONS	COMMON STOCK	11133T103	4,327	28,246	SH		DFND	1	18,645	0	9,601
BROADRIDGE FINANCIAL SOLUTIONS	COMMON STOCK	11133T103	1,451	9,474	SH		OTR	1	7,863	1,023	588
BROADRIDGE FINANCIAL SOLUTIONS	COMMON STOCK	11133T103	91	592	SH		DFND	5	592	0	0
BROADCOM INC	COMMON STOCK	11135F101	30,954	70,697	SH		DFND	1	62,704	0	7,993
BROADCOM INC	COMMON STOCK	11135F101	9,163	20,928	SH		OTR	1	19,871	688	369
BROADCOM INC	COMMON STOCK	11135F101	157	358	SH		DFND	5	341	0	17
BROADWIND INC	COMMON STOCK	11161T207	785	99,010	SH		DFND	1	99,010	0	0
BROOKFIELD INFRASTRUCTURE CORP	COMMON STOCK	11275Q107	6	79	SH		DFND	1	79	0	0
BROOKFIELD INFRASTRUCTURE CORP	COMMON STOCK	11275Q107	14	194	SH		OTR	1	33	0	161
BROOKFIELD PROPERTY REIT I A	COMMON STOCK	11282X103	22	1,500	SH		OTR	1	1,500	0	0
BROOKFIELD RENEWABLE CORP CL A	COMMON STOCK	11284V105	143	2,453	SH		DFND	1	1,598	0	855
BROOKFIELD RENEWABLE CORP CL A	COMMON STOCK	11284V105	141	2,423	SH		OTR	1	863	0	1,560
BROOKLINE BANCORP INC	COMMON STOCK	11373M107	968	80,429	SH		DFND	1	80,388	0	41
BROOKLINE BANCORP INC	COMMON STOCK	11373M107	16	1,365	SH		OTR	1	1,365	0	0
B2GOLD CORP	COMMON STOCK	11777Q209	15	2,674	SH		DFND	1	0	0	2,674
BUILDERS FIRSTSOURCE INC	COMMON STOCK	12008R107	642	15,745	SH		DFND	1	14,675	0	1,070
BUILDERS FIRSTSOURCE INC	COMMON STOCK	12008R107	9	232	SH		OTR	1	202	0	30
C3 AI INC CL A	COMMON STOCK	12468P104	57	410	SH		DFND	1	410	0	0
CAI INTERNATIONAL INC	COMMON STOCK	12477X106	6	188	SH		DFND	1	0	0	188
CAI INTERNATIONAL INC	COMMON STOCK	12477X106	45	1,444	SH		OTR	1	0	1,444	0
CBOE HOLDINGS INC	COMMON STOCK	12503M108	83	894	SH		DFND	1	523	0	371
CBOE HOLDINGS INC	COMMON STOCK	12503M108	51	549	SH		OTR	1	29	0	520
CBRE GROUP INC	COMMON STOCK	12504L109	3,270	52,128	SH		DFND	1	49,678	0	2,450
CBRE GROUP INC	COMMON STOCK	12504L109	832	13,278	SH		OTR	1	13,120	16	142
CBRE GROUP INC	COMMON STOCK	12504L109	55	878	SH		DFND	5	878	0	0
CDW CORP DE	COMMON STOCK	12514G108	4,611	34,983	SH		DFND	1	29,281	0	5,702
CDW CORP DE	COMMON STOCK	12514G108	932	7,068	SH		OTR	1	6,140	72	856
CDW CORP	COMMON STOCK	12514G108	271	2,054	SH		DFND	5	2,054	0	0
CGI INC	COMMON STOCK	12532H104	217	2,734	SH		DFND	1	1,471	0	1,263
CGI INC	COMMON STOCK	12532H104	60	754	SH		OTR	1	754	0	0
C H ROBINSON WORLDWIDE INC	COMMON STOCK	12541W209	67,707	721,286	SH		DFND	1	715,353	0	5,933
C H ROBINSON WORLDWIDE INC	COMMON STOCK	12541W209	11,672	124,338	SH		OTR	1	115,802	3,473	5,063
CMC MATLS INC	COMMON STOCK	12571T100	2,811	18,577	SH		DFND	1	18,573	0	4
CMC MATLS INC	COMMON STOCK	12571T100	409	2,706	SH		OTR	1	2,690	0	16
CMC MATERIALS INC	COMMON STOCK	12571T100	40	263	SH		DFND	5	263	0	0
CME GROUP INC	COMMON STOCK	12572Q105	66,444	364,979	SH		DFND	1	362,552	0	2,427
CME GROUP INC	COMMON STOCK	12572Q105	7,942	43,626	SH		OTR	1	37,627	1,306	4,693
CME GROUP INC	COMMON STOCK	12572Q105	56	306	SH		DFND	5	306	0	0
CRA INTERNATIONAL INC	COMMON STOCK	12618T105	14	288	SH		OTR	1	25	263	0
CRH SPON A D R	COMMON STOCK	12626K203	3,349	78,652	SH		DFND	1	75,647	0	3,005
CRH SPON A D R	COMMON STOCK	12626K203	699	16,401	SH		OTR	1	16,022	353	26
CRH PLC	COMMON STOCK	12626K203	22	527	SH		DFND	5	527	0	0
CNX RESOURCES CORP	COMMON STOCK	12653C108	147	13,615	SH		DFND	1	9,062	0	4,553
CNX RESOURCES CORP	COMMON STOCK	12653C108	6	537	SH		OTR	1	0	18	519
CVR ENERGY INC	COMMON STOCK	12662P108	46	3,080	SH		DFND	1	3,000	0	80
CABALETTA BIO INC	COMMON STOCK	12674W109	11	900	SH		DFND	1	900	0	0
CABLE ONE INC	COMMON STOCK	12685J105	929	417	SH		DFND	1	243	0	174
CABLE ONE INC	COMMON STOCK	12685J105	179	80	SH		OTR	1	0	76	4
CABLE ONE INC	COMMON STOCK	12685J105	45	20	SH		DFND	5	20	0	0
CADENCE BANCORP	COMMON STOCK	12739A100	45	2,711	SH		DFND	1	1,982	0	729
CADENCE BANCORP	COMMON STOCK	12739A100	7	446	SH		OTR	1	0	446	0
CAESARS ENTERTAINMENT INC	COMMON STOCK	12769G100	57	773	SH		DFND	1	676	0	97
CAESARS ENTERTAINMENT INC	COMMON STOCK	12769G100	15	208	SH		OTR	1	0	0	208
CAESARS ENTERTAINMENT INC	COMMON STOCK	12769G100	1	14	SH		DFND	5	14	0	0
CALAMOS GLOBAL DYNAMIC INCOME	OTHER	12811L107	3	319	SH		DFND	1	319	0	0
CALAMOS CONV & HIGH INCOME FD	OTHER	12811P108	85	5,959	SH		OTR	1	5,959	0	0
CALIX NETWORKS INC	COMMON STOCK	13100M509	20	680	SH		DFND	1	420	0	260
CALLON PETROLEUM COMPANY	COMMON STOCK	13123X508	14	1,073	SH		DFND	1	1,073	0	0
CAMECO CORP	COMMON STOCK	13321L108	108	8,039	SH		DFND	1	5,306	0	2,733
CAMECO CORP	COMMON STOCK	13321L108	159	11,899	SH		OTR	1	0	0	11,899
CAMPING WORLD HOLDINGS INC A	COMMON STOCK	13462K109	3	129	SH		DFND	1	0	0	129
CAMPING WORLD HOLDINGS INC A	COMMON STOCK	13462K109	29	1,118	SH		OTR	1	0	1,118	0
CANADIAN PACIFIC RAILWAY LTD	COMMON STOCK	13645T100	1,222	3,525	SH		DFND	1	2,965	0	560
CANADIAN PACIFIC RAILWAY LTD	COMMON STOCK	13645T100	581	1,677	SH		OTR	1	1,443	0	234
CANADIAN PACIFIC RAILWAY LTD	COMMON STOCK	13645T100	397	1,142	SH		DFND	5	1,142	0	0
CANNAE HOLDINGS INC	COMMON STOCK	13765N107	16	362	SH		DFND	1	362	0	0
CAPITAL ONE FINANCIAL CORP	COMMON STOCK	14040H105	6,391	64,652	SH		DFND	1	59,769	0	4,883
CAPITAL ONE FINANCIAL CORP	COMMON STOCK	14040H105	1,767	17,874	SH		OTR	1	10,478	221	7,175
CAPITAL ONE FINANCIAL CORP	COMMON STOCK	14040H105	3	28	SH		DFND	5	28	0	0
CAPITOL FEDERAL FINANCIAL INC	COMMON STOCK	14057J101	52	4,185	SH		DFND	1	4,185	0	0
CAPITOL FEDERAL FINANCIAL INC	COMMON STOCK	14057J101	33	2,637	SH		OTR	1	2,637	0	0
CAPSTEAD MORTGAGE CORP	COMMON STOCK	14067E506	2	286	SH		DFND	1	286	0	0
CAPRICOR THERAPEUTICS INC	COMMON STOCK	14070B309	26	7,462	SH		OTR	1	7,462	0	0
CARDINAL HEALTH INC	COMMON STOCK	14149Y108	4,683	87,435	SH		DFND	1	79,212	0	8,223
CARDINAL HEALTH INC	COMMON STOCK	14149Y108	1,324	24,702	SH		OTR	1	14,859	7,886	1,957
CARDINAL HEALTH INC	COMMON STOCK	14149Y108	15	276	SH		DFND	5	94	0	182
CARDLYTICS INC	COMMON STOCK	14161W105	13	89	SH		DFND	1	0	0	89
CAREDX INC	COMMON STOCK	14167L103	32	436	SH		DFND	1	436	0	0
CARETRUST REIT INC	COMMON STOCK	14174T107	27	1,200	SH		DFND	1	1,200	0	0
CARETRUST REIT INC	COMMON STOCK	14174T107	1	42	SH		OTR	1	42	0	0
CARLYLE GROUP INC THE	COMMON STOCK	14316J108	58	1,850	SH		OTR	1	1,850	0	0
CARNIVAL PLC A D R	COMMON STOCK	14365C103	547	29,188	SH		DFND	1	20,778	0	8,410
CARNIVAL PLC A D R	COMMON STOCK	14365C103	40	2,117	SH		OTR	1	410	1,707	0
CARRIER GLOBAL CORPORATION COM	COMMON STOCK	14448C104	8,982	238,116	SH		DFND	1	231,210	0	6,906
CARRIER GLOBAL CORPORATION COM	COMMON STOCK	14448C104	5,748	152,381	SH		OTR	1	138,967	6,192	7,222
CARRIER GLOBAL CORP	COMMON STOCK	14448C104	13	334	SH		DFND	5	334	0	0
CARROLS RESTAURANT GROUP INC	COMMON STOCK	14574X104	9	1,388	SH		DFND	1	1,388	0	0
CASA SYSTEMS INC	COMMON STOCK	14713L102	1	182	SH		DFND	1	0	0	182
CASS INFORMATION SYSTEMS INC	COMMON STOCK	14808P109	215	5,525	SH		DFND	1	5,525	0	0
CASS INFORMATION SYSTEMS INC	COMMON STOCK	14808P109	57	1,468	SH		OTR	1	1,468	0	0
CASS INFORMATION SYSTEMS INC	COMMON STOCK	14808P109	2	55	SH		DFND	5	55	0	0
CATCHMARK TIMBER TRUST INC A	COMMON STOCK	14912Y202	11	1,174	SH		DFND	1	1,174	0	0
CELESTICA INC	COMMON STOCK	15101Q108	84	10,444	SH		DFND	1	236	0	10,208
CELCUITY INC	COMMON STOCK	15102K100	5	548	SH		OTR	1	548	0	0
CELLDEX THERAPEUTICS INC	COMMON STOCK	15117B202	6	356	SH		DFND	1	356	0	0
CELLECTAR BIOSCIENCES INC	COMMON STOCK	15117F500	0	1	SH		DFND	1	1	0	0
CELLECTAR BIOSCIENCES INC	COMMON STOCK	15117F500	6	2,857	SH		OTR	1	2,857	0	0
CELLECTIS A D R	COMMON STOCK	15117K103	30	1,110	SH		DFND	1	930	0	180
CELLECTIS A D R	COMMON STOCK	15117K103	23	854	SH		OTR	1	0	854	0
CENTENE CORP	COMMON STOCK	15135B101	75,092	1,250,892	SH		DFND	1	1,236,671	0	14,221
CENTENE CORP	COMMON STOCK	15135B101	10,639	177,242	SH		OTR	1	167,646	7,394	2,202
CENOVUS ENERGY INC	COMMON STOCK	15135U109	536	88,714	SH		DFND	1	75,586	0	13,128
CENOVUS ENERGY INC	COMMON STOCK	15135U109	102	16,845	SH		OTR	1	16,588	0	257
CENTERPOINT ENERGY INC	COMMON STOCK	15189T107	1,129	52,160	SH		DFND	1	46,070	0	6,090
CENTERPOINT ENERGY INC	COMMON STOCK	15189T107	426	19,671	SH		OTR	1	18,590	38	1,043
CENTERPOINT ENERGY INC	COMMON STOCK	15189T107	19	882	SH		DFND	5	882	0	0
CENTERSPACE	COMMON STOCK	15202L107	174	2,467	SH		DFND	1	2,467	0	0
CENTERSPACE	COMMON STOCK	15202L107	126	1,787	SH		OTR	1	1,787	0	0
CENTERSPACE	COMMON STOCK	15202L107	0	5	SH		DFND	5	5	0	0
CENTRAIS ELETRICAS BRASILEIRAS	COMMON STOCK	15234Q207	9	1,254	SH		DFND	1	1,254	0	0
CERIDIAN HCM HOLDING INC	COMMON STOCK	15677J108	49	457	SH		DFND	1	333	0	124
CERIDIAN HCM HOLDING INC	COMMON STOCK	15677J108	8	77	SH		OTR	1	0	45	32
CERTARA INC	COMMON STOCK	15687V109	22	645	SH		DFND	1	645	0	0
CHAMPIONX CORPORATION COM	COMMON STOCK	15872M104	1,336	87,288	SH		DFND	1	27,043	0	60,245
CHAMPIONX CORPORATION COM	COMMON STOCK	15872M104	265	17,339	SH		OTR	1	17,339	0	0
CHANGE HEALTHCARE INC	COMMON STOCK	15912K100	22	1,162	SH		DFND	1	307	0	855
CHANGE HEALTHCARE INC	COMMON STOCK	15912K100	167	8,961	SH		OTR	1	4,352	4,609	0
CHANGE HEALTHCARE INC	COMMON STOCK	15912K100	45	2,437	SH		DFND	5	2,437	0	0
CHART INDUSTRIES INC	COMMON STOCK	16115Q308	2,001	16,986	SH		DFND	1	13,683	0	3,303
CHART INDUSTRIES INC	COMMON STOCK	16115Q308	283	2,403	SH		OTR	1	1,346	1,046	11
CHART INDUSTRIES INC	COMMON STOCK	16115Q308	1	6	SH		DFND	5	6	0	0
CHARTER COMMUNICATIONS INC NEW	COMMON STOCK	16119P108	4,127	6,238	SH		DFND	1	4,830	0	1,408
CHARTER COMMUNICATIONS INC NEW	COMMON STOCK	16119P108	1,072	1,622	SH		OTR	1	490	640	492
CHARTER COMMUNICATIONS INC	COMMON STOCK	16119P108	3	4	SH		DFND	5	4	0	0
CHASE CORP	COMMON STOCK	16150R104	4	44	SH		OTR	1	44	0	0
CHATHAM LODGING TRUST	COMMON STOCK	16208T102	4	391	SH		DFND	1	391	0	0
CHEMED CORP	COMMON STOCK	16359R103	5,056	9,493	SH		DFND	1	9,410	0	83
CHEMED CORP	COMMON STOCK	16359R103	414	778	SH		OTR	1	761	10	7
CHEMED CORPORATION	COMMON STOCK	16359R103	398	747	SH		DFND	5	747	0	0
CHEMOCENTRYX INC	COMMON STOCK	16383L106	400	6,463	SH		DFND	1	6,437	0	26
CHEMOCENTRYX INC	COMMON STOCK	16383L106	5	79	SH		OTR	1	79	0	0
CHENIERE ENERGY PARTNERS LP	COMMON STOCK	16411Q101	159	4,500	SH		DFND	1	4,500	0	0
CHENIERE ENERGY PARTNERS LP	COMMON STOCK	16411Q101	35	1,000	SH		OTR	1	1,000	0	0
CHENIERE ENERGY INC	COMMON STOCK	16411R208	489	8,139	SH		DFND	1	4,721	0	3,418
CHENIERE ENERGY INC	COMMON STOCK	16411R208	115	1,915	SH		OTR	1	638	873	404
CHENIERE ENERGY INC	COMMON STOCK	16411R208	8	132	SH		DFND	5	132	0	0
CHEWY INC CLASS A	COMMON STOCK	16679L109	710	7,902	SH		DFND	1	7,902	0	0
CHEWY INC CLASS A	COMMON STOCK	16679L109	40	450	SH		OTR	1	100	0	350
CHEWY INC	COMMON STOCK	16679L109	26	291	SH		DFND	5	291	0	0
CHIMERA INVESTMENT CORP	COMMON STOCK	16934Q208	41	4,000	SH		OTR	1	4,000	0	0
CHIMERA INVESTMENT CORP	COMMON STOCK	16934Q208	37	3,644	SH		DFND	5	3,644	0	0
CHINA EASTERN AIRLINES A D R	COMMON STOCK	16937R104	24	1,115	SH		DFND	1	1,000	0	115
CHINA LIFE INSURANCE CO A D R	COMMON STOCK	16939P106	175	15,776	SH		DFND	1	8,850	0	6,926
CHINA LIFE INSURANCE CO A D R	COMMON STOCK	16939P106	0	1	SH		OTR	1	1	0	0
CHINA MOBILE LIMITED A D R	COMMON STOCK	16941M109	1,351	47,311	SH		DFND	1	46,959	0	352
CHINA MOBILE LIMITED A D R	COMMON STOCK	16941M109	303	10,625	SH		OTR	1	10,021	0	604
CHINA PETROLEUM CHEM A D R	COMMON STOCK	16941R108	438	9,819	SH		DFND	1	9,695	0	124
CHINA PETROLEUM CHEM A D R	COMMON STOCK	16941R108	104	2,329	SH		OTR	1	2,329	0	0
CHINA UNICOM HONG KONG LTD A D	COMMON STOCK	16945R104	83	14,666	SH		DFND	1	11,889	0	2,777
CHINA UNICOM LTD	COMMON STOCK	16945R104	1	240	SH		DFND	5	240	0	0
CHIPMOS TECHNOLOGIES INC ADR A	COMMON STOCK	16965P202	12	507	SH		DFND	1	0	0	507
CHUNGHWA TELECOM CO LTD A D R	COMMON STOCK	17133Q502	1,382	35,796	SH		DFND	1	23,169	0	12,627
CHUNGHWA TELECOM CO LTD A D R	COMMON STOCK	17133Q502	90	2,326	SH		OTR	1	718	0	1,608
CINEMARK HLDGS INC	COMMON STOCK	17243V102	95	5,455	SH		DFND	1	3,550	0	1,905
CIRCOR INTERNATIONAL INC	COMMON STOCK	17273K109	84	2,183	SH		DFND	1	84	0	2,099
CISCO SYSTEMS INC	COMMON STOCK	17275R102	149,710	3,345,474	SH		DFND	1	3,218,869	0	126,605
CISCO SYSTEMS INC	COMMON STOCK	17275R102	40,921	914,438	SH		OTR	1	851,702	27,712	35,024
CISCO SYSTEMS INC	COMMON STOCK	17275R102	673	15,025	SH		DFND	5	14,759	0	266
CITI TRENDS INC	COMMON STOCK	17306X102	9	175	SH		OTR	1	175	0	0
CITIUS PHARMACEUTICALS INC	COMMON STOCK	17322U207	10	10,000	SH		DFND	1	10,000	0	0
CLEANSPARK INC	COMMON STOCK	18452B209	29	1,000	SH		DFND	1	1,000	0	0
CLEARBRIDGE ENERGY MIDSTREAM	OTHER	18469P209	110	8,161	SH		DFND	1	968	0	7,193
CLEARWATER PAPER CORP	COMMON STOCK	18538R103	3	81	SH		DFND	1	81	0	0
CLEARWATER PAPER CORP	COMMON STOCK	18538R103	20	537	SH		OTR	1	537	0	0
CLEARWAY ENERGY INC CLASS A	COMMON STOCK	18539C105	12	400	SH		DFND	1	400	0	0
CLEARWAY ENERGY INC CLASS A	COMMON STOCK	18539C105	7	250	SH		OTR	1	250	0	0
CLEARWAY ENERGY INC C	COMMON STOCK	18539C204	45	1,397	SH		DFND	1	1,397	0	0
CLEARWAY ENERGY INC C	COMMON STOCK	18539C204	38	1,180	SH		OTR	1	1,000	55	125
CLEARWAY ENERGY INC	COMMON STOCK	18539C204	1	42	SH		DFND	5	42	0	0
CLOUDERA INC	COMMON STOCK	18914U100	6	400	SH		DFND	1	400	0	0
CLOUDFLARE INC CLASS A	COMMON STOCK	18915M107	2,571	33,828	SH		DFND	1	33,828	0	0
CLOUDFLARE INC CLASS A	COMMON STOCK	18915M107	927	12,204	SH		OTR	1	12,204	0	0
COGENT COMMUNICATIONS HOLDIN	COMMON STOCK	19239V302	1,555	25,967	SH		DFND	1	24,900	0	1,067
COGENT COMMUNICATIONS HOLDIN	COMMON STOCK	19239V302	72	1,206	SH		OTR	1	1,206	0	0
COGENT COMMUNICATIONS GROUP	COMMON STOCK	19239V302	8	129	SH		DFND	5	129	0	0
COHEN STEERS INC	COMMON STOCK	19247A100	1,467	19,740	SH		DFND	1	19,051	0	689
COHEN STEERS INC	COMMON STOCK	19247A100	49	665	SH		OTR	1	619	46	0
COHEN & STEERS INC	COMMON STOCK	19247A100	15	196	SH		DFND	5	196	0	0
COHEN & STEERS QUALITY INCOME	OTHER	19247L106	91	7,365	SH		DFND	1	4,088	0	3,277
COHEN STEERS REIT AND PREFERRE	OTHER	19247X100	26	1,125	SH		OTR	1	1,125	0	0
COHEN & STEERS INFRASTRUCTURE	OTHER	19248A109	187	7,242	SH		DFND	1	7,242	0	0
COHEN STEERS LIMITED DURAT	OTHER	19248C105	10	391	SH		DFND	1	391	0	0
COHEN STEERS LIMITED DURAT	OTHER	19248C105	118	4,429	SH		OTR	1	4,429	0	0
COHERUS BIOSCIENCES INC	COMMON STOCK	19249H103	49	2,840	SH		OTR	1	0	2,840	0
COHBAR INC	COMMON STOCK	19249J109	11	7,915	SH		OTR	1	7,915	0	0
COLLECTORS UNIVERSE INC	COMMON STOCK	19421R200	1	15	SH		DFND	1	0	0	15
COLONY CREDIT REAL ESTATE IN	COMMON STOCK	19625T101	11	1,441	SH		DFND	1	808	0	633
COLONY CAPITAL INC	COMMON STOCK	19626G108	41	8,583	SH		DFND	1	3,427	0	5,156
COLONY CAPITAL INC	COMMON STOCK	19626G108	1	258	SH		OTR	1	0	0	258
COMCAST CORP CLASS A	COMMON STOCK	20030N101	121,275	2,314,408	SH		DFND	1	2,232,198	0	82,210
COMCAST CORP CLASS A	COMMON STOCK	20030N101	28,333	540,689	SH		OTR	1	517,941	11,077	11,671
COMCAST CORPORATION	COMMON STOCK	20030N101	769	14,652	SH		DFND	5	14,510	0	142
COMMSCOPE HLDG CO INC	COMMON STOCK	20337X109	32	2,407	SH		DFND	1	1,733	0	674
COMMSCOPE HLDG CO INC	COMMON STOCK	20337X109	1	107	SH		OTR	1	0	107	0
COMMUNITY HEALTHCARE TRUST I	COMMON STOCK	20369C106	11	232	SH		DFND	1	128	0	104
CIA BRASILEIRA DE DIS ADR A D	COMMON STOCK	20440T300	123	8,556	SH		DFND	1	6,663	0	1,893
CIA BRASILEIRA DE DIS ADR A D	COMMON STOCK	20440T300	3	200	SH		OTR	1	200	0	0
COMPANHIA SIDERURGICA NACIONAL	COMMON STOCK	20440W105	160	26,978	SH		DFND	1	16,927	0	10,051
COMPANHIA SIDERURGICA NACIONAL	COMMON STOCK	20440W105	20	3,432	SH		OTR	1	3,301	0	131
CIA SIDERURGICA NACIONAL	COMMON STOCK	20440W105	1	222	SH		DFND	5	222	0	0
CIA SANEAMENTO BASICO DE A D R	COMMON STOCK	20441A102	209	24,414	SH		DFND	1	22,704	0	1,710
CIA SANEAMENTO BASICO DE A D R	COMMON STOCK	20441A102	29	3,358	SH		OTR	1	3,358	0	0
COMPANHIA PARANAENSE ENER SP A	COMMON STOCK	20441B407	540	37,735	SH		DFND	1	33,303	0	4,432
COMPANHIA PARANAENSE ENER SP A	COMMON STOCK	20441B407	35	2,399	SH		OTR	1	2,399	0	0
COMPANHIA PARANAENSE DE ENERGI	COMMON STOCK	20441B407	1	41	SH		DFND	5	41	0	0
COMPASS MINERALS INTERNATIONAL	COMMON STOCK	20451N101	389	6,301	SH		DFND	1	5,803	0	498
COMPASS MINERALS INTERNATIONAL	COMMON STOCK	20451N101	830	13,447	SH		OTR	1	13,447	0	0
COMPASS DIVERSIFIED HOLDINGS	COMMON STOCK	20451Q104	375	19,255	SH		DFND	1	19,255	0	0
COMPASS DIVERSIFIED HOLDINGS	COMMON STOCK	20451Q104	19	1,000	SH		OTR	1	1,000	0	0
CONCENTRIX CORP	COMMON STOCK	20602D101	383	3,883	SH		DFND	1	3,535	0	348
CONCENTRIX CORP	COMMON STOCK	20602D101	280	2,835	SH		OTR	1	2,568	252	15
CONCHO RES INC	COMMON STOCK	20605P101	243	4,166	SH		DFND	1	2,404	0	1,762
CONCHO RES INC	COMMON STOCK	20605P101	180	3,085	SH		OTR	1	24	0	3,061
CONNECTONE BANCORP INC	COMMON STOCK	20786W107	3	144	SH		DFND	1	144	0	0
CONOCOPHILLIPS	COMMON STOCK	20825C104	18,520	463,124	SH		DFND	1	451,687	0	11,437
CONOCOPHILLIPS	COMMON STOCK	20825C104	8,691	217,331	SH		OTR	1	207,267	1,701	8,363
CONOCOPHILLIPS	COMMON STOCK	20825C104	98	2,453	SH		DFND	5	2,277	0	176
CONSTELLATION BRANDS INC A	COMMON STOCK	21036P108	4,688	21,404	SH		DFND	1	19,677	0	1,727
CONSTELLATION BRANDS INC A	COMMON STOCK	21036P108	2,667	12,174	SH		OTR	1	12,075	7	92
CONSTELLATION BRANDS INC	COMMON STOCK	21036P108	7	30	SH		DFND	5	0	0	30
CONSTRUCTION PARTNERS INC A	COMMON STOCK	21044C107	8	261	SH		DFND	1	0	0	261
CONSTRUCTION PARTNERS INC A	COMMON STOCK	21044C107	218	7,490	SH		OTR	1	380	0	7,110
COOPER STANDARD HOLDING	COMMON STOCK	21676P103	3	81	SH		DFND	1	0	0	81
COOTEK CAYMAN INC A D R	COMMON STOCK	21718L102	0	20	SH		DFND	1	0	0	20
CORBUS PHARMACEUTICALS HLDGS I	COMMON STOCK	21833P103	0	200	SH		DFND	1	0	0	200
CORESITE REALTY CORP	COMMON STOCK	21870Q105	300	2,398	SH		DFND	1	1,813	0	585
CORESITE REALTY CORP	COMMON STOCK	21870Q105	551	4,399	SH		OTR	1	2,189	0	2,210
CORESITE REALTY CORP	COMMON STOCK	21870Q105	36	286	SH		DFND	5	286	0	0
CORELOGIC INC	COMMON STOCK	21871D103	57	738	SH		DFND	1	723	0	15
CORELOGIC INC	COMMON STOCK	21871D103	155	2,000	SH		OTR	1	2,000	0	0
COREPOINT LODGING INC	COMMON STOCK	21872L104	6	845	SH		DFND	1	845	0	0
CORNERSTONE BUILDING BRANDS IN	COMMON STOCK	21925D109	11	1,215	SH		DFND	1	898	0	317
CORNERSTONE BUILDING BRANDS IN	COMMON STOCK	21925D109	1	125	SH		OTR	1	0	0	125
CORNERSTONE ONDEMAND INC	COMMON STOCK	21925Y103	26	583	SH		DFND	1	0	0	583
CORPORATE OFFICE PPTYS TR	COMMON STOCK	22002T108	121	4,631	SH		DFND	1	1,612	0	3,019
CORPORATE OFFICE PPTYS TR	COMMON STOCK	22002T108	6	243	SH		OTR	1	167	0	76
CORTEVA INC	COMMON STOCK	22052L104	4,857	125,435	SH		DFND	1	118,206	0	7,229
CORTEVA INC	COMMON STOCK	22052L104	5,692	147,014	SH		OTR	1	139,365	2,830	4,819
CORTEVA INC	COMMON STOCK	22052L104	11	283	SH		DFND	5	283	0	0
COSTCO WHSL CORP	COMMON STOCK	22160K105	169,056	448,686	SH		DFND	1	435,841	0	12,845
COSTCO WHSL CORP	COMMON STOCK	22160K105	46,012	122,121	SH		OTR	1	86,284	1,566	34,271
COSTCO WHOLESALE CORP	COMMON STOCK	22160K105	1,002	2,643	SH		DFND	5	2,620	0	23
COSTAR GROUP INC	COMMON STOCK	22160N109	6,041	6,536	SH		DFND	1	6,093	0	443
COSTAR GROUP INC	COMMON STOCK	22160N109	1,620	1,753	SH		OTR	1	518	1,195	40
COSTAR GROUP INC	COMMON STOCK	22160N109	389	421	SH		DFND	5	419	0	2
COUPA SOFTWARE INC	COMMON STOCK	22266L106	2,525	7,452	SH		DFND	1	6,389	0	1,063
COUPA SOFTWARE INC	COMMON STOCK	22266L106	568	1,676	SH		OTR	1	143	1,493	40
COUPA SOFTWARE INCORPORATED	COMMON STOCK	22266L106	18	53	SH		DFND	5	53	0	0
COVETRUS INC NPV	COMMON STOCK	22304C100	49	1,699	SH		DFND	1	875	0	824
COVETRUS INC NPV	COMMON STOCK	22304C100	24	836	SH		OTR	1	654	0	182
CRACKER BARREL OLD CTRY STORE	COMMON STOCK	22410J106	243	1,840	SH		DFND	1	1,077	0	763
CRACKER BARREL OLD CTRY STORE	COMMON STOCK	22410J106	34	259	SH		OTR	1	245	0	14
CREATIVE REALITIES INC	COMMON STOCK	22530J200	0	4	SH		OTR	1	4	0	0
CREDIT SUISSE FI LARGE CAP GR	OTHER	22542D423	109	200	SH		DFND	1	200	0	0
CRINETICS PHARMACEUTICALS IN	COMMON STOCK	22663K107	1	63	SH		DFND	1	0	0	63
CRONOS GROUP INC	COMMON STOCK	22717L101	13	1,900	SH		OTR	1	1,900	0	0
CROWDSTRIKE HOLDINGS INC A	COMMON STOCK	22788C105	3,028	14,296	SH		DFND	1	14,098	0	198
CROWDSTRIKE HOLDINGS INC A	COMMON STOCK	22788C105	1,917	9,050	SH		OTR	1	8,187	619	244
CROWDSTRIKE HOLDINGS INC	COMMON STOCK	22788C105	4	18	SH		DFND	5	18	0	0
CROWN CASTLE INTL CORP	COMMON STOCK	22822V101	21,934	137,789	SH		DFND	1	127,397	0	10,392
CROWN CASTLE INTL CORP	COMMON STOCK	22822V101	5,030	31,597	SH		OTR	1	28,405	735	2,457
CROWN CASTLE INTL CORP	COMMON STOCK	22822V101	312	1,958	SH		DFND	5	1,958	0	0
CTO REALTY GROWTH INC	COMMON STOCK	22948P103	358	8,489	SH		DFND	1	8,489	0	0
CUSTOMERS BANCORP INC	COMMON STOCK	23204G100	18	993	SH		DFND	1	643	0	350
CYMABAY THERAPEUTICS INC	COMMON STOCK	23257D103	8	1,333	SH		DFND	1	856	0	477
CYTOKINETICS INC	COMMON STOCK	23282W605	14	689	SH		DFND	1	275	0	414
CYRUSONE INC	COMMON STOCK	23283R100	541	7,401	SH		DFND	1	7,134	0	267
CYRUSONE INC	COMMON STOCK	23283R100	340	4,647	SH		OTR	1	1,672	2,421	554
CYRUSONE INC	COMMON STOCK	23283R100	3	42	SH		DFND	5	42	0	0
DMC GLOBAL INC	COMMON STOCK	23291C103	58	1,345	SH		DFND	1	21	0	1,324
DBV TECHNOLOGIES SA A D R	COMMON STOCK	23306J101	4	1,576	SH		DFND	1	0	0	1,576
DBV TECHNOLOGIES SA A D R	COMMON STOCK	23306J101	26	10,000	SH		OTR	1	10,000	0	0
DCP MIDSTREAM LP	COMMON STOCK	23311P100	185	10,000	SH		DFND	1	10,000	0	0
DCP MIDSTREAM PARTNERS	COMMON STOCK	23311P100	2	130	SH		DFND	5	130	0	0
DNP SELECT INCOME FUND INC	OTHER	23325P104	976	95,160	SH		DFND	1	90,706	0	4,454
DNP SELECT INCOME FUND INC	OTHER	23325P104	497	48,419	SH		OTR	1	44,719	0	3,700
DNP SELECT INCOME FUND INC	OTHER	23325P104	30	2,947	SH		DFND	5	0	0	2,947
D R HORTON INC	COMMON STOCK	23331A109	11,204	162,560	SH		DFND	1	159,574	0	2,986
D R HORTON INC	COMMON STOCK	23331A109	2,116	30,710	SH		OTR	1	28,988	1,047	675
D.R. HORTON INC	COMMON STOCK	23331A109	115	1,673	SH		DFND	5	1,673	0	0
D S P GROUP INC	COMMON STOCK	23332B106	3	179	SH		DFND	1	0	0	179
DEUTSCHE STRATEGIC MUNICIPAL I	OTHER	23342Q101	55	4,806	SH		DFND	1	4,806	0	0
DXC TECHNOLOGY CO	COMMON STOCK	23355L106	310	12,021	SH		DFND	1	9,938	0	2,083
DXC TECHNOLOGY CO	COMMON STOCK	23355L106	217	8,467	SH		OTR	1	7,590	230	647
DAQO NEW ENERGY CORP A D R	COMMON STOCK	23703Q203	35	615	SH		DFND	1	90	0	525
DASEKE INC	COMMON STOCK	23753F107	1,801	310,000	SH		DFND	1	310,000	0	0
DATADOG INCCLASS A	COMMON STOCK	23804L103	149	1,517	SH		DFND	1	1,428	0	89
DATADOG INCCLASS A	COMMON STOCK	23804L103	32	330	SH		OTR	1	187	106	37
DATADOG INC	COMMON STOCK	23804L103	5	50	SH		DFND	5	50	0	0
DAVITA INC	COMMON STOCK	23918K108	179	1,526	SH		DFND	1	1,103	0	423
DAVITA INC	COMMON STOCK	23918K108	48	410	SH		OTR	1	380	0	30
DAVITA INC	COMMON STOCK	23918K108	40	338	SH		DFND	5	338	0	0
DECIPHERA PHARMACEUTICALS IN	COMMON STOCK	24344T101	704	12,339	SH		DFND	1	12,330	0	9
DECIPHERA PHARMACEUTICALS IN	COMMON STOCK	24344T101	12	204	SH		OTR	1	204	0	0
DELAWARE INVTS MINN MUN INCOME	OTHER	24610V103	5	352	SH		DFND	1	352	0	0
DELAWARE INVTS MINN MUN INCOME	OTHER	24610V103	36	2,706	SH		OTR	1	2,706	0	0
DELEK LOGISTICS PARTNERSHIP LP	COMMON STOCK	24664T103	32	1,000	SH		OTR	1	1,000	0	0
DELEK HOLDCO INC	COMMON STOCK	24665A103	120	7,421	SH		DFND	1	6,567	0	854
DELEK HOLDCO INC	COMMON STOCK	24665A103	1	92	SH		OTR	1	92	0	0
DELL TECHNOLOGIES C	COMMON STOCK	24703L202	571	7,797	SH		DFND	1	5,870	0	1,927
DELL TECHNOLOGIES C	COMMON STOCK	24703L202	255	3,486	SH		OTR	1	3,054	295	137
DENALI THERAPEUTICS INC	COMMON STOCK	24823R105	33	395	SH		DFND	1	346	0	49
DENNYS CORP	COMMON STOCK	24869P104	590	40,220	SH		DFND	1	40,220	0	0
DENNYS CORP	COMMON STOCK	24869P104	108	7,364	SH		OTR	1	7,364	0	0
DENTSPLY SIRONA INC	COMMON STOCK	24906P109	1,121	21,410	SH		DFND	1	19,236	0	2,174
DENTSPLY SIRONA INC	COMMON STOCK	24906P109	406	7,761	SH		OTR	1	7,757	0	4
DENTSPLY SIRONA INC	COMMON STOCK	24906P109	52	991	SH		DFND	5	991	0	0
DERMTECH INC	COMMON STOCK	24984K105	3	100	SH		OTR	1	100	0	0
DESKTOP METAL INC COM CL A	COMMON STOCK	25058X105	206	11,972	SH		DFND	1	11,972	0	0
DESKTOP METAL INC COM CL A	COMMON STOCK	25058X105	3	148	SH		OTR	1	148	0	0
DEVON ENERGY CORP	COMMON STOCK	25179M103	1,017	64,319	SH		DFND	1	50,108	0	14,211
DEVON ENERGY CORP	COMMON STOCK	25179M103	119	7,523	SH		OTR	1	5,882	0	1,641
DEVON ENERGY CORP	COMMON STOCK	25179M103	33	2,087	SH		DFND	5	2,087	0	0
DIAGEO PLC SPONSORED A D R	COMMON STOCK	25243Q205	11,917	75,040	SH		DFND	1	69,531	0	5,509
DIAGEO PLC SPONSORED A D R	COMMON STOCK	25243Q205	4,359	27,444	SH		OTR	1	25,697	1,126	621
DIAGEO PLC	COMMON STOCK	25243Q205	48	300	SH		DFND	5	287	0	13
DIAMEDICA THERAPEUTICS INC	COMMON STOCK	25253X207	44	4,335	SH		OTR	1	4,335	0	0
DIAMOND HILL INVT GROUP INC	COMMON STOCK	25264R207	14	92	SH		DFND	1	28	0	64
DIAMONDBACK ENERGY INC	COMMON STOCK	25278X109	546	11,285	SH		DFND	1	9,158	0	2,127
DIAMONDBACK ENERGY INC	COMMON STOCK	25278X109	30	607	SH		OTR	1	0	238	369
DIAMONDBACK ENERGY INC	COMMON STOCK	25278X109	2	46	SH		DFND	5	46	0	0
DIGITAL TURBINE INC	COMMON STOCK	25400W102	1,204	21,284	SH		DFND	1	21,068	0	216
DIGITAL TURBINE INC	COMMON STOCK	25400W102	7	121	SH		OTR	1	121	0	0
DIREXION DAILY MSCI REAL EST E	OTHER	25459W755	0	25	SH		DFND	1	25	0	0
DIREXION DAILY SMALL CAP ETF	OTHER	25459W847	0	2	SH		DFND	1	2	0	0
DIREXION DAILY 20 YR TRSY BEAR	OTHER	25460G849	6	115	SH		DFND	1	115	0	0
DISCOVERY INC A	COMMON STOCK	25470F104	1,604	53,317	SH		DFND	1	53,317	0	0
DISCOVERY INC A	COMMON STOCK	25470F104	45	1,489	SH		OTR	1	1,489	0	0
DISCOVERY COMMUNICATIONS	COMMON STOCK	25470F104	32	1,069	SH		DFND	5	1,069	0	0
DISCOVERY INC C	COMMON STOCK	25470F302	246	9,380	SH		DFND	1	9,274	0	106
DISCOVERY INC C	COMMON STOCK	25470F302	597	22,796	SH		OTR	1	5,848	0	16,948
DISH NETWORK CORP CL A	COMMON STOCK	25470M109	40	1,249	SH		DFND	1	336	0	913
DISH NETWORK CORP CL A	COMMON STOCK	25470M109	3	94	SH		OTR	1	53	0	41
DIVERSIFIED HEALTHCARE TRUST	COMMON STOCK	25525P107	21	4,992	SH		DFND	1	1,761	0	3,231
DOLBY LABORATORIES INC CL A	COMMON STOCK	25659T107	236	2,427	SH		DFND	1	1,746	0	681
DOLBY LABORATORIES INC CL A	COMMON STOCK	25659T107	264	2,717	SH		OTR	1	345	34	2,338
DOMINION ENERGY INC	COMMON STOCK	25746U109	8,461	112,521	SH		DFND	1	102,655	0	9,866
DOMINION ENERGY INC	COMMON STOCK	25746U109	4,426	58,846	SH		OTR	1	58,126	114	606
DOMINION RESOURCES INC/VA	COMMON STOCK	25746U109	178	2,369	SH		DFND	5	2,369	0	0
DOMINION ENERGY INC	COMMON STOCK	25746U109	2	18	SH	Put	DFND	1	18	0	0
DOMINOS PIZZA INC	COMMON STOCK	25754A201	1,026	2,675	SH		DFND	1	2,481	0	194
DOMINOS PIZZA INC	COMMON STOCK	25754A201	323	842	SH		OTR	1	516	269	57
DOMINOS PIZZA INC	COMMON STOCK	25754A201	8	22	SH		DFND	5	22	0	0
DONNELLEY FINANCIAL SOLUTIONS	COMMON STOCK	25787G100	2	144	SH		DFND	1	144	0	0
DORCHESTER MINERALS L P	COMMON STOCK	25820R105	26	2,400	SH		DFND	1	2,400	0	0
DORCHESTER MINERALS L P	COMMON STOCK	25820R105	11	1,000	SH		OTR	1	0	1,000	0
DOUGLAS EMMITT INC	COMMON STOCK	25960P109	58	1,972	SH		DFND	1	838	0	1,134
DOUGLAS EMMITT INC	COMMON STOCK	25960P109	1	42	SH		OTR	1	42	0	0
DOUGLAS DYNAMICS INC	COMMON STOCK	25960R105	147	3,435	SH		DFND	1	3,435	0	0
DOUGLAS DYNAMICS INC	COMMON STOCK	25960R105	142	3,310	SH		OTR	1	210	0	3,100
DOUYU INTL HLDGS LTD A D R	COMMON STOCK	25985W105	60	5,394	SH		DFND	1	5,394	0	0
DRAFTKINGS INC COM CL A	COMMON STOCK	26142R104	188	4,036	SH		DFND	1	4,036	0	0
DRAFTKINGS INC COM CL A	COMMON STOCK	26142R104	49	1,050	SH		OTR	1	0	850	200
DRDGOLD LTD A D R	COMMON STOCK	26152H301	1	93	SH		DFND	1	93	0	0
DROPBOX INC CLASS A	COMMON STOCK	26210C104	500	22,523	SH		DFND	1	22,422	0	101
DROPBOX INC CLASS A	COMMON STOCK	26210C104	9	410	SH		OTR	1	354	0	56
DUFF & PHELPS UTIL & CORPORATE	OTHER	26432K108	14	1,400	SH		OTR	1	1,400	0	0
DUFF PHELPS SEL ENRG MLP MIDST	OTHER	26433F207	0	75	SH		DFND	1	75	0	0
DUKE ENERGY HOLDING CORP	COMMON STOCK	26441C204	24,859	271,504	SH		DFND	1	257,873	0	13,631
DUKE ENERGY HOLDING CORP	COMMON STOCK	26441C204	9,813	107,172	SH		OTR	1	99,533	2,006	5,633
DUKE ENERGY CORP	COMMON STOCK	26441C204	362	3,953	SH		DFND	5	3,928	0	25
DULUTH HOLDINGS INC CL B	COMMON STOCK	26443V101	7	700	SH		DFND	1	700	0	0
DULUTH HOLDINGS INC CL B	COMMON STOCK	26443V101	26	2,500	SH		OTR	1	2,500	0	0
DUPONT DE NEMOURS INC WI	COMMON STOCK	26614N102	14,571	204,909	SH		DFND	1	196,436	0	8,473
DUPONT DE NEMOURS INC WI	COMMON STOCK	26614N102	11,895	167,274	SH		OTR	1	158,980	3,536	4,758
DUPONT DE NEMOURS INC	COMMON STOCK	26614N102	53	742	SH		DFND	5	629	0	113
DYNEX CAPITAL INC	COMMON STOCK	26817Q886	76	4,288	SH		DFND	1	277	0	4,011
E L F BEAUTY INC	COMMON STOCK	26856L103	48	1,890	SH		OTR	1	0	1,890	0
E N I SPA A D R	COMMON STOCK	26874R108	290	14,088	SH		DFND	1	4,216	0	9,872
E N I SPA A D R	COMMON STOCK	26874R108	62	3,011	SH		OTR	1	428	873	1,710
ENI SPA	COMMON STOCK	26874R108	2	87	SH		DFND	5	87	0	0
E O G RES INC	COMMON STOCK	26875P101	2,896	58,076	SH		DFND	1	54,110	0	3,966
E O G RES INC	COMMON STOCK	26875P101	572	11,453	SH		OTR	1	7,895	388	3,170
EOG RESOURCES INC	COMMON STOCK	26875P101	25	497	SH		DFND	5	466	0	31
EQT CORP	COMMON STOCK	26884L109	211	16,587	SH		DFND	1	9,358	0	7,229
EQT CORP	COMMON STOCK	26884L109	98	7,729	SH		OTR	1	2,164	0	5,565
EPR PROPERTIES	COMMON STOCK	26884U109	297	9,148	SH		DFND	1	8,154	0	994
EPR PROPERTIES	COMMON STOCK	26884U109	103	3,172	SH		OTR	1	2,667	246	259
DEFIANCE 5G NEXT GEN ETF	OTHER	26922A289	1	20	SH		DFND	1	20	0	0
US VEGAN CLIMATE ETF	OTHER	26922A297	1	40	SH		DFND	1	40	0	0
DISTILLATE US FDMTL STBLTY VAL	OTHER	26922A321	155	4,256	SH		OTR	1	4,256	0	0
CBOE VESTS P500 DIV ARSTCRTS T	OTHER	26922A537	13	263	SH		DFND	1	263	0	0
PREMISE CAPITAL FRONTIER	OTHER	26922A768	93	3,116	SH		DFND	5	3,116	0	0
US GLOBAL JETS ETF	OTHER	26922A842	179	8,000	SH		DFND	1	8,000	0	0
BIOSHARES BIOTECHNOLOGY ETF	OTHER	26923G301	32	645	SH		DFND	1	645	0	0
BIOSHARES BIOTECHNOLOGY ETF	OTHER	26923G301	18	355	SH		OTR	1	355	0	0
VIRTUS REAVES UTILITIES ETF	OTHER	26923G806	20	500	SH		OTR	1	500	0	0
VIRTUS INFRACAP US PREFERRED E	OTHER	26923G822	25	1,150	SH		OTR	1	1,150	0	0
PRIME JUNIOR SILVER MINERS ETF	OTHER	26924G102	127	7,800	SH		DFND	1	7,800	0	0
PRIME JUNIOR SILVER MINERS ETF	OTHER	26924G102	16	981	SH		OTR	1	981	0	0
ETFMG ALTERNATIVE HARVEST ETF	OTHER	26924G508	49	3,427	SH		DFND	1	3,427	0	0
ETFMG ALTERNATIVE HARVEST ETF	OTHER	26924G508	14	1,000	SH		OTR	1	1,000	0	0
ETFMG VIDEO GAME TECH ETF	OTHER	26924G706	79	1,000	SH		DFND	1	1,000	0	0
BLUESTAR TA BIGITECH ISRAEL ET	OTHER	26924G870	775	11,273	SH		DFND	1	11,273	0	0
BLUESTAR TA BIGITECH ISRAEL ET	OTHER	26924G870	15	220	SH		OTR	1	0	0	220
ETHO CLIMATE LEADERSHIP US ETF	OTHER	26924G888	96	1,805	SH		DFND	1	1,805	0	0
EVI INDUSTRIES INC	COMMON STOCK	26929N102	13	420	SH		OTR	1	70	0	350
EAGLE BANCORP MONTANA INC	COMMON STOCK	26942G100	4	211	SH		DFND	1	211	0	0
EAGLE MATERIALS INC	COMMON STOCK	26969P108	1,718	16,953	SH		DFND	1	16,806	0	147
EAGLE MATERIALS INC	COMMON STOCK	26969P108	174	1,716	SH		OTR	1	1,638	60	18
EAGLE MATERIALS INC	COMMON STOCK	26969P108	41	405	SH		DFND	5	405	0	0
EAST WEST BANCORP INC	COMMON STOCK	27579R104	300	5,908	SH		DFND	1	5,494	0	414
EAST WEST BANCORP INC	COMMON STOCK	27579R104	2	41	SH		OTR	1	41	0	0
EASTERLY GOVERNMENT PROPERTI	COMMON STOCK	27616P103	316	13,911	SH		DFND	1	13,795	0	116
EASTERLY GOVERNMENT PROPERTI	COMMON STOCK	27616P103	63	2,760	SH		OTR	1	2,760	0	0
EASTERLY GOVERNMENT PROPERTIES	COMMON STOCK	27616P103	60	2,655	SH		DFND	5	2,655	0	0
EATON VANCE SR INCOME TR	OTHER	27826S103	4	700	SH		DFND	1	0	0	700
EATON VANCE MUN INCOME TR	OTHER	27826U108	174	13,000	SH		DFND	1	13,000	0	0
EATON VANCE INSURED MUNI BND	OTHER	27827X101	11	830	SH		DFND	1	830	0	0
EATON VANCE INSURED MUNI BND	OTHER	27827X101	57	4,200	SH		OTR	1	4,200	0	0
EATON VANCE INSURED NEW YORK	OTHER	27827Y109	86	7,000	SH		DFND	1	7,000	0	0
EATON VANCE TAX ADV DVD INC	OTHER	27828G107	160	6,760	SH		DFND	1	6,760	0	0
EATON VANCE LTD DURATION FND	OTHER	27828H105	28	2,200	SH		DFND	1	2,200	0	0
EATON VANCE LTD DURATION FND	OTHER	27828H105	208	16,600	SH		OTR	1	16,600	0	0
EATON VANCE TX MGD DIV EQ INCO	OTHER	27828N102	6	487	SH		DFND	1	487	0	0
EATON VANCE SR FLTG RATE TR	OTHER	27828Q105	19	1,500	SH		DFND	1	0	0	1,500
EATON VANCE TAX ADV GLBL DVD I	OTHER	27828S101	43	2,335	SH		DFND	1	2,335	0	0
EATON VANCE T/M BUY-WRITE OP	OTHER	27828Y108	61	3,954	SH		DFND	1	3,954	0	0
EV TAX MANAGED GLB B W OPPS	OTHER	27829C105	65	6,715	SH		DFND	1	6,715	0	0
EATON VANCE TAX MGD DIVERSIFIE	OTHER	27829F108	53	6,035	SH		DFND	1	6,035	0	0
EATON VANCE RISK MANAGED DIVER	OTHER	27829G106	202	19,517	SH		DFND	1	19,517	0	0
ECHO GLOBAL LOGISTICS INC	COMMON STOCK	27875T101	20	732	SH		DFND	1	698	0	34
EDGEWELL PERSONAL CARE CO	COMMON STOCK	28035Q102	810	23,414	SH		DFND	1	22,468	0	946
EDGEWELL PERSONAL CARE CO	COMMON STOCK	28035Q102	28	820	SH		OTR	1	820	0	0
EDITAS MEDICINE INC	COMMON STOCK	28106W103	630	8,983	SH		DFND	1	8,837	0	146
EDITAS MEDICINE INC	COMMON STOCK	28106W103	11	150	SH		OTR	1	150	0	0
EHEALTH INC	COMMON STOCK	28238P109	46	649	SH		DFND	1	649	0	0
EIDOS THERAPEUTICS INC	COMMON STOCK	28249H104	3	21	SH		DFND	1	21	0	0
ELANCO ANIMAL HEALTH INC	COMMON STOCK	28414H103	483	15,740	SH		DFND	1	9,276	0	6,464
ELANCO ANIMAL HEALTH INC	COMMON STOCK	28414H103	114	3,712	SH		OTR	1	3,562	0	150
ELEMENT SOLUTIONS INC	COMMON STOCK	28618M106	22	1,224	SH		DFND	1	240	0	984
ELEMENT SOLUTIONS INC	COMMON STOCK	28618M106	44	2,473	SH		DFND	5	2,473	0	0
ELLINGTON FINANCIAL INC	COMMON STOCK	28852N109	37	2,500	SH		DFND	1	2,500	0	0
EMBOTELLADORA ANDINA B A D R	COMMON STOCK	29081P303	4	273	SH		DFND	1	272	0	1
EMBRAER SA SPON A D R	COMMON STOCK	29082A107	132	19,426	SH		DFND	1	18,390	0	1,036
EMBRAER SA SPON A D R	COMMON STOCK	29082A107	17	2,510	SH		OTR	1	249	0	2,261
EMCOR GROUP INC	COMMON STOCK	29084Q100	1,004	10,973	SH		DFND	1	9,619	0	1,354
EMCOR GROUP INC	COMMON STOCK	29084Q100	200	2,192	SH		OTR	1	305	0	1,887
EMERGENT BIOSOLUTIONS INC	COMMON STOCK	29089Q105	1,009	11,259	SH		DFND	1	7,905	0	3,354
EMERGENT BIOSOLUTIONS INC	COMMON STOCK	29089Q105	146	1,632	SH		OTR	1	105	1,527	0
ENBRIDGE INC	COMMON STOCK	29250N105	12,092	378,000	SH		DFND	1	366,546	0	11,454
ENBRIDGE INC	COMMON STOCK	29250N105	3,661	114,440	SH		OTR	1	104,879	906	8,655
ENBRIDGE INC	COMMON STOCK	29250N105	557	17,397	SH		DFND	5	17,333	0	64
ENANTA PHARMACEUTICALS INC	COMMON STOCK	29251M106	41	977	SH		DFND	1	134	0	843
ENDEAVOUR SILVER CORP	COMMON STOCK	29258Y103	5	1,000	SH		DFND	1	1,000	0	0
ENDAVA PLC A D R	COMMON STOCK	29260V105	3	37	SH		DFND	1	29	0	8
ENDAVA PLC A D R	COMMON STOCK	29260V105	104	1,356	SH		OTR	1	800	556	0
ENCOMPASS HEALTH CORPORATION	COMMON STOCK	29261A100	221	2,678	SH		DFND	1	2,403	0	275
ENCOMPASS HEALTH CORPORATION	COMMON STOCK	29261A100	208	2,513	SH		OTR	1	2,030	177	306
ENCOMPASS HEALTH CORP	COMMON STOCK	29261A100	47	574	SH		DFND	5	574	0	0
ENERGY RECOVERY INC	COMMON STOCK	29270J100	10	700	SH		OTR	1	700	0	0
ENDURANCE INTERNATINAL GROUP H	COMMON STOCK	29272B105	4	437	SH		DFND	1	437	0	0
ENERGIZER HLDGS INC	COMMON STOCK	29272W109	1,889	44,787	SH		DFND	1	42,459	0	2,328
ENERGIZER HLDGS INC	COMMON STOCK	29272W109	581	13,779	SH		OTR	1	10,423	3,056	300
ENERGY TRANSFER L P	COMMON STOCK	29273V100	1,035	167,481	SH		DFND	1	167,227	0	254
ENERGY TRANSFER L P	COMMON STOCK	29273V100	228	36,880	SH		OTR	1	17,822	809	18,249
ENERGY TRANSFER EQUITY LP	COMMON STOCK	29273V100	5	771	SH		DFND	5	771	0	0
ENEL AMERICAS SA A D R	COMMON STOCK	29274F104	110	13,351	SH		DFND	1	12,185	0	1,166
ENEL AMERICAS SA A D R	COMMON STOCK	29274F104	14	1,671	SH		OTR	1	1,632	0	39
ENERSYS	COMMON STOCK	29275Y102	1,540	18,544	SH		DFND	1	18,248	0	296
ENERSYS	COMMON STOCK	29275Y102	95	1,145	SH		OTR	1	1,145	0	0
ENERSYS INC	COMMON STOCK	29275Y102	7	82	SH		DFND	5	82	0	0
ENEL CHILE SA A D R	COMMON STOCK	29278D105	32	8,401	SH		DFND	1	4,459	0	3,942
ENEL CHILE SA A D R	COMMON STOCK	29278D105	18	4,736	SH		OTR	1	0	547	4,189
ENLINK MIDSTREAM LLC	COMMON STOCK	29336T100	1	323	SH		DFND	5	323	0	0
ENPHASE ENERGY INC	COMMON STOCK	29355A107	6,395	36,446	SH		DFND	1	36,063	0	383
ENPHASE ENERGY INC	COMMON STOCK	29355A107	296	1,689	SH		OTR	1	441	1,170	78
ENPRO INDUSTRIES INC	COMMON STOCK	29355X107	787	10,417	SH		DFND	1	8,967	0	1,450
ENPRO INDUSTRIES INC	COMMON STOCK	29355X107	123	1,626	SH		OTR	1	1,626	0	0
ENSIGN GROUP INC THE	COMMON STOCK	29358P101	662	9,072	SH		DFND	1	8,843	0	229
ENSIGN GROUP INC THE	COMMON STOCK	29358P101	91	1,254	SH		OTR	1	1,254	0	0
ENSIGN GROUP INC (THE)	COMMON STOCK	29358P101	23	315	SH		DFND	5	315	0	0
ENTEGRIS INC	COMMON STOCK	29362U104	3,617	37,648	SH		DFND	1	36,402	0	1,246
ENTEGRIS INC	COMMON STOCK	29362U104	1,100	11,437	SH		OTR	1	7,755	744	2,938
ENTEGRIS INC	COMMON STOCK	29362U104	98	1,023	SH		DFND	5	983	0	40
ENTERGY CORPORATION	COMMON STOCK	29364G103	945	9,460	SH		DFND	1	8,036	0	1,424
ENTERGY CORPORATION	COMMON STOCK	29364G103	311	3,117	SH		OTR	1	3,056	61	0
ENVESTNET INC	COMMON STOCK	29404K106	1,550	18,830	SH		DFND	1	16,026	0	2,804
ENVESTNET INC	COMMON STOCK	29404K106	894	10,869	SH		OTR	1	611	8,645	1,613
ENVESTNET INC	COMMON STOCK	29404K106	35	420	SH		DFND	5	420	0	0
EPAM SYSTEMS INC	COMMON STOCK	29414B104	1,502	4,192	SH		DFND	1	4,041	0	151
EPAM SYSTEMS INC	COMMON STOCK	29414B104	882	2,463	SH		OTR	1	1,011	132	1,320
EPAM SYSTEMS INC	COMMON STOCK	29414B104	130	363	SH		DFND	5	363	0	0
ENVIVA PARTNERS L P	COMMON STOCK	29414J107	30	650	SH		OTR	1	650	0	0
ENVISTA HOLDINGS CORP	COMMON STOCK	29415F104	157	4,657	SH		DFND	1	50	0	4,607
EPIZYME INC	COMMON STOCK	29428V104	12	1,071	SH		DFND	1	1,071	0	0
EPIZYME INC	COMMON STOCK	29428V104	1	54	SH		OTR	1	54	0	0
EQUINIX INC	COMMON STOCK	29444U700	7,011	9,817	SH		DFND	1	9,142	0	675
EQUINIX INC	COMMON STOCK	29444U700	1,585	2,219	SH		OTR	1	1,586	358	275
EQUINIX INC	COMMON STOCK	29444U700	446	624	SH		DFND	5	624	0	0
EQUILLIUM INC	COMMON STOCK	29446K106	3	494	SH		DFND	1	494	0	0
EQUINOR ASA SPON ADR	COMMON STOCK	29446M102	348	21,178	SH		DFND	1	12,079	0	9,099
EQUINOR ASA SPON ADR	COMMON STOCK	29446M102	53	3,216	SH		OTR	1	1,750	1,431	35
EQUINOR ASA	COMMON STOCK	29446M102	4	246	SH		DFND	5	246	0	0
EQUITY LIFESTYLE PPTYS INC	COMMON STOCK	29472R108	739	11,655	SH		DFND	1	11,223	0	432
EQUITY LIFESTYLE PPTYS INC	COMMON STOCK	29472R108	211	3,330	SH		OTR	1	3,248	52	30
EQUITY RESIDENTIAL	COMMON STOCK	29476L107	799	13,487	SH		DFND	1	10,812	0	2,675
EQUITY RESIDENTIAL	COMMON STOCK	29476L107	702	11,835	SH		OTR	1	9,004	0	2,831
EQUITY RESIDENTIAL COM	COMMON STOCK	29476L107	6	99	SH		DFND	5	0	0	99
ERIE INDEMNITY CO	COMMON STOCK	29530P102	159	648	SH		DFND	1	490	0	158
ERIE INDEMNITY CO	COMMON STOCK	29530P102	84	342	SH		OTR	1	7	39	296
ESPERION THERAPEUTICS INC	COMMON STOCK	29664W105	399	15,333	SH		DFND	1	13,311	0	2,022
ESPERION THERAPEUTICS INC	COMMON STOCK	29664W105	32	1,233	SH		OTR	1	1,233	0	0
ESSENTIAL UTILITIES INC	COMMON STOCK	29670G102	655	13,857	SH		DFND	1	13,226	0	631
ESSENTIAL UTILITIES INC	COMMON STOCK	29670G102	500	10,580	SH		OTR	1	8,710	1,419	451
ETSY INC	COMMON STOCK	29786A106	6,880	38,671	SH		DFND	1	37,959	0	712
ETSY INC	COMMON STOCK	29786A106	1,879	10,558	SH		OTR	1	7,393	64	3,101
ETSY INC	COMMON STOCK	29786A106	37	207	SH		DFND	5	190	0	17
EVERCORE INC	COMMON STOCK	29977A105	2,190	19,980	SH		DFND	1	18,451	0	1,529
EVERCORE INC	COMMON STOCK	29977A105	405	3,690	SH		OTR	1	3,690	0	0
EVERCORE PARTNERS INC	COMMON STOCK	29977A105	20	178	SH		DFND	5	178	0	0
EVERBRIDGE INC	COMMON STOCK	29978A104	392	2,628	SH		DFND	1	2,619	0	9
EVERBRIDGE INC	COMMON STOCK	29978A104	283	1,899	SH		OTR	1	749	66	1,084
EVERI HOLDINGS INC	COMMON STOCK	30034T103	8	561	SH		DFND	1	561	0	0
EVERI HOLDINGS INC	COMMON STOCK	30034T103	70	5,069	SH		OTR	1	0	5,069	0
EVERI HOLDINGS INC	COMMON STOCK	30034T103	0	4	SH		DFND	5	4	0	0
EVERGY INC	COMMON STOCK	30034W106	1,300	23,414	SH		DFND	1	21,533	0	1,881
EVERGY INC	COMMON STOCK	30034W106	969	17,455	SH		OTR	1	16,222	17	1,216
EVERTEC INC	COMMON STOCK	30040P103	51	1,307	SH		OTR	1	0	1,307	0
EVERSOURCE ENERGY	COMMON STOCK	30040W108	3,821	44,168	SH		DFND	1	40,967	0	3,201
EVERSOURCE ENERGY	COMMON STOCK	30040W108	823	9,516	SH		OTR	1	8,241	1,145	130
EVERSOURCE ENERGY	COMMON STOCK	30040W108	64	742	SH		DFND	5	742	0	0
EVERQUOTE INC CLASS A	COMMON STOCK	30041R108	20	541	SH		DFND	1	541	0	0
EVOFEM BIOSCIENCES INC	COMMON STOCK	30048L104	2	820	SH		DFND	1	820	0	0
EVOFEM BIOSCIENCES INC	COMMON STOCK	30048L104	0	1	SH		DFND	5	1	0	0
EVOLUTION PETROLEUM CORP	COMMON STOCK	30049A107	1	250	SH		DFND	1	250	0	0
EVOLENT HEALTH INC A	COMMON STOCK	30050B101	400	24,976	SH		DFND	1	23,996	0	980
EVOLENT HEALTH INC A	COMMON STOCK	30050B101	95	5,952	SH		OTR	1	393	5,559	0
EVOLUS INC	COMMON STOCK	30052C107	5	1,375	SH		DFND	1	1,375	0	0
EVOQUA WATER TECHNOLOGIES CO	COMMON STOCK	30057T105	4	139	SH		DFND	1	139	0	0
EVOQUA WATER TECHNOLOGIES CO	COMMON STOCK	30057T105	1	37	SH		OTR	1	0	0	37
EXACT SCIENCES CORP	COMMON STOCK	30063P105	10,556	79,678	SH		DFND	1	75,932	0	3,746
EXACT SCIENCES CORP	COMMON STOCK	30063P105	2,474	18,674	SH		OTR	1	12,276	5,055	1,343
EXACT SCIENCES CORPORATION	COMMON STOCK	30063P105	58	434	SH		DFND	5	415	0	19
EXANTAS CAPITAL CORP	COMMON STOCK	30068N105	48	12,133	SH		DFND	1	12,133	0	0
EXELON CORPORATION	COMMON STOCK	30161N101	6,479	153,441	SH		DFND	1	145,912	0	7,529
EXELON CORPORATION	COMMON STOCK	30161N101	1,531	36,271	SH		OTR	1	34,359	1,053	859
EXELIXIS INC	COMMON STOCK	30161Q104	448	22,338	SH		DFND	1	19,253	0	3,085
EXELIXIS INC	COMMON STOCK	30161Q104	179	8,915	SH		OTR	1	319	0	8,596
EXPEDIA INC	COMMON STOCK	30212P303	254	1,920	SH		DFND	1	983	0	937
EXPEDIA INC	COMMON STOCK	30212P303	697	5,260	SH		OTR	1	80	188	4,992
EXPEDIA INC	COMMON STOCK	30212P303	3	21	SH		DFND	5	11	0	10
EXP WORLD HOLDINGS INC	COMMON STOCK	30212W100	2	34	SH		DFND	1	0	0	34
EXP WORLD HOLDINGS INC	COMMON STOCK	30212W100	3	50	SH		OTR	1	50	0	0
EXPONENT INC	COMMON STOCK	30214U102	1,939	21,529	SH		DFND	1	17,948	0	3,581
EXPONENT INC	COMMON STOCK	30214U102	446	4,955	SH		OTR	1	1,212	873	2,870
EXPONENT INC	COMMON STOCK	30214U102	274	3,036	SH		DFND	5	3,036	0	0
EXTENDED STAY AMERICA INC	COMMON STOCK	30224P200	104	7,054	SH		DFND	1	2,990	0	4,064
EXTENDED STAY AMERICA INC	COMMON STOCK	30224P200	2	145	SH		OTR	1	0	0	145
EXTRA SPACE STORAGE INC	COMMON STOCK	30225T102	2,436	21,031	SH		DFND	1	20,478	0	553
EXTRA SPACE STORAGE INC	COMMON STOCK	30225T102	595	5,132	SH		OTR	1	5,102	12	18
EXTRA SPACE STORAGE INC	COMMON STOCK	30225T102	66	566	SH		DFND	5	566	0	0
EXXON MOBIL CORP	COMMON STOCK	30231G102	108,186	2,624,624	SH		DFND	1	2,220,094	0	404,530
EXXON MOBIL CORP	COMMON STOCK	30231G102	72,967	1,770,192	SH		OTR	1	1,682,638	49,267	38,287
EXXON MOBIL CORPORATION	COMMON STOCK	30231G102	310	7,510	SH		DFND	5	7,468	0	42
EXXON MOBIL CORP	COMMON STOCK	30231G102	1	14	SH	Put	DFND	1	14	0	0
EYEGATE PHARMACEUTICALS INC	COMMON STOCK	30233M503	13	2,710	SH		OTR	1	2,710	0	0
FBL FINL GROUP INC CL A	COMMON STOCK	30239F106	29	551	SH		DFND	1	140	0	411
FB FINANCIAL CORP	COMMON STOCK	30257X104	2	54	SH		DFND	1	54	0	0
FACEBOOK INC CLASS A	COMMON STOCK	30303M102	220,747	808,125	SH		DFND	1	769,862	0	38,263
FACEBOOK INC CLASS A	COMMON STOCK	30303M102	46,026	168,493	SH		OTR	1	140,900	15,116	12,477
FACEBOOK	COMMON STOCK	30303M102	2,241	8,204	SH		DFND	5	8,074	0	130
FANHUA INC A D R	COMMON STOCK	30712A103	36	2,991	SH		DFND	1	2,841	0	150
FARMLAND PARTNERS INC	COMMON STOCK	31154R109	1	100	SH		OTR	1	0	0	100
FASTLY INC CLASS A	COMMON STOCK	31188V100	103	1,177	SH		DFND	1	1,177	0	0
FASTLY INC CLASS A	COMMON STOCK	31188V100	70	800	SH		OTR	1	800	0	0
FATE THERAPEUTICS INC	COMMON STOCK	31189P102	9	96	SH		DFND	1	0	0	96
FATE THERAPEUTICS INC	COMMON STOCK	31189P102	2	19	SH		OTR	1	0	0	19
FEDERATED HERMES PREMIER MUNIC	OTHER	31423P108	1,339	90,504	SH		DFND	1	90,504	0	0
FED EX CORP	COMMON STOCK	31428X106	19,163	73,812	SH		DFND	1	71,571	0	2,241
FED EX CORP	COMMON STOCK	31428X106	8,037	30,956	SH		OTR	1	28,651	1,135	1,170
FEDEX CORPORATION	COMMON STOCK	31428X106	9	35	SH		DFND	5	35	0	0
FIBROGEN INC	COMMON STOCK	31572Q808	419	11,296	SH		DFND	1	11,285	0	11
FIBROGEN INC	COMMON STOCK	31572Q808	24	654	SH		OTR	1	180	0	474
FIDELITY NATIONAL INFO SERV	COMMON STOCK	31620M106	16,892	119,412	SH		DFND	1	112,951	0	6,461
FIDELITY NATIONAL INFO SERV	COMMON STOCK	31620M106	3,609	25,517	SH		OTR	1	23,102	1,583	832
FIDELITY NATL INFO SVCS INC	COMMON STOCK	31620M106	597	4,215	SH		DFND	5	4,215	0	0
FIDELITY NATIONAL INFO SERV	COMMON STOCK	31620M106	1	7	SH	Put	DFND	1	7	0	0
FIDELITY NATIONAL FINANCIAL IN	COMMON STOCK	31620R303	1,106	28,303	SH		DFND	1	26,813	0	1,490
FIDELITY NATIONAL FINANCIAL IN	COMMON STOCK	31620R303	222	5,684	SH		OTR	1	1,975	3,709	0
FNF GROUP	COMMON STOCK	31620R303	66	1,689	SH		DFND	5	1,689	0	0
FID CLAYMORE ENERGY INFRASTRUC	OTHER	31647Q205	0	47	SH		DFND	1	47	0	0
FINVOLUTION GROUP A D R	COMMON STOCK	31810T101	476	178,168	SH		DFND	1	174,716	0	3,452
FINVOLUTION GROUP A D R	COMMON STOCK	31810T101	80	29,909	SH		OTR	1	29,909	0	0
FINVOLUTION GROUP	COMMON STOCK	31810T101	1	367	SH		DFND	5	367	0	0
FIREEYE INC	COMMON STOCK	31816Q101	1,944	84,303	SH		DFND	1	84,303	0	0
FIREEYE INC	COMMON STOCK	31816Q101	332	14,386	SH		OTR	1	14,386	0	0
FIRST AMERICAN FINANCIAL	COMMON STOCK	31847R102	254	4,920	SH		DFND	1	4,513	0	407
FIRST AMERICAN FINANCIAL	COMMON STOCK	31847R102	114	2,200	SH		OTR	1	2,000	0	200
FIRST AMERICAN FINANCIAL	COMMON STOCK	31847R102	4	70	SH		DFND	5	70	0	0
FIRST CTZNS BANCSHARES INC CL	COMMON STOCK	31946M103	86	150	SH		DFND	1	101	0	49
FIRST FINL BANKSHARES INC	COMMON STOCK	32020R109	53	1,456	SH		DFND	1	1,255	0	201
FIRST FINL BANKSHARES INC	COMMON STOCK	32020R109	11	283	SH		OTR	1	212	0	71
FIRST FOUNDATION INC	COMMON STOCK	32026V104	2	97	SH		DFND	1	97	0	0
FIRST HAWAIIAN INC	COMMON STOCK	32051X108	57	2,429	SH		DFND	1	1,541	0	888
FIRST HAWAIIAN INC	COMMON STOCK	32051X108	3	140	SH		OTR	1	0	0	140
FIRST INDL RLTY TR INC	COMMON STOCK	32054K103	1,604	38,068	SH		DFND	1	38,019	0	49
FIRST INDL RLTY TR INC	COMMON STOCK	32054K103	389	9,224	SH		OTR	1	8,135	1,089	0
FIRST INDUSTRIAL REALTY TRUST	COMMON STOCK	32054K103	2	50	SH		DFND	5	50	0	0
FIRST INTERSTATE BANCSYS MT	COMMON STOCK	32055Y201	388	9,509	SH		DFND	1	5,891	0	3,618
FIRST INTERSTATE BANCSYS MT	COMMON STOCK	32055Y201	65	1,600	SH		OTR	1	0	1,600	0
FIRST MAJESTIC SILVER CORP	COMMON STOCK	32076V103	3	216	SH		DFND	1	92	0	124
FIRST REPUBLIC BANK SAN FRAN	COMMON STOCK	33616C100	6,071	41,324	SH		DFND	1	39,896	0	1,428
FIRST REPUBLIC BANK SAN FRAN	COMMON STOCK	33616C100	2,422	16,478	SH		OTR	1	6,371	0	10,107
FIRST REP BK SAN FRANCISCO CA	COMMON STOCK	33616C100	651	4,429	SH		DFND	5	4,413	0	16
FIRST TRUST INTERMEDIATE DUR	OTHER	33718W103	10	397	SH		DFND	1	397	0	0
FIRST TRUST ISE WATER IDX ETF	OTHER	33733B100	6,123	84,896	SH		DFND	1	82,872	0	2,024
FIRST TRUST ISE WATER IDX ETF	OTHER	33733B100	844	11,700	SH		OTR	1	10,900	0	800
FIRST TRUST NASDAQ 100 EX TECH	OTHER	33733E401	43	545	SH		DFND	1	545	0	0
FIRST TRUST NASDAQ CLEAN EDGE	OTHER	33733E500	84	1,195	SH		DFND	5	0	0	1,195
FIRST TR VALUE LINE DVD INDX F	OTHER	33734H106	350	9,981	SH		DFND	1	9,981	0	0
FIRST TRUST VALUE LINE DVD	OTHER	33734H106	15,375	438,350	SH		DFND	5	122,811	0	315,539
FIRST TRUST LARGE CAP CORE ETF	OTHER	33734K109	417	5,682	SH		DFND	1	5,682	0	0
FIRST TRUST LARGE CAP CORE ETF	OTHER	33734K109	35	480	SH		OTR	1	480	0	0
FIRST TRUST LARGE CAP CORE	OTHER	33734K109	4,276	58,240	SH		DFND	5	1,893	0	56,347
FIRST TRUST CONSUMER DISCRET E	OTHER	33734X101	25	490	SH		DFND	1	490	0	0
FIRST TRUST CONSMR DISCRETNARY	OTHER	33734X101	46	904	SH		DFND	5	0	0	904
FIRST TRUST CONSUMER STAPLES E	OTHER	33734X119	45	870	SH		DFND	1	870	0	0
FIRST TRUST CONSUMER STAPLES	OTHER	33734X119	58	1,120	SH		DFND	5	0	0	1,120
FIRST TRUST FINANCIAL ALPHAD E	OTHER	33734X135	6	174	SH		DFND	1	174	0	0
FIRST TRUST FINANCIAL ALPHADEX	OTHER	33734X135	215	6,286	SH		DFND	5	0	0	6,286
FIRST TR EXCHANGE TRADED FD ET	OTHER	33734X143	65	605	SH		DFND	1	605	0	0
FIRST TRUST HEALTH CARE	OTHER	33734X143	84	780	SH		DFND	5	0	0	780
FIRST TRUST TECHNOLOGY ALPHA E	OTHER	33734X176	115	1,030	SH		DFND	1	1,030	0	0
FIRST TRUST TECH ALPHADEX	OTHER	33734X176	3,610	32,286	SH		DFND	5	0	0	32,286
FIRST TRUST CLOUD COMPUTING ET	OTHER	33734X192	408	4,296	SH		DFND	1	4,296	0	0
FIRST TRUST CLOUD COMPUTING ET	OTHER	33734X192	404	4,250	SH		OTR	1	4,250	0	0
FIRST TRUST ISE CLOUD COMPUTNG	OTHER	33734X192	13	138	SH		DFND	5	0	0	138
FIRST TRUST DOW JONES GL SEL E	OTHER	33734X200	58	2,635	SH		DFND	1	2,635	0	0
FIRST TRUST DOW JONES GL SEL E	OTHER	33734X200	112	5,100	SH		OTR	1	5,100	0	0
FIRST TRUST NASDAQ CYBERSECURI	OTHER	33734X846	74	1,675	SH		DFND	1	1,675	0	0
FIRST TRUST INTL IPO	OTHER	33734X853	1,007	14,639	SH		DFND	5	0	0	14,639
FIRST TR SM CAP CORE ALPHADEX	OTHER	33734Y109	130	1,688	SH		DFND	5	0	0	1,688
FIRST TRUST MID CAP CORE ALP E	OTHER	33735B108	26	309	SH		DFND	1	309	0	0
FIRST TRUST MID CAP CORE	OTHER	33735B108	81	978	SH		DFND	5	381	0	597
FIRST TRUST LARGE CAP VALUE ET	OTHER	33735J101	1,224	22,289	SH		DFND	1	22,289	0	0
FIRST TRUST LARGE CAP VALUE	OTHER	33735J101	49	897	SH		DFND	5	0	0	897
FIRST TRUST LARGE CAP GR ETF	OTHER	33735K108	80	823	SH		DFND	1	823	0	0
FIRST TRUST LRG CAP GROWTH OPP	OTHER	33735K108	1,127	11,659	SH		DFND	5	11,606	0	53
FIRST TR GLB WIND ENGY ETF	OTHER	33736G106	374	16,059	SH		DFND	1	16,059	0	0
FIRST TR GLB WIND ENGY ETF	OTHER	33736G106	35	1,500	SH		OTR	1	1,500	0	0
FIRST TRUST ASIAPAC EX JPN ETF	OTHER	33737J109	6	190	SH		DFND	1	190	0	0
FIRST TRUST EUROPE ALPHADEX ET	OTHER	33737J117	12	310	SH		DFND	1	310	0	0
FIRST TR EUROPE ALPHADEX FUND	OTHER	33737J117	170	4,468	SH		DFND	5	0	0	4,468
FIRST TRUST INDXX NEXTG ETF	OTHER	33737K205	30	432	SH		DFND	1	432	0	0
FIRST TRUST NASDAQ SMARTPHONE	OTHER	33737K205	1,474	21,300	SH		DFND	5	0	0	21,300
FIRST TRUST MID CAP GROWTH ETF	OTHER	33737M102	29	443	SH		DFND	1	443	0	0
FT SMALL CAP VALUE ALPHADEX FD	OTHER	33737M409	1,160	27,870	SH		DFND	5	0	0	27,870
FIRST TRUST NORTH AMERICAN ETF	OTHER	33738D101	10	500	SH		DFND	1	500	0	0
FIRST TRUST SENIOR LOAN ETF	OTHER	33738D309	51	1,067	SH		DFND	1	1,067	0	0
FIRST TRUST SENIOR LOAN	OTHER	33738D309	1	20	SH		DFND	5	0	0	20
FIRST TRUST HIGH YIELD LONG	OTHER	33738D408	1,818	37,307	SH		DFND	5	0	0	37,307
FIRST TRUST MULTI ASSET DIV ET	OTHER	33738R100	91	6,050	SH		DFND	1	4,250	0	1,800
FIRST TRUST NASDAQ TECHNOLOGY	OTHER	33738R118	6	122	SH		DFND	1	122	0	0
FT	OTHER	33738R118	35	705	SH		DFND	5	0	0	705
FIRST TRUST LOW BETA INCOME	OTHER	33738R407	15	738	SH		DFND	5	0	0	738
FT NASDAQ RISING DVD ACHIEVERS	OTHER	33738R506	5,827	145,843	SH		DFND	5	17,755	0	128,088
FIRST TRUST DORSEY WRIGHT FOCU	OTHER	33738R605	51	1,254	SH		DFND	1	1,254	0	0
FT DORSEY WRIGHT FOCUS FIVE	OTHER	33738R605	85	2,111	SH		DFND	5	0	0	2,111
FIRST TRUST S P INT DIV ETF	OTHER	33738R688	19	1,140	SH		DFND	1	1,140	0	0
FIRST TRUST NASDAQ BANK	OTHER	33738R860	38	1,601	SH		DFND	5	0	0	1,601
FIRST TRUST MANAGED MUNICIPAL	OTHER	33739N108	370	6,524	SH		DFND	5	1,220	0	5,304
FIRST TRUST LOW DURATION MTGE	OTHER	33739Q200	11,633	226,015	SH		DFND	5	606	0	225,409
FIRST TRUST ENHANCED SHORT	OTHER	33739Q408	4,355	72,537	SH		DFND	5	0	0	72,537
FIRST TRUST SSI STRATEGIC	OTHER	33739Q507	453	9,240	SH		DFND	5	0	0	9,240
FIRST TRUST TCW OPPORTUNISTIC	OTHER	33740F805	94	1,725	SH		DFND	1	1,725	0	0
FIRST TRUST TCW OPPORTUNISTIC	OTHER	33740F805	5,609	102,467	SH		DFND	5	0	0	102,467
FIRSTCASH INC	COMMON STOCK	33767D105	701	10,013	SH		DFND	1	10,013	0	0
FIRSTCASH INC	COMMON STOCK	33767D105	116	1,653	SH		OTR	1	1,095	558	0
FITBIT INC A	COMMON STOCK	33812L102	13	1,848	SH		DFND	1	1,739	0	109
FIVE BELOW	COMMON STOCK	33829M101	10,136	57,926	SH		DFND	1	57,256	0	670
FIVE BELOW	COMMON STOCK	33829M101	2,801	16,005	SH		OTR	1	15,509	432	64
FIVE BELOW INC	COMMON STOCK	33829M101	535	3,058	SH		DFND	5	3,041	0	17
FIVEPRIME THERAPEUTICS INC	COMMON STOCK	33830X104	2	114	SH		DFND	1	0	0	114
FIVE STAR SENIOR LIVING INC	COMMON STOCK	33832D205	0	31	SH		DFND	1	31	0	0
FIVE POINT HOLDINGS LLC CL A	COMMON STOCK	33833Q106	1	189	SH		DFND	1	0	0	189
FLEXSHARES GLOBAL UPSTREAM N E	OTHER	33939L407	7	223	SH		DFND	1	223	0	0
FLEXSHARES TR IBOXX 5YR TRGT E	OTHER	33939L605	16	570	SH		DFND	1	570	0	0
FLEXSHARES GLBL QUALITY REAL E	OTHER	33939L787	3	50	SH		DFND	1	50	0	0
FLEXSHARES STOXX GLOBAL BROAD	OTHER	33939L795	9	160	SH		DFND	1	160	0	0
FLEXSHARES QUALITY DIVIDEND ET	OTHER	33939L845	14	293	SH		DFND	1	293	0	0
FLEXSHARES QUALITY DIVIDEND ET	OTHER	33939L845	101	2,100	SH		OTR	1	2,100	0	0
FLEXSHARES QUALITY DIVIDEND ET	OTHER	33939L860	3	68	SH		DFND	1	68	0	0
FLEXSHARES QUALITY DIVIDEND ET	OTHER	33939L860	235	4,750	SH		OTR	1	4,750	0	0
FLOWSERVE CORP	COMMON STOCK	34354P105	174	4,734	SH		DFND	1	3,417	0	1,317
FLOWSERVE CORP	COMMON STOCK	34354P105	144	3,907	SH		OTR	1	3,904	0	3
FLY LEASING LTD A D R	COMMON STOCK	34407D109	2,317	235,234	SH		DFND	1	234,561	0	673
FLY LEASING LTD A D R	COMMON STOCK	34407D109	573	58,236	SH		OTR	1	58,236	0	0
FOCUS FINANCIAL PARTNERS A	COMMON STOCK	34417P100	2,437	56,015	SH		DFND	1	55,973	0	42
FOCUS FINANCIAL PARTNERS A	COMMON STOCK	34417P100	61	1,406	SH		OTR	1	1,406	0	0
FORTIVE CORP WI	COMMON STOCK	34959J108	6,021	85,020	SH		DFND	1	81,901	0	3,119
FORTIVE CORP WI	COMMON STOCK	34959J108	827	11,681	SH		OTR	1	4,975	31	6,675
FORTIVE CORP	COMMON STOCK	34959J108	148	2,085	SH		DFND	5	1,933	0	152
FORTERRA INC	COMMON STOCK	34960W106	3	179	SH		DFND	1	179	0	0
FORTUNE BRANDS HOME & SECURI	COMMON STOCK	34964C106	2,204	25,707	SH		DFND	1	24,283	0	1,424
FORTUNE BRANDS HOME & SECURI	COMMON STOCK	34964C106	761	8,882	SH		OTR	1	7,845	0	1,037
FORTUNE BRANDS HOME & SEC INC	COMMON STOCK	34964C106	30	344	SH		DFND	5	344	0	0
FOSSIL GROUP INC	COMMON STOCK	34988V106	4	418	SH		DFND	1	0	0	418
FOUR CORNERS PROPERTY TRUST	COMMON STOCK	35086T109	257	8,647	SH		DFND	1	4,048	0	4,599
FOUR CORNERS PROPERTY TRUST	COMMON STOCK	35086T109	523	17,584	SH		OTR	1	5,190	12,359	35
FOX CORP CLASS A W I	COMMON STOCK	35137L105	191	6,548	SH		DFND	1	4,790	0	1,758
FOX CORP CLASS A W I	COMMON STOCK	35137L105	45	1,547	SH		OTR	1	1,503	0	44
FOX CORP	COMMON STOCK	35137L105	2	64	SH		DFND	5	0	0	64
FOX CORP CLASS B	COMMON STOCK	35137L204	91	3,172	SH		DFND	1	1,912	0	1,260
FOX CORP CLASS B	COMMON STOCK	35137L204	1	41	SH		OTR	1	0	0	41
FOX FACTORY HOLDING CORP	COMMON STOCK	35138V102	7,782	73,613	SH		DFND	1	70,670	0	2,943
FOX FACTORY HOLDING CORP	COMMON STOCK	35138V102	892	8,437	SH		OTR	1	5,016	586	2,835
FRANCHISE GROUP INC	COMMON STOCK	35180X105	7	242	SH		DFND	1	0	0	242
FRANKLIN STR PPTYS CORP	COMMON STOCK	35471R106	14	3,210	SH		DFND	1	3,210	0	0
FRANKLIN LIBERTY US LOW	OTHER	35473P504	33	773	SH		DFND	5	0	0	773
FREEPORT MCMORAN INC	COMMON STOCK	35671D857	1,430	54,935	SH		DFND	1	46,039	0	8,896
FREEPORT MCMORAN INC	COMMON STOCK	35671D857	1,001	38,471	SH		OTR	1	37,018	921	532
FREEPORT MCMORAN COPPER & GOLD	COMMON STOCK	35671D857	0	0	SH		DFND	5	0	0	0
FRONT YARD RESIDENTIAL CORP	COMMON STOCK	35904G107	8	479	SH		DFND	1	0	0	479
FRONTDOOR INC	COMMON STOCK	35905A109	1,775	35,355	SH		DFND	1	35,296	0	59
FRONTDOOR INC	COMMON STOCK	35905A109	51	1,020	SH		OTR	1	1,020	0	0
FRONTDOOR INC	COMMON STOCK	35905A109	40	794	SH		DFND	5	794	0	0
FUELCELL ENERGY INC	COMMON STOCK	35952H601	19	1,719	SH		DFND	1	1,719	0	0
FS KKR CAPITAL CORP II	COMMON STOCK	35952V303	9	524	SH		DFND	5	0	0	524
FUTUREFUEL CORPORATION	COMMON STOCK	36116M106	2	142	SH		DFND	1	9	0	133
FUTUREFUEL CORPORATION	COMMON STOCK	36116M106	157	12,400	SH		OTR	1	12,400	0	0
FUTU HOLDINGS LTD A D R	COMMON STOCK	36118L106	200	4,377	SH		DFND	1	4,167	0	210
FUTU HOLDINGS LTD A D R	COMMON STOCK	36118L106	8	170	SH		OTR	1	0	170	0
GEO GROUP INC THE	COMMON STOCK	36162J106	6	639	SH		DFND	1	301	0	338
GCP APPLIED TECHNOLOGIES INC	COMMON STOCK	36164Y101	10	440	SH		DFND	1	440	0	0
GDS HOLDING LTD A D R	COMMON STOCK	36165L108	1,019	10,877	SH		DFND	1	7,242	0	3,635
GDS HOLDING LTD A D R	COMMON STOCK	36165L108	26	278	SH		OTR	1	267	0	11
GW PHARMACEUTICALS A D R	COMMON STOCK	36197T103	99	860	SH		DFND	1	860	0	0
GW PHARMACEUTICALS A D R	COMMON STOCK	36197T103	1,808	15,670	SH		OTR	1	15,550	0	120
G1 THERAPEUTICS INC	COMMON STOCK	3621LQ109	21	1,184	SH		DFND	1	0	0	1,184
G I I I APPAREL GROUP LTD	COMMON STOCK	36237H101	698	29,401	SH		DFND	1	28,073	0	1,328
G I I I APPAREL GROUP LTD	COMMON STOCK	36237H101	126	5,304	SH		OTR	1	4,947	357	0
G-III APPAREL	COMMON STOCK	36237H101	7	314	SH		DFND	5	314	0	0
THE GABELLI MULTIMEDIA TRUST I	OTHER	36239Q109	0	35	SH		OTR	1	35	0	0
GABELLI UTIL FD	OTHER	36240A101	1	138	SH		OTR	1	138	0	0
GSI TECHNOLOGY INC	COMMON STOCK	36241U106	7	1,000	SH		DFND	1	1,000	0	0
GABELLI HEALTHCARE WELLNESS	OTHER	36246K103	1	87	SH		OTR	1	87	0	0
GMS INC	COMMON STOCK	36251C103	4	130	SH		DFND	1	130	0	0
GALAPAGOS NV SPON A D R	COMMON STOCK	36315X101	22	223	SH		DFND	1	90	0	133
GALAPAGOS NV SPON A D R	COMMON STOCK	36315X101	13	131	SH		OTR	1	31	100	0
GAMCO GLOBAL GOLD NAT RES INC	OTHER	36465A109	19	5,354	SH		DFND	1	5,354	0	0
GAMCO GLOBAL GOLD NAT RES INC	OTHER	36465A109	63	17,930	SH		OTR	1	17,930	0	0
GAMING & LEISURE PROPE W I	COMMON STOCK	36467J108	649	15,314	SH		DFND	1	13,930	0	1,384
GAMING & LEISURE PROPE W I	COMMON STOCK	36467J108	58	1,378	SH		OTR	1	1,232	0	146
GAMING AND LEISURE PROPERTIES	COMMON STOCK	36467J108	1	28	SH		DFND	5	28	0	0
GAMESTOP CORP CLASS A	COMMON STOCK	36467W109	5	288	SH		DFND	1	0	0	288
GAMESTOP CORP CLASS A	COMMON STOCK	36467W109	38	2,000	SH		OTR	1	2,000	0	0
GANNETT CO INC	COMMON STOCK	36472T109	1	237	SH		DFND	1	237	0	0
GANNETT CO INC	COMMON STOCK	36472T109	0	27	SH		OTR	1	27	0	0
GENERAL MOTORS CO	COMMON STOCK	37045V100	1,177	28,262	SH		DFND	1	19,904	0	8,358
GENERAL MOTORS CO	COMMON STOCK	37045V100	691	16,599	SH		OTR	1	16,599	0	0
GENERAL MOTORS CO	COMMON STOCK	37045V100	4	98	SH		DFND	5	98	0	0
GENESIS HEALTHCARE INC	COMMON STOCK	37185X106	0	1,000	SH		DFND	1	1,000	0	0
GENWORTH FINANCIAL INC CL A	COMMON STOCK	37247D106	10	2,565	SH		DFND	1	2,344	0	221
GENWORTH FINANCIAL INC CL A	COMMON STOCK	37247D106	15	4,071	SH		OTR	1	0	0	4,071
GENTHERM INC	COMMON STOCK	37253A103	38	583	SH		DFND	1	583	0	0
GENTHERM INC	COMMON STOCK	37253A103	5	69	SH		OTR	1	69	0	0
GLACIER BANCORP INC NEW	COMMON STOCK	37637Q105	364	7,908	SH		DFND	1	4,708	0	3,200
GLACIER BANCORP INC NEW	COMMON STOCK	37637Q105	2	49	SH		OTR	1	0	0	49
GLACIER BANCORP INC	COMMON STOCK	37637Q105	34	737	SH		DFND	5	737	0	0
GLAXO SMITHKLINE P L C A D R	COMMON STOCK	37733W105	3,229	87,761	SH		DFND	1	79,147	0	8,614
GLAXO SMITHKLINE P L C A D R	COMMON STOCK	37733W105	704	19,122	SH		OTR	1	16,847	847	1,428
GLAXOSMITHKLINE PLC	COMMON STOCK	37733W105	17	454	SH		DFND	5	454	0	0
GLOBAL BLOOD THERAPEUTICS IN	COMMON STOCK	37890U108	18	406	SH		DFND	1	222	0	184
GLOBAL PAYMENTS INC	COMMON STOCK	37940X102	8,375	38,880	SH		DFND	1	37,330	0	1,550
GLOBAL PAYMENTS INC	COMMON STOCK	37940X102	2,312	10,736	SH		OTR	1	7,871	724	2,141
GLOBAL PAYMENT INC	COMMON STOCK	37940X102	94	438	SH		DFND	5	425	0	13
GLOBAL X FDS	OTHER	37950E549	13	1,000	SH		DFND	1	1,000	0	0
GLOBAL X FDS	OTHER	37950E549	417	32,500	SH		OTR	1	32,500	0	0
GLOBAL MEDICAL REIT INC	COMMON STOCK	37954A204	2	143	SH		DFND	1	143	0	0
GLOBAL X FDS GLOBAL ETF	OTHER	37954Y343	9	313	SH		DFND	1	313	0	0
GLOBAL X S P 500 COVE CALL E E	OTHER	37954Y475	41	882	SH		DFND	1	882	0	0
GLOBAL X NASD 100 COV CALL ETF	OTHER	37954Y483	46	2,000	SH		DFND	1	2,000	0	0
GLOBAL X NASD 100 COVERED CALL	OTHER	37954Y483	700	30,711	SH		DFND	5	525	0	30,186
GLOBAL X DAX GERMANY ETF	OTHER	37954Y491	60	1,952	SH		DFND	1	1,952	0	0
GLOBAL X ATNMS ELCTRC VHCLS ET	OTHER	37954Y624	58	2,400	SH		DFND	1	2,400	0	0
GLOBAL X YIELDCO RENEWABLE ETF	OTHER	37954Y707	10	550	SH		DFND	1	550	0	0
GLOBAL X ROBOTICS ARTFCL ETF	OTHER	37954Y715	26	775	SH		DFND	1	775	0	0
GLOBAL X FINTECH ETF	OTHER	37954Y814	9	200	SH		OTR	1	200	0	0
GLOBAL X FDS GLOBAL X SILVER E	OTHER	37954Y848	132	2,900	SH		DFND	1	2,900	0	0
GLOBAL X FDS GLOBAL X SILVER E	OTHER	37954Y848	29	635	SH		OTR	1	635	0	0
GLOBAL X LITHIUM BATT TH ETF	OTHER	37954Y855	440	7,103	SH		DFND	1	7,103	0	0
GLOBAL X LITHIUM BATT TH ETF	OTHER	37954Y855	110	1,782	SH		OTR	1	1,782	0	0
GOL LINHAS AEREAS A D R	COMMON STOCK	38045R206	59	6,030	SH		DFND	1	4,712	0	1,318
GOLD FIELDS LTD SPSD A D R	COMMON STOCK	38059T106	173	18,626	SH		DFND	1	13,953	0	4,673
GOLDEN NUGGET ONLINE GAMING	COMMON STOCK	38113L107	4	200	SH		OTR	1	0	0	200
GOLDMAN SACHS GROUP INC	COMMON STOCK	38141G104	22,113	83,858	SH		DFND	1	80,925	0	2,933
GOLDMAN SACHS GROUP INC	COMMON STOCK	38141G104	10,362	39,292	SH		OTR	1	33,426	2,066	3,800
GOLDMAN SACHS GROUP INC	COMMON STOCK	38141G104	27	102	SH		DFND	5	102	0	0
GOLDMAN SACHS MLP ENERGY RENAI	OTHER	38148G206	6	747	SH		DFND	1	747	0	0
GOLDMAN SACHS MLP ENERGY RENAI	OTHER	38148G206	4	555	SH		OTR	1	555	0	0
GOLDMAN SACHS PHYSICAL GOLD ET	OTHER	38150K103	14	764	SH		DFND	1	764	0	0
GOLUB CAPITAL BDC INC	COMMON STOCK	38173M102	311	21,998	SH		DFND	1	21,614	0	384
GOODRX HLDGS INC	COMMON STOCK	38246G108	499	12,361	SH		DFND	1	12,361	0	0
GOODRX HLDGS INC	COMMON STOCK	38246G108	68	1,681	SH		OTR	1	1,357	120	204
GOODRX HOLDINGS INC	COMMON STOCK	38246G108	7	180	SH		DFND	5	180	0	0
GOOSEHEAD INSURANCE INC A	COMMON STOCK	38267D109	2,674	21,428	SH		DFND	1	21,328	0	100
GOOSEHEAD INSURANCE INC A	COMMON STOCK	38267D109	278	2,227	SH		OTR	1	2,227	0	0
GOOSEHEAD INSURANCE INC	COMMON STOCK	38267D109	13	102	SH		DFND	5	102	0	0
GOPRO INC CLASS A	COMMON STOCK	38268T103	3	368	SH		DFND	1	368	0	0
GOSSAMER BIO INC	COMMON STOCK	38341P102	17	1,796	SH		DFND	1	1,500	0	296
GRACE W R & CO	COMMON STOCK	38388F108	217	3,970	SH		DFND	1	610	0	3,360
GRACE W R & CO	COMMON STOCK	38388F108	6	113	SH		OTR	1	0	82	31
GRAND CANYON EDUCATION INC	COMMON STOCK	38526M106	3,137	33,688	SH		DFND	1	33,524	0	164
GRAND CANYON EDUCATION INC	COMMON STOCK	38526M106	244	2,627	SH		OTR	1	1,199	9	1,419
GRAND CANYON EDUCATION INC	COMMON STOCK	38526M106	298	3,201	SH		DFND	5	3,201	0	0
GRANITE POINT MORTGAGE TRUST	COMMON STOCK	38741L107	20	1,982	SH		DFND	1	1,270	0	712
GRAVITY CO LTD A D R	COMMON STOCK	38911N206	5	27	SH		DFND	1	19	0	8
GRAVITY CO LTD	COMMON STOCK	38911N206	0	1	SH		DFND	5	1	0	0
GREEN DOT CORP CL A	COMMON STOCK	39304D102	423	7,583	SH		DFND	1	7,583	0	0
GREENTREE HOSPITALTY GR A D R	COMMON STOCK	39579V100	9	689	SH		DFND	1	470	0	219
GROCERY OUTLET HOLDING CORP	COMMON STOCK	39874R101	990	25,232	SH		DFND	1	25,232	0	0
GROCERY OUTLET HOLDING CORP	COMMON STOCK	39874R101	68	1,734	SH		OTR	1	1,734	0	0
GROWGENERATION CORP	COMMON STOCK	39986L109	80	2,000	SH		DFND	1	2,000	0	0
GROWGENERATION CORP	COMMON STOCK	39986L109	80	2,000	SH		OTR	1	2,000	0	0
GRUPO TELEVISA SA SPON A D R	COMMON STOCK	40049J206	108	13,177	SH		DFND	1	12,438	0	739
GRUPO AVAL ACCIONES Y VALORE A	COMMON STOCK	40053W101	14	2,088	SH		DFND	1	564	0	1,524
GUARANTY FED BANCSHARES INC	COMMON STOCK	40108P101	2	89	SH		OTR	1	89	0	0
GUARDANT HEALTH INC	COMMON STOCK	40131M109	81	628	SH		DFND	1	569	0	59
GUARDANT HEALTH INC	COMMON STOCK	40131M109	149	1,150	SH		OTR	1	22	36	1,092
GUGGENHEIM STRATEGIC OPP FUND	OTHER	40167F101	12	600	SH		DFND	1	0	0	600
GUGGENHEIM CREDIT ALLOCATION	OTHER	40168G108	3	166	SH		DFND	1	166	0	0
GUIDEWIRE SOFTWARE INC	COMMON STOCK	40171V100	585	4,538	SH		DFND	1	4,460	0	78
GUIDEWIRE SOFTWARE INC	COMMON STOCK	40171V100	813	6,319	SH		OTR	1	108	5,134	1,077
GUIDEWIRE SOFTWARE INC	COMMON STOCK	40171V100	292	2,268	SH		DFND	5	2,268	0	0
HCA HEALTHCARE INC	COMMON STOCK	40412C101	1,389	8,443	SH		DFND	1	6,660	0	1,783
HCA HEALTHCARE INC	COMMON STOCK	40412C101	233	1,420	SH		OTR	1	910	68	442
HCA HOLDINGS INC	COMMON STOCK	40412C101	24	144	SH		DFND	5	144	0	0
HDFC BANK LTD A D R	COMMON STOCK	40415F101	5,860	81,097	SH		DFND	1	80,425	0	672
HDFC BANK LTD A D R	COMMON STOCK	40415F101	1,308	18,100	SH		OTR	1	16,843	1,149	108
HDFC BANK LTD	COMMON STOCK	40415F101	958	13,248	SH		DFND	5	13,248	0	0
HF FOODS GROUP INC	COMMON STOCK	40417F109	2	291	SH		DFND	1	0	0	291
HMS HLDGS CORP	COMMON STOCK	40425J101	84	2,276	SH		DFND	1	2,276	0	0
HMS HLDGS CORP	COMMON STOCK	40425J101	44	1,203	SH		OTR	1	1,203	0	0
HMS HOLDINGS CORP	COMMON STOCK	40425J101	5	123	SH		DFND	5	123	0	0
HP INC	COMMON STOCK	40434L105	4,761	193,603	SH		DFND	1	181,627	0	11,976
HP INC	COMMON STOCK	40434L105	3,390	137,826	SH		OTR	1	128,800	1,762	7,264
HP INC	COMMON STOCK	40434L105	4	170	SH		DFND	5	170	0	0
HALOZYME THERAPEUTICS INC	COMMON STOCK	40637H109	153	3,581	SH		DFND	1	2,332	0	1,249
HALOZYME THERAPEUTICS INC	COMMON STOCK	40637H109	281	6,594	SH		OTR	1	4,045	2,549	0
JOHN HANCOCK PREMIUM DIVIDEND	OTHER	41013T105	55	3,883	SH		OTR	1	3,508	0	375
JOHN HANCOCK PREMIUM DIVIDEND	OTHER	41013T105	100	7,023	SH		DFND	5	1,667	0	5,356
HANGER INC	COMMON STOCK	41043F208	11	500	SH		DFND	1	500	0	0
HANNON ARMSTRONG SUSTAINABLE	COMMON STOCK	41068X100	1,167	18,406	SH		DFND	1	13,941	0	4,465
HANNON ARMSTRONG SUSTAINABLE	COMMON STOCK	41068X100	152	2,399	SH		OTR	1	1,619	680	100
HARBORONE BANCORP INC	COMMON STOCK	41165Y100	1	126	SH		DFND	1	126	0	0
HARPOON THERAPEUTICS INC	COMMON STOCK	41358P106	28	1,705	SH		DFND	1	1,200	0	505
HEALTHSTREAM INC	COMMON STOCK	42222N103	4	164	SH		DFND	1	103	0	61
HEALTHCARE TRUST OF AME CL A	COMMON STOCK	42225P501	126	4,576	SH		DFND	1	4,233	0	343
HEALTHCARE TRUST OF AME CL A	COMMON STOCK	42225P501	44	1,590	SH		OTR	1	1,533	0	57
HEALTHCARE TRST OF AMERICA	COMMON STOCK	42225P501	2	65	SH		DFND	5	65	0	0
HEALTH CATALYST INC	COMMON STOCK	42225T107	360	8,260	SH		DFND	1	8,260	0	0
HEALTH CATALYST INC	COMMON STOCK	42225T107	61	1,403	SH		OTR	1	1,403	0	0
HEALTHEQUITY INC	COMMON STOCK	42226A107	1,734	24,877	SH		DFND	1	22,489	0	2,388
HEALTHEQUITY INC	COMMON STOCK	42226A107	40	567	SH		OTR	1	484	43	40
HEARTLAND FINANCIAL USA INC	COMMON STOCK	42234Q102	791	19,604	SH		DFND	1	19,586	0	18
HEALTHPEAK PROPERTIES INC	COMMON STOCK	42250P103	669	22,144	SH		DFND	1	16,397	0	5,747
HEALTHPEAK PROPERTIES INC	COMMON STOCK	42250P103	740	24,466	SH		OTR	1	22,150	800	1,516
HEALTHPEAK PROPERTIES INC	COMMON STOCK	42250P103	26	862	SH		DFND	5	862	0	0
HELIOS TECHNOLOGIES INC	COMMON STOCK	42328H109	51	950	SH		DFND	1	950	0	0
HELIOS TECHNOLOGIES INC	COMMON STOCK	42328H109	133	2,505	SH		OTR	1	130	0	2,375
HELIUS MEDICAL TECHNOLOGIES	COMMON STOCK	42328V504	0	500	SH		DFND	1	500	0	0
HELIX ENERGY SOLUTIONS GROUP I	COMMON STOCK	42330P107	20	4,756	SH		DFND	1	4,756	0	0
HELIX ENERGY SOLUTIONS GROUP I	COMMON STOCK	42330P107	49	11,675	SH		OTR	1	814	10,861	0
HERC HOLDINGS INC	COMMON STOCK	42704L104	48	719	SH		DFND	1	697	0	22
HERC HOLDINGS INC	COMMON STOCK	42704L104	1	20	SH		OTR	1	0	0	20
HERITAGE FINL CORP WASH	COMMON STOCK	42722X106	5	201	SH		DFND	1	0	0	201
HERITAGE FINL CORP WASH	COMMON STOCK	42722X106	16	705	SH		OTR	1	0	705	0
HESKA CORP	COMMON STOCK	42805E306	407	2,789	SH		DFND	1	2,789	0	0
HESKA CORP	COMMON STOCK	42805E306	62	423	SH		OTR	1	423	0	0
HESS CORP	COMMON STOCK	42809H107	647	12,240	SH		DFND	1	9,907	0	2,333
HESS CORP	COMMON STOCK	42809H107	471	8,921	SH		OTR	1	8,914	0	7
HESS CORP	COMMON STOCK	42809H107	1	27	SH		DFND	5	27	0	0
HEWLETT PACKARD ENTERPRIS CO	COMMON STOCK	42824C109	2,137	180,342	SH		DFND	1	174,692	0	5,650
HEWLETT PACKARD ENTERPRIS CO	COMMON STOCK	42824C109	1,435	121,050	SH		OTR	1	115,812	0	5,238
HIGHLAND INCOME FUND	OTHER	43010E404	86	8,319	SH		DFND	1	8,319	0	0
HILTON GRAND VACATIONS	COMMON STOCK	43283X105	136	4,353	SH		DFND	1	3,871	0	482
HILTON GRAND VACATIONS	COMMON STOCK	43283X105	1	43	SH		OTR	1	0	34	9
HIMAX TECHNOLOGIES INC A D R	COMMON STOCK	43289P106	13	1,760	SH		DFND	1	0	0	1,760
HIMAX TECHNOLOGIES INC	COMMON STOCK	43289P106	6	865	SH		DFND	5	865	0	0
HILTON WORLDWIDE HLDGS WI	COMMON STOCK	43300A203	856	7,695	SH		DFND	1	6,299	0	1,396
HILTON WORLDWIDE HLDGS WI	COMMON STOCK	43300A203	243	2,185	SH		OTR	1	1,682	18	485
HOMESTREET INC	COMMON STOCK	43785V102	1	44	SH		DFND	1	44	0	0
HOOKIPA PHARMA INC	COMMON STOCK	43906K100	8	750	SH		DFND	1	750	0	0
HOPE BANCORP INC	COMMON STOCK	43940T109	26	2,367	SH		DFND	1	2,166	0	201
HOST HOTELS RESORTS INC	COMMON STOCK	44107P104	627	42,889	SH		DFND	1	39,639	0	3,250
HOST HOTELS RESORTS INC	COMMON STOCK	44107P104	47	3,219	SH		OTR	1	2,689	530	0
HOST HOTELS & RESORTS INC	COMMON STOCK	44107P104	1	72	SH		DFND	5	72	0	0
HOSTESS BRANDS INC	COMMON STOCK	44109J106	605	41,345	SH		DFND	1	31,300	0	10,045
HOSTESS BRANDS INC	COMMON STOCK	44109J106	6	383	SH		OTR	1	383	0	0
HOWARD HUGHES CORP	COMMON STOCK	44267D107	97	1,230	SH		DFND	1	722	0	508
HOWARD HUGHES CORP	COMMON STOCK	44267D107	9	110	SH		OTR	1	17	59	34
HUAMI CORP ADR A D R	COMMON STOCK	44331K103	1	80	SH		DFND	1	0	0	80
HUAZHU GROUP LTD A D R	COMMON STOCK	44332N106	298	6,618	SH		DFND	1	5,557	0	1,061
HUTCHISON CHINA MEDITECH LTD A	COMMON STOCK	44842L103	42	1,305	SH		DFND	1	1,216	0	89
HUYA INC ADS	COMMON STOCK	44852D108	69	3,466	SH		DFND	1	2,831	0	635
HYCROFT MNG HLDG CORP WT 9	OTHER	44862P125	0	7	SH		DFND	1	7	0	0
IAC INTERACTIVECORP NEW	COMMON STOCK	44891N109	1,163	6,143	SH		DFND	1	5,640	0	503
IAC INTERACTIVECORP NEW	COMMON STOCK	44891N109	242	1,277	SH		OTR	1	1,229	42	6
IAC/INTERACTIVECORP	COMMON STOCK	44891N109	235	1,240	SH		DFND	5	1,240	0	0
HYRECAR INC	COMMON STOCK	44916T107	4	500	SH		DFND	1	500	0	0
ICF INTERNATIONAL INC	COMMON STOCK	44925C103	300	4,044	SH		DFND	1	4,005	0	39
ICF INTERNATIONAL INC	COMMON STOCK	44925C103	129	1,733	SH		OTR	1	1,234	499	0
ICF INTERNATIONAL INC	COMMON STOCK	44925C103	10	139	SH		DFND	5	139	0	0
ICU MEDICAL INC	COMMON STOCK	44930G107	738	3,439	SH		DFND	1	3,192	0	247
ICU MEDICAL INC	COMMON STOCK	44930G107	71	330	SH		OTR	1	112	180	38
IPG PHOTONICS CORP	COMMON STOCK	44980X109	2,833	12,661	SH		DFND	1	10,989	0	1,672
IPG PHOTONICS CORP	COMMON STOCK	44980X109	127	569	SH		OTR	1	569	0	0
IPG PHOTONICS CORP	COMMON STOCK	44980X109	9	41	SH		DFND	5	41	0	0
INVO BIOSCIENCE INC	COMMON STOCK	44984F302	32	10,625	SH		OTR	1	10,625	0	0
ISTAR INC	COMMON STOCK	45031U101	21	1,443	SH		DFND	1	734	0	709
ISTAR INC	COMMON STOCK	45031U101	1	75	SH		OTR	1	0	0	75
ITT CORP NEW	COMMON STOCK	45073V108	1,895	24,610	SH		DFND	1	19,798	0	4,812
ITT CORP NEW	COMMON STOCK	45073V108	409	5,304	SH		OTR	1	4,107	805	392
ICICI BANK LTD A D R	COMMON STOCK	45104G104	5,108	343,742	SH		DFND	1	320,800	0	22,942
ICICI BANK LTD A D R	COMMON STOCK	45104G104	791	53,255	SH		OTR	1	51,216	1,891	148
ICICI BANK LTD	COMMON STOCK	45104G104	25	1,699	SH		DFND	5	1,699	0	0
ICLICK INTERACTIVE ASIA ADR	COMMON STOCK	45113Y104	1	125	SH		DFND	1	0	0	125
IDEX CORP	COMMON STOCK	45167R104	8,913	44,740	SH		DFND	1	41,962	0	2,778
IDEX CORP	COMMON STOCK	45167R104	963	4,836	SH		OTR	1	4,315	440	81
IDEX CORP	COMMON STOCK	45167R104	50	252	SH		DFND	5	241	0	11
IDEXX LABS INC	COMMON STOCK	45168D104	18,558	37,127	SH		DFND	1	35,639	0	1,488
IDEXX LABS INC	COMMON STOCK	45168D104	2,998	5,999	SH		OTR	1	2,884	93	3,022
IDEXX LABS CORP	COMMON STOCK	45168D104	614	1,229	SH		DFND	5	1,229	0	0
IKONICS CORP	COMMON STOCK	45172K102	30	3,000	SH		DFND	1	3,000	0	0
IHEARTMEDIA INC	COMMON STOCK	45174J509	1	100	SH		DFND	1	100	0	0
IMMUNOGEN INC	COMMON STOCK	45253H101	3	473	SH		DFND	1	0	0	473
INARI MED INC	COMMON STOCK	45332Y109	118	1,354	SH		OTR	1	0	0	1,354
INCYTE CORP	COMMON STOCK	45337C102	2,285	26,262	SH		DFND	1	25,154	0	1,108
INCYTE CORP	COMMON STOCK	45337C102	3,029	34,815	SH		OTR	1	34,023	83	709
INCYTE CORP	COMMON STOCK	45337C102	2	23	SH		DFND	5	23	0	0
INDEPENDENCE REALTY TRUST IN	COMMON STOCK	45378A106	3	214	SH		DFND	1	214	0	0
INDEPENDENCE REALTY TRUST IN	COMMON STOCK	45378A106	81	6,040	SH		OTR	1	0	6,040	0
INDEPENDENT BANK GROUP INC	COMMON STOCK	45384B106	476	7,609	SH		DFND	1	7,580	0	29
INDEPENDENT BANK GROUP INC	COMMON STOCK	45384B106	110	1,753	SH		OTR	1	1,522	231	0
INDEXIQ ETF TR IQ MRGR ARB ETF	OTHER	45409B800	20	602	SH		DFND	1	0	0	602
IQ ARB MERGER ARBITAGE	OTHER	45409B800	51	1,527	SH		DFND	5	1,527	0	0
INFINERA CORP	COMMON STOCK	45667G103	17	1,578	SH		DFND	1	1,578	0	0
INGERSOLL RAND INC	COMMON STOCK	45687V106	2,438	53,522	SH		DFND	1	50,344	0	3,178
INGERSOLL RAND INC	COMMON STOCK	45687V106	559	12,276	SH		OTR	1	11,214	630	432
INGERSOLL RAND INC	COMMON STOCK	45687V106	67	1,475	SH		DFND	5	1,475	0	0
INGEVITY CORP	COMMON STOCK	45688C107	260	3,428	SH		DFND	1	1,737	0	1,691
INGEVITY CORP	COMMON STOCK	45688C107	96	1,259	SH		OTR	1	1,028	145	86
INSIGHT ENTERPRISES INC	COMMON STOCK	45765U103	9	117	SH		DFND	1	117	0	0
INNOSPEC INC	COMMON STOCK	45768S105	86	944	SH		DFND	1	54	0	890
INNOSPEC INC	COMMON STOCK	45768S105	69	755	SH		OTR	1	0	755	0
INPHI CORP	COMMON STOCK	45772F107	356	2,219	SH		DFND	1	1,177	0	1,042
INPHI CORP	COMMON STOCK	45772F107	14	89	SH		OTR	1	40	49	0
INOVIO PHARMACEUTICALS INC	COMMON STOCK	45773H201	62	7,036	SH		DFND	1	7,036	0	0
INSTEEL INDUSTRIES INC	COMMON STOCK	45774W108	2	92	SH		DFND	1	92	0	0
INSPERITY INC	COMMON STOCK	45778Q107	9	114	SH		DFND	1	114	0	0
INSPERITY INC	COMMON STOCK	45778Q107	231	2,841	SH		OTR	1	0	0	2,841
INOGEN INC	COMMON STOCK	45780L104	130	2,895	SH		DFND	1	2,895	0	0
INOGEN INC	COMMON STOCK	45780L104	17	382	SH		OTR	1	382	0	0
INSTALLED BUILDING PRODUCTS	COMMON STOCK	45780R101	528	5,182	SH		DFND	1	5,166	0	16
INSTALLED BUILDING PRODUCTS	COMMON STOCK	45780R101	28	274	SH		OTR	1	77	186	11
INOVALON HOLDINGS INC	COMMON STOCK	45781D101	249	13,713	SH		DFND	1	12,786	0	927
INOVALON HOLDINGS INC	COMMON STOCK	45781D101	40	2,204	SH		OTR	1	2,204	0	0
INNOVATIVE INDUSTRIAL PROPER	COMMON STOCK	45781V101	113	617	SH		DFND	1	542	0	75
INNOVATIVE INDUSTRIAL PPTYS	COMMON STOCK	45781V101	14	77	SH		DFND	5	77	0	0
INSIGHT SELECT INCOME FUND	OTHER	45781W109	64	3,000	SH		OTR	1	3,000	0	0
INSEEGO CORP	COMMON STOCK	45782B104	72	4,685	SH		DFND	1	4,685	0	0
INNOVATOR IBD 50	OTHER	45782C102	7	170	SH		DFND	5	170	0	0
INSULET CORP	COMMON STOCK	45784P101	2,122	8,300	SH		DFND	1	8,098	0	202
INSULET CORP	COMMON STOCK	45784P101	1,043	4,077	SH		OTR	1	3,707	323	47
INSULET CORP	COMMON STOCK	45784P101	38	148	SH		DFND	5	148	0	0
INTELLIGENT SYSTEMS CORP	COMMON STOCK	45816D100	20	500	SH		DFND	1	500	0	0
INTEGER HOLDINGS CORPORATION	COMMON STOCK	45826H109	30	365	SH		DFND	1	365	0	0
INTEGER HOLDINGS CORPORATION	COMMON STOCK	45826H109	1	11	SH		OTR	1	11	0	0
INTEGER HOLDINGS CORP	COMMON STOCK	45826H109	34	413	SH		DFND	5	413	0	0
INTELLIA THERAPEUTICS INC	COMMON STOCK	45826J105	27	500	SH		DFND	1	500	0	0
INTELLIA THERAPEUTICS INC	COMMON STOCK	45826J105	10	175	SH		OTR	1	175	0	0
INTERACTIVE BROKERS GROUP CL A	COMMON STOCK	45841N107	2,286	37,516	SH		DFND	1	37,452	0	64
INTERACTIVE BROKERS GROUP CL A	COMMON STOCK	45841N107	310	5,086	SH		OTR	1	5,086	0	0
INTERCEPT PHARMACEUTICALS IN	COMMON STOCK	45845P108	5	214	SH		DFND	1	68	0	146
INTERCONTINENTAL HOTELS A D R	COMMON STOCK	45857P806	391	6,013	SH		DFND	1	3,343	0	2,670
INTERCONTINENTAL HOTELS A D R	COMMON STOCK	45857P806	86	1,317	SH		OTR	1	790	478	49
INTERCONTINENTAL EXCHANGE IN	COMMON STOCK	45866F104	7,249	62,876	SH		DFND	1	56,864	0	6,012
INTERCONTINENTAL EXCHANGE IN	COMMON STOCK	45866F104	4,736	41,075	SH		OTR	1	35,158	2,670	3,247
INTERCONTINENTAL EXCHANGE INC	COMMON STOCK	45866F104	407	3,529	SH		DFND	5	3,529	0	0
INTERDIGITAL INC	COMMON STOCK	45867G101	337	5,568	SH		DFND	1	5,518	0	50
INTERDIGITAL INC	COMMON STOCK	45867G101	50	837	SH		OTR	1	837	0	0
INTERSECT ENT INC	COMMON STOCK	46071F103	13	572	SH		DFND	1	572	0	0
INTERSECT ENT INC	COMMON STOCK	46071F103	4	175	SH		OTR	1	0	175	0
INVESCO S P 500 DOWNSIDE HEDGE	OTHER	46090A705	17	500	SH		DFND	1	500	0	0
INVESCO TOTAL RETURN BOND ETF	OTHER	46090A804	17	296	SH		DFND	1	296	0	0
INVESCO BLDRS EMERGING MKTS	OTHER	46090C305	0	0	SH		DFND	5	0	0	0
INVESCO OPTIMUM YIELD ETF	OTHER	46090F100	112	7,321	SH		DFND	1	7,321	0	0
INVESCO OPTIMUM YIELD	OTHER	46090F100	2,075	135,968	SH		DFND	5	0	0	135,968
INTRA CELLULAR THERAPIES INC	COMMON STOCK	46116X101	8	266	SH		DFND	1	148	0	118
INTUITIVE SURGICAL INC	COMMON STOCK	46120E602	13,797	16,864	SH		DFND	1	16,190	0	674
INTUITIVE SURGICAL INC	COMMON STOCK	46120E602	4,992	6,102	SH		OTR	1	3,322	1,124	1,656
INTUITIVE SURGICAL INC	COMMON STOCK	46120E602	290	354	SH		DFND	5	333	0	21
INTRICON CORP	COMMON STOCK	46121H109	632	34,894	SH		DFND	1	34,894	0	0
INTRICON CORP	COMMON STOCK	46121H109	12	667	SH		OTR	1	667	0	0
INTREPID POTASH INC	COMMON STOCK	46121Y201	1	60	SH		DFND	1	60	0	0
INVESCO MORTGAGE CAPITAL	COMMON STOCK	46131B100	2	667	SH		DFND	1	667	0	0
INVESCO MORTGAGE CAPITAL	COMMON STOCK	46131B100	4	1,054	SH		OTR	1	0	0	1,054
INVESCO VAN KAMPEN SENIOR INCO	OTHER	46131H107	102	25,714	SH		DFND	1	24,914	0	800
INVESCO VAN KAMPEN MUNICIPAL T	OTHER	46131J103	8	612	SH		DFND	1	612	0	0
INVESCO MUNICIPAL OPPORTUNITY	OTHER	46132C107	20	1,582	SH		DFND	1	1,582	0	0
INVESCO MUNICIPAL OPPORTUNITY	OTHER	46132C107	13	1,018	SH		OTR	1	1,018	0	0
INVESCO VAN KAMPEN CA VALUE	OTHER	46132H106	19	1,454	SH		DFND	1	1,454	0	0
INVESCO VALUE MUNICIPAL INCOME	OTHER	46132P108	205	13,097	SH		OTR	1	13,097	0	0
INVESCO QUALITY MUNI INC TR	OTHER	46133G107	52	4,019	SH		DFND	1	4,019	0	0
INVESCO HIGH INCOME 2023 TAR	OTHER	46135X108	9	1,060	SH		DFND	1	1,060	0	0
INVESTCO INDIA EXCHANGE TRADED	OTHER	46137R109	51	2,200	SH		DFND	1	2,200	0	0
INVESCO EXCHANGE TRADED FD TR	OTHER	46137V100	18	268	SH		DFND	1	268	0	0
INVESTCO EXCHANGE TRADED FD TR	OTHER	46137V134	55	536	SH		DFND	1	536	0	0
INVESCO EXCHANGE TRADED FD ETF	OTHER	46137V142	615	13,250	SH		DFND	1	12,950	0	300
INVESCO EXCHANGE TRADED FD ETF	OTHER	46137V142	47	1,015	SH		OTR	1	815	200	0
INVESCO S P MIDCAP 400 PURE ET	OTHER	46137V191	58	790	SH		DFND	1	790	0	0
INVESCO S P MIDCAP 400 PURE ET	OTHER	46137V217	152	750	SH		DFND	1	750	0	0
INVESCO S P 500 TOP 50 ETF	OTHER	46137V233	427	1,497	SH		DFND	1	1,497	0	0
INVESCO POWERSHARES S&P 500	OTHER	46137V233	2,099	7,359	SH		DFND	5	0	0	7,359
INVESCO EXCHANGE TRADED FD TR	OTHER	46137V241	71	1,681	SH		DFND	1	1,681	0	0
INVESCO EXCHANGE TRADED FD TR	OTHER	46137V241	19	447	SH		OTR	1	0	0	447
INVESCO POWERSHARES S&P 500	OTHER	46137V241	2,900	68,858	SH		DFND	5	0	0	68,858
INVESCO S P 500 PURE VALUE ETF	OTHER	46137V258	64	1,037	SH		DFND	1	1,037	0	0
INVESCO S P 500 PURE GROWTH ET	OTHER	46137V266	1,069	6,562	SH		DFND	1	6,562	0	0
INVESCO POWERSHARES S&P EQUAL	OTHER	46137V274	21	213	SH		DFND	5	213	0	0
INVESCO S P 500 EQUAL WEIGHT E	OTHER	46137V282	70	277	SH		DFND	1	277	0	0
INVESCO S P 500 EQUAL WEIGHT E	OTHER	46137V282	48	190	SH		OTR	1	190	0	0
INVESCO EXCHANGE TRADED FD TR	OTHER	46137V308	255	3,475	SH		DFND	1	3,475	0	0
INVESCO BUYBACK ACHIEVERS	OTHER	46137V308	263	3,586	SH		DFND	5	3,181	0	405
INVESCO S P 500 EQUAL WEIGHT E	OTHER	46137V316	37	264	SH		DFND	1	264	0	0
INVESCO S P 500 EQUAL WEIGHT E	OTHER	46137V324	8	53	SH		DFND	1	53	0	0
INVESCO S P 500 EQUAL WEIGHT E	OTHER	46137V332	24	92	SH		DFND	1	92	0	0
INVESCO S P 500 EQUAL WEIGHT E	OTHER	46137V357	1,240	9,726	SH		DFND	1	9,226	0	500
INVESCO S P 500 EQUAL WEIGHT E	OTHER	46137V357	1	10	SH		OTR	1	10	0	0
INVESCO S&P 500 EQUAL WEIGHT	OTHER	46137V357	88	693	SH		DFND	5	0	0	693
INVESCO S P 500 EQUAL WEIGHT E	OTHER	46137V365	4	125	SH		DFND	1	125	0	0
INVESCO EXCHANGE TRADED FD TR	OTHER	46137V407	1,109	14,755	SH		DFND	1	1,005	0	13,750
INVESCO RUSSELL MIDCAP PURE	OTHER	46137V464	55	699	SH		DFND	5	369	0	330
INVESCO SP SMALLCAP MOMENTUM E	OTHER	46137V498	14	300	SH		DFND	1	300	0	0
INVESTCO EXCHANGE TRADED FD TR	OTHER	46137V506	12	375	SH		DFND	1	375	0	0
INVESCO EXCHANGE TRADED FD TR	OTHER	46137V530	276	1,210	SH		DFND	1	1,210	0	0
INVESCO EXCHANGE TRADED FD TR	OTHER	46137V548	43	2,800	SH		OTR	1	2,800	0	0
INVESCO POWERSHARES INTL	OTHER	46137V548	1	92	SH		DFND	5	0	0	92
INVESCO EXCHANGE TRADED FD TR	OTHER	46137V563	84	4,819	SH		DFND	1	4,819	0	0
INVESCO FTSE RAFI US SML MID E	OTHER	46137V597	2,762	18,301	SH		DFND	1	14,801	0	3,500
POWERSHARES FTSE RAFI US 1000	OTHER	46137V613	2,365	17,753	SH		DFND	1	17,753	0	0
POWERSHARES FTSE RAFI US 1000	OTHER	46137V613	220	1,655	SH		OTR	1	1,655	0	0
INVESCO POWERSHARES FTSE RAFI	OTHER	46137V613	10	72	SH		DFND	5	0	0	72
INVESCO EXCHANGE TRADED FD TR	OTHER	46137V621	54	2,800	SH		DFND	1	2,800	0	0
INVESCO EXCHANGE TRADED FD TR	OTHER	46137V621	165	8,600	SH		OTR	1	0	0	8,600
INVESCO EXCHANGE TRADED FD TR	OTHER	46137V647	24	230	SH		DFND	1	230	0	0
INVESCO EXCHANGE TRADED FD TR	OTHER	46137V662	118	1,650	SH		DFND	1	1,650	0	0
INVESCO DYNAMIC LEISURE AND ET	OTHER	46137V720	168	4,200	SH		DFND	1	4,200	0	0
INVESCO EXCHANGE TRADED FD TR	OTHER	46137V738	99	2,572	SH		DFND	1	2,572	0	0
INVESCO POWERSHARES DYNAMIC	OTHER	46137V753	21	588	SH		DFND	5	588	0	0
INVESCO DYNAMIC BUILDING CONST	OTHER	46137V779	1	18	SH		DFND	1	18	0	0
INVESCO EXCHANGE TRADED FD TR	OTHER	46137V787	1,190	16,965	SH		DFND	1	16,500	0	465
INVESCO EXCHANGE TRADED FD TR	OTHER	46137V837	67	761	SH		DFND	1	761	0	0
INVESCO DWA MOMENTUM	OTHER	46137V837	24	268	SH		DFND	5	0	0	268
INVESTCO DB COMMODITY INDEX ET	OTHER	46138B103	27	1,865	SH		DFND	1	1,865	0	0
INVESCO DB COMMODITY INDEX	OTHER	46138B103	78	5,284	SH		DFND	5	12	0	5,272
INVESCO S P 500 LOW VOLATILITY	OTHER	46138E354	717	12,753	SH		DFND	1	9,498	0	3,255
INVESCO POWERSHARES S&P 500	OTHER	46138E354	454	8,077	SH		DFND	5	477	0	7,600
INVESCO S P 500 HIGH DIVIDEND	OTHER	46138E362	28	743	SH		DFND	1	743	0	0
INVESCO POWERSHARES S&P 500	OTHER	46138E362	95	2,526	SH		DFND	5	0	0	2,526
INVESCO S P 500 ENHANCED VALUE	OTHER	46138E396	7	194	SH		DFND	1	194	0	0
INVESCO CEF INCOME COMPOSITE E	OTHER	46138E404	18	800	SH		DFND	1	800	0	0
INVESCO CEF INCOME COMPOSITE E	OTHER	46138E404	121	5,400	SH		OTR	1	0	0	5,400
INVESCO PREFERRED ETF	OTHER	46138E511	212	13,888	SH		DFND	1	12,538	0	1,350
INVESCO POWERSHARES	OTHER	46138E511	318	20,824	SH		DFND	5	20,824	0	0
INVESCO NATIONAL AMT FREE ETF	OTHER	46138E537	47	1,737	SH		DFND	1	1,737	0	0
INVESCO NATIONAL AMT FREE ETF	OTHER	46138E537	22	800	SH		OTR	1	0	0	800
INVESCO POWERSHARES NATIONAL	OTHER	46138E537	397	14,631	SH		DFND	5	14,631	0	0
INVESCO INTERNATIONAL	OTHER	46138E636	11	380	SH		DFND	5	380	0	0
INVESCO INTERNATIONAL BUYBACK	OTHER	46138E644	47	1,236	SH		DFND	1	1,236	0	0
INVESCO GLOBAL WATER ETF	OTHER	46138E651	31	900	SH		DFND	1	0	0	900
INVESCO GLOBAL WATER ETF	OTHER	46138E651	39	1,113	SH		OTR	1	0	0	1,113
INVESCO FTSE RAFI EMERGING ETF	OTHER	46138E727	2	118	SH		DFND	1	118	0	0
INVESCO FTSE RAFI DEVELOPED ET	OTHER	46138E743	8,521	200,778	SH		DFND	1	195,896	0	4,882
INVESCO FTSE RAFI DEVELOPED ET	OTHER	46138E743	133	3,142	SH		OTR	1	2,937	205	0
INVESCO POWERSHARES FTSE RAFI	OTHER	46138E743	46	1,081	SH		DFND	5	0	0	1,081
INVESCO EMERGING MARKETS ETF	OTHER	46138E784	970	33,632	SH		DFND	1	33,632	0	0
INVESCO EMERGING MARKETS ETF	OTHER	46138E784	531	18,418	SH		OTR	1	18,418	0	0
INVESCO EMERGING MARKETS	OTHER	46138E784	227	7,862	SH		DFND	5	0	0	7,862
INVESCO S P SMALLCAP LOW ETF	OTHER	46138G102	15	362	SH		DFND	1	362	0	0
INVESCO POWERSHARES S&P	OTHER	46138G102	8	200	SH		DFND	5	0	0	200
INVESCO POWERSHARES SENIOR	OTHER	46138G508	2	102	SH		DFND	5	0	0	102
INVESCO FUNDS	OTHER	46138G664	370	4,568	SH		DFND	5	0	0	4,568
INVESCO FUNDS	OTHER	46138G672	55	770	SH		DFND	5	0	0	770
INVESCO S P 500 REVENUE ETF	OTHER	46138G698	46	731	SH		DFND	1	731	0	0
INVESCO EXCHANGE TRADED FD TR	OTHER	46138G706	476	4,629	SH		DFND	1	4,629	0	0
INVESCO EXCHANGE TRADED FD TR	OTHER	46138G706	26	250	SH		OTR	1	250	0	0
INVESCO TAXABLE MUNICIPAL ETF	OTHER	46138G805	137	4,100	SH		DFND	1	4,100	0	0
INVESCO TAXABLE MUNICIPAL ETF	OTHER	46138G805	23	674	SH		OTR	1	0	0	674
INVESCO GLOBAL CLEAN ENERGY ET	OTHER	46138G847	306	8,797	SH		DFND	1	8,797	0	0
INVESCO BULLETSHARES 2021 ETF	OTHER	46138J700	163	7,697	SH		DFND	1	1,822	0	5,875
INVESCO DEFENSIVE EQUITY ETF	OTHER	46138J775	91	1,544	SH		DFND	1	1,544	0	0
INVESCO POWERSHARES DEFENSIVE	OTHER	46138J775	3,534	59,742	SH		DFND	5	0	0	59,742
INVESCO BULLETSHARES 2021 HIGH	OTHER	46138J809	14	601	SH		DFND	1	601	0	0
INVESCO BULLETSHARES 2024 ETF	OTHER	46138J841	139	6,200	SH		DFND	1	0	0	6,200
INVESCO BULLETSHARES 2023 HIGH	OTHER	46138J858	14	570	SH		DFND	1	570	0	0
INVESCO BULLETSHARES 2023 ETF	OTHER	46138J866	135	6,150	SH		DFND	1	0	0	6,150
INVESCO BULLETSHARES 2023 ETF	OTHER	46138J866	11	500	SH		OTR	1	0	0	500
INVESCO BULLETSHARES 2022 HIGH	OTHER	46138J874	14	606	SH		DFND	1	606	0	0
INVESCO BULLETSHARES 2022 ETF	OTHER	46138J882	183	8,421	SH		DFND	1	2,596	0	5,825
INVESCO CURRENCYSHARES SWISS E	OTHER	46138R108	103	1,000	SH		DFND	1	1,000	0	0
INVESCO POWERSHARES DB	OTHER	46140H106	15	899	SH		DFND	5	0	0	899
INVESCO POWERSHARES DB	OTHER	46140H502	148	2,809	SH		DFND	5	2,809	0	0
INVESTORS BANCORP INC	COMMON STOCK	46146L101	7	690	SH		DFND	1	690	0	0
INVITAE CORP	COMMON STOCK	46185L103	1,355	32,411	SH		DFND	1	32,136	0	275
INVITAE CORP	COMMON STOCK	46185L103	42	1,000	SH		OTR	1	960	19	21
INVITATION HOMES INC	COMMON STOCK	46187W107	1,483	49,946	SH		DFND	1	48,496	0	1,450
INVITATION HOMES INC	COMMON STOCK	46187W107	429	14,449	SH		OTR	1	14,449	0	0
INVITATION HOMES INC	COMMON STOCK	46187W107	5	171	SH		DFND	5	171	0	0
IQVIA HOLDINGS INC	COMMON STOCK	46266C105	2,181	12,173	SH		DFND	1	9,780	0	2,393
IQVIA HOLDINGS INC	COMMON STOCK	46266C105	609	3,399	SH		OTR	1	1,879	1,444	76
IQVIA HOLDINGS INC	COMMON STOCK	46266C105	381	2,128	SH		DFND	5	2,128	0	0
IQIYI INC A D R	COMMON STOCK	46267X108	226	12,939	SH		DFND	1	10,980	0	1,959
IQIYI INC A D R	COMMON STOCK	46267X108	22	1,295	SH		OTR	1	1,262	0	33
IRIDIUM COMMUNICATION INC	COMMON STOCK	46269C102	368	9,349	SH		DFND	1	9,153	0	196
IRIDIUM COMMUNICATION INC	COMMON STOCK	46269C102	16	402	SH		OTR	1	402	0	0
IRON MOUNTAIN INC	COMMON STOCK	46284V101	198	6,730	SH		DFND	1	3,022	0	3,708
IRON MOUNTAIN INC	COMMON STOCK	46284V101	454	15,399	SH		OTR	1	14,711	643	45
IRON MOUNTAIN	COMMON STOCK	46284V101	2	57	SH		DFND	5	57	0	0
IRONWOOD PHARMACEUTICALS INC	COMMON STOCK	46333X108	113	9,895	SH		DFND	1	9,895	0	0
IRONWOOD PHARMACEUTICALS INC	COMMON STOCK	46333X108	3	262	SH		OTR	1	262	0	0
ISHARES SILVER TRUST ETF	OTHER	46428Q109	1,173	47,756	SH		DFND	1	47,756	0	0
ISHARES SILVER TRUST ETF	OTHER	46428Q109	64	2,609	SH		OTR	1	2,609	0	0
ISHARES SILVER TRUST	OTHER	46428Q109	99	4,047	SH		DFND	5	1,207	0	2,840
ISHARES US TREASURY BOND ETF	OTHER	46429B267	54,300	1,993,364	SH		DFND	1	1,964,092	0	29,272
ISHARES US TREASURY BOND ETF	OTHER	46429B267	6,298	231,197	SH		OTR	1	229,157	2,040	0
ISHARES AAA A RATED CORPORATE	OTHER	46429B291	88	1,502	SH		DFND	1	1,502	0	0
ISHARES CMBS ETF	OTHER	46429B366	221	4,000	SH		DFND	1	4,000	0	0
ISHARES CMBS ETF	OTHER	46429B366	122	2,200	SH		OTR	1	2,200	0	0
ISHARES MSCI INDIA ETF	OTHER	46429B598	30	750	SH		DFND	1	750	0	0
ISHARES FLOATING RATE BOND ETF	OTHER	46429B655	25,623	505,184	SH		DFND	1	503,989	0	1,195
ISHARES FLOATING RATE BOND ETF	OTHER	46429B655	1,868	36,832	SH		OTR	1	36,617	0	215
ISHARES CORE HIGH DIVIDEND ETF	OTHER	46429B663	99	1,131	SH		DFND	1	1,131	0	0
ISHARES CORE HIGH DIVIDEND ETF	OTHER	46429B663	43	490	SH		OTR	1	490	0	0
ISHARES HIGH DIVIDEND EQTY FD	OTHER	46429B663	37	425	SH		DFND	5	425	0	0
ISHARES MSCI CHINA ETF	OTHER	46429B671	20	252	SH		DFND	1	252	0	0
ISHARES MSCI EAFE MIN VOL FACT	OTHER	46429B689	341	4,639	SH		DFND	1	4,639	0	0
ISHARES MSCI EAFE MIN VOL FACT	OTHER	46429B689	146	1,985	SH		OTR	1	0	0	1,985
ISHARES MSCI EAFE MINIUM	OTHER	46429B689	640	8,717	SH		DFND	5	505	0	8,212
ISHARES MSCI USA MIN VOL FACTO	OTHER	46429B697	200,123	2,948,192	SH		DFND	1	2,929,238	0	18,954
ISHARES MSCI USA MIN VOL FACTO	OTHER	46429B697	21,663	319,129	SH		OTR	1	304,524	10,828	3,777
ISHARES MSCI USA MINIMUM	OTHER	46429B697	386	5,691	SH		DFND	5	1,339	0	4,352
ISHARES 0 5 YEAR TIPS BOND ETF	OTHER	46429B747	2,937	28,123	SH		DFND	1	28,123	0	0
ISHARES BARCLAYS 0-5 YEAR TIPS	OTHER	46429B747	110	1,054	SH		DFND	5	0	0	1,054
BLACKROCK SHORT MATURITY BOND	OTHER	46431W507	65	1,288	SH		DFND	1	1,288	0	0
BLACKROCK SHORT MATURITY BOND	OTHER	46431W507	515	10,270	SH		OTR	1	10,270	0	0
ISHARES INT HEDG CORP BD	OTHER	46431W705	25	261	SH		DFND	5	261	0	0
ISHARES MSCI USA QUALITY FACTO	OTHER	46432F339	61,196	526,597	SH		DFND	1	523,659	0	2,938
ISHARES MSCI USA QUALITY FACTO	OTHER	46432F339	4,133	35,561	SH		OTR	1	34,521	1,040	0
ISHARES MSCI USA QUALITY	OTHER	46432F339	141	1,215	SH		DFND	5	432	0	783
ISHARES MSCI USA SIZE FACTOR E	OTHER	46432F370	150	1,352	SH		DFND	1	1,352	0	0
ISHARES MSCI USA SIZE FACTOR E	OTHER	46432F370	10	89	SH		OTR	1	89	0	0
ISHARES MSCI USA VALUE FACTOR	OTHER	46432F388	7,923	91,168	SH		DFND	1	89,924	0	1,244
ISHARES MSCI USA VALUE FACTOR	OTHER	46432F388	543	6,244	SH		OTR	1	5,451	793	0
ISHARES MSCI USA VALUE FACTR	OTHER	46432F388	72	823	SH		DFND	5	0	0	823
ISHARES MSCI USA MOMENTUM FACT	OTHER	46432F396	12,744	79,016	SH		DFND	1	78,226	0	790
ISHARES MSCI USA MOMENTUM FACT	OTHER	46432F396	1,183	7,336	SH		OTR	1	6,835	501	0
ISHARES CORE MSCI TOTAL ETF	OTHER	46432F834	45,824	682,002	SH		DFND	1	675,832	0	6,170
ISHARES CORE MSCI TOTAL ETF	OTHER	46432F834	7,848	116,799	SH		OTR	1	116,799	0	0
ISHARES CORE MSCI EAFE ETF	OTHER	46432F842	786,115	11,378,124	SH		DFND	1	11,300,078	0	78,046
ISHARES CORE MSCI EAFE ETF	OTHER	46432F842	112,468	1,627,841	SH		OTR	1	1,589,126	20,290	18,425
ISHARES CORE MSCI EAFE	OTHER	46432F842	181,210	2,622,756	SH		DFND	5	36,751	0	2,586,005
ISHARES CORE 1 5 YEAR USD BOND	OTHER	46432F859	72	1,403	SH		DFND	1	1,403	0	0
ISHARES IBONDS MAR 2023 TERM C	OTHER	46432FAZ0	5,171	189,597	SH		DFND	1	184,597	0	5,000
ISHARES IBONDS MAR 2023 TERM C	OTHER	46432FAZ0	227	8,320	SH		OTR	1	8,320	0	0
ISHARES CORE MSCI EMERGING MKT	OTHER	46434G103	616,329	9,934,367	SH		DFND	1	9,863,683	0	70,684
ISHARES CORE MSCI EMERGING MKT	OTHER	46434G103	90,732	1,462,485	SH		OTR	1	1,441,811	9,826	10,848
ISHARES CORE MSCI EMERGING MKT	OTHER	46434G103	149,674	2,412,468	SH		DFND	5	23,821	0	2,388,647
ISHARES MSCI TAIWAN ETF	OTHER	46434G772	106	2,000	SH		DFND	1	2,000	0	0
ISHARES MSCI TAIWAN ETF	OTHER	46434G772	48	913	SH		OTR	1	913	0	0
ISHARES MSCI SINGAPORE ETF	OTHER	46434G780	240	11,173	SH		DFND	1	11,173	0	0
ISHARES MSCI SINGAPORE ETF	OTHER	46434G780	35	1,625	SH		OTR	1	1,625	0	0
ISHARES MSCI JAPAN ETF	OTHER	46434G822	1,428	21,142	SH		DFND	1	21,142	0	0
ISHARES MSCI JAPAN ETF	OTHER	46434G822	1,803	26,686	SH		OTR	1	23,686	3,000	0
ISHARES ESG AWARE MSCI EM ETF	OTHER	46434G863	6,039	143,857	SH		DFND	1	138,473	0	5,384
ISHARES ESG AWARE MSCI EM ETF	OTHER	46434G863	1,244	29,633	SH		OTR	1	29,633	0	0
ISHARES MSCI EM ESG SELECT	OTHER	46434G863	28	673	SH		DFND	5	673	0	0
ISHARES 0 5 YEAR INVESTMENT ET	OTHER	46434V100	125	2,388	SH		DFND	1	2,388	0	0
ISHARES MSCI INTL MULTIFACTOR	OTHER	46434V274	60	2,225	SH		DFND	1	2,225	0	0
ISHARES EXPONENTIAL TECHNOLOGI	OTHER	46434V381	20	350	SH		OTR	1	0	0	350
ISHARES 0 5 YEAR HIGH YIELD ET	OTHER	46434V407	55	1,205	SH		DFND	1	1,205	0	0
ISHARES 0-5 YEAR HI YLD CRP BD	OTHER	46434V407	37	817	SH		DFND	5	817	0	0
ISHARES MSCI INTL QUALITY FACT	OTHER	46434V456	5	133	SH		DFND	1	133	0	0
ISHARES MSCL INTL DEVELPD QLTY	OTHER	46434V456	13	373	SH		DFND	5	373	0	0
ISHARES MSCI ACWI LOW CARBON E	OTHER	46434V464	113	760	SH		DFND	1	760	0	0
ISHARES CORE TOTAL USD BOND ET	OTHER	46434V613	35	646	SH		DFND	1	646	0	0
ISHARES CORE DIVIDEND GROWTH E	OTHER	46434V621	2,960	66,032	SH		DFND	1	66,032	0	0
ISHARES CORE DIVIDEND GROWTH	OTHER	46434V621	878	19,593	SH		DFND	5	1,571	0	18,022
ISHARES CURRENCY HEDGED MSCI E	OTHER	46434V639	871	27,929	SH		DFND	1	27,929	0	0
ISHARES CURRENCY HEDGED MSCI E	OTHER	46434V639	92	2,946	SH		OTR	1	2,946	0	0
ISHARES GLOBAL REIT ETF	OTHER	46434V647	31	1,300	SH		DFND	1	1,300	0	0
ISHARES GLOBAL REIT ETF	OTHER	46434V647	442	18,453	SH		OTR	1	18,453	0	0
ISHARES CORE MSCI PACIFIC ETF	OTHER	46434V696	524	8,130	SH		DFND	1	8,130	0	0
ISHARES CORE MSCI PACIFIC ETF	OTHER	46434V696	226	3,502	SH		OTR	1	3,502	0	0
ISHARES CORE MSCI EUROPE ETF	OTHER	46434V738	8,768	171,000	SH		DFND	1	170,500	0	500
ISHARES CORE MSCI EUROPE ETF	OTHER	46434V738	781	15,232	SH		OTR	1	15,232	0	0
ISHARES CURRENCY HEDGED MSCI E	OTHER	46434V803	42,104	1,381,807	SH		DFND	1	1,368,623	0	13,184
ISHARES CURRENCY HEDGED MSCI E	OTHER	46434V803	5,973	196,033	SH		OTR	1	191,114	3,419	1,500
ISHARES CURRENCY HEDGED MSCI	OTHER	46434V803	42	1,373	SH		DFND	5	283	0	1,090
BLACKROCK ULTRA SHORT TERM BON	OTHER	46434V878	121	2,400	SH		DFND	1	0	0	2,400
BLACKROCK ULTRA SHORT TERM BON	OTHER	46434V878	10	200	SH		OTR	1	0	200	0
ISHARES LIQUIDITY INCOME	OTHER	46434V878	3	55	SH		DFND	5	0	0	55
ISHARES CURRENCY HEDGED MSCI E	OTHER	46434V886	3,419	95,323	SH		DFND	1	95,323	0	0
ISHARES CURRENCY HEDGED MSCI E	OTHER	46434V886	239	6,650	SH		OTR	1	6,650	0	0
ISHARES IBONDS DEC 2023 TERM C	OTHER	46434VAX8	31,363	1,193,846	SH		DFND	1	1,173,223	0	20,623
ISHARES IBONDS DEC 2023 TERM C	OTHER	46434VAX8	3,262	124,188	SH		OTR	1	121,746	0	2,442
ISHARES IBONDS DEC 2022 TERM C	OTHER	46434VBA7	31,042	1,214,014	SH		DFND	1	1,197,254	0	16,760
ISHARES IBONDS DEC 2022 TERM C	OTHER	46434VBA7	3,210	125,546	SH		OTR	1	123,305	941	1,300
ISHARES IBONDS DEC 2025 TERM C	OTHER	46434VBD1	17,461	637,947	SH		DFND	1	630,206	0	7,741
ISHARES IBONDS DEC 2025 TERM C	OTHER	46434VBD1	1,631	59,599	SH		OTR	1	59,599	0	0
ISHARES IBONDS DEC 2025 CORP	OTHER	46434VBD1	35	1,274	SH		DFND	5	0	0	1,274
ISHARES IBONDS DEC 2024 TERM C	OTHER	46434VBG4	25,728	967,949	SH		DFND	1	948,866	0	19,083
ISHARES IBONDS DEC 2024 TERM C	OTHER	46434VBG4	2,389	89,894	SH		OTR	1	87,475	0	2,419
ISHARES IBONDS DEC 2021 TERM C	OTHER	46434VBK5	37,243	1,495,107	SH		DFND	1	1,472,915	0	22,192
ISHARES IBONDS DEC 2021 TERM C	OTHER	46434VBK5	2,032	81,578	SH		OTR	1	77,139	1,573	2,866
ISHARES EDGE HIGH YIELD	OTHER	46435G250	123	2,399	SH		DFND	5	2,399	0	0
ISHARES RUSSELL 2500 ETF	OTHER	46435G268	7,974	140,483	SH		DFND	1	136,709	0	3,774
ISHARES RUSSELL 2500 ETF	OTHER	46435G268	160	2,821	SH		OTR	1	2,547	0	274
ISHARES DEC 2023 MUNI BD ETF	OTHER	46435G318	11,929	455,556	SH		DFND	1	455,556	0	0
ISHARES DEC 2023 MUNI BD ETF	OTHER	46435G318	1,886	72,025	SH		OTR	1	68,525	0	3,500
ISHARES CORE MSCI INTERNATIONA	OTHER	46435G326	400	6,494	SH		DFND	1	6,494	0	0
ISHARES MSCI UNITED KINGDOM ET	OTHER	46435G334	35	1,200	SH		DFND	1	1,200	0	0
ISHARES MORTGAGE REAL ESTATE E	OTHER	46435G342	6	180	SH		DFND	1	180	0	0
ISHARES US FALLEN ANGELS USD	OTHER	46435G474	94	3,207	SH		DFND	5	0	0	3,207
ISHARES TRUST ISHARES ESG AWAR	OTHER	46435G516	7,828	107,168	SH		DFND	1	103,148	0	4,020
ISHARES TRUST ISHARES ESG AWAR	OTHER	46435G516	1,986	27,189	SH		OTR	1	27,189	0	0
ISHARES MSCI GLOBAL IMPACT ETF	OTHER	46435G532	59	630	SH		DFND	1	630	0	0
ISHARES IBONDS DEC 2022 TERM E	OTHER	46435G755	21,463	814,241	SH		DFND	1	814,241	0	0
ISHARES IBONDS DEC 2022 TERM E	OTHER	46435G755	5,260	199,554	SH		OTR	1	189,134	0	10,420
ISHARES IBONDS DEC 2021 TERM E	OTHER	46435G789	11,930	462,853	SH		DFND	1	462,853	0	0
ISHARES IBONDS DEC 2021 TERM E	OTHER	46435G789	3,004	116,537	SH		OTR	1	103,067	7,120	6,350
ISHARES IBONDS DEC 2026 TERM C	OTHER	46435GAA0	10,325	380,987	SH		DFND	1	379,562	0	1,425
ISHARES IBONDS DEC 2026 TERM C	OTHER	46435GAA0	990	36,522	SH		OTR	1	36,522	0	0
ISHARES IBONDS DEC 2026 TERM	OTHER	46435GAA0	35	1,287	SH		DFND	5	0	0	1,287
ISHARES CYBERSECURITY AND TECH	OTHER	46435U135	6	140	SH		OTR	1	140	0	0
ISHARES IBONDS DEC 2026 TERM E	OTHER	46435U259	1,787	65,978	SH		DFND	1	65,978	0	0
ISHARES IBONDS DEC 2026 TERM E	OTHER	46435U259	208	7,687	SH		OTR	1	7,687	0	0
ISHARES IBONDS DEC 2027 TERM E	OTHER	46435U283	3,658	133,949	SH		DFND	1	133,949	0	0
ISHARES IBONDS DEC 2027 TERM E	OTHER	46435U283	209	7,640	SH		OTR	1	7,640	0	0
ISHARES IBONDS DEC 2028 TERM E	OTHER	46435U325	2,129	77,139	SH		DFND	1	77,139	0	0
ISHARES IBONDS DEC 2028 TERM E	OTHER	46435U325	471	17,072	SH		OTR	1	17,072	0	0
ISHARES SELF DRIVING EV AND TE	OTHER	46435U366	30	700	SH		DFND	1	700	0	0
ISHARES IBONDS DEC 2025 TERM E	OTHER	46435U432	5,111	181,425	SH		DFND	1	181,425	0	0
ISHARES IBONDS DEC 2025 TERM E	OTHER	46435U432	333	11,809	SH		OTR	1	9,348	0	2,461
ISHARES IBONDS DEC 2028 TERM C	OTHER	46435U515	4,749	160,702	SH		DFND	1	160,222	0	480
ISHARES IBONDS DEC 2028 TERM C	OTHER	46435U515	21	700	SH		OTR	1	700	0	0
I SHARES IBONDS DEC 2028	OTHER	46435U515	35	1,181	SH		DFND	5	0	0	1,181
ISHARES IBONDS DEC 2024 TERM E	OTHER	46435U697	6,067	223,925	SH		DFND	1	223,925	0	0
ISHARES IBONDS DEC 2024 TERM E	OTHER	46435U697	684	25,242	SH		OTR	1	25,242	0	0
ISHARES BROAD USD HIGH YIELD C	OTHER	46435U853	14,815	359,063	SH		DFND	1	359,063	0	0
ISHARES BROAD USD HIGH YIELD	OTHER	46435U853	71	1,709	SH		DFND	5	1,709	0	0
ISHARES IBONDS DEC 2027 TERM C	OTHER	46435UAA9	6,398	232,065	SH		DFND	1	232,065	0	0
ISHARES IBONDS DEC 2027 TERM C	OTHER	46435UAA9	394	14,300	SH		OTR	1	14,300	0	0
ISHARES IBONDS DEC 2027 TERM	OTHER	46435UAA9	35	1,270	SH		DFND	5	0	0	1,270
I3 VERTICALS INC CLASS A	COMMON STOCK	46571Y107	125	3,766	SH		DFND	1	0	0	3,766
JBG SMITH PROPERTIES	COMMON STOCK	46590V100	79	2,540	SH		DFND	1	1,426	0	1,114
JBG SMITH PROPERTIES	COMMON STOCK	46590V100	9	286	SH		OTR	1	40	0	246
JOYY INC A D R	COMMON STOCK	46591M109	242	3,027	SH		DFND	1	1,931	0	1,096
JOYY INC A D R	COMMON STOCK	46591M109	0	6	SH		OTR	1	0	0	6
J P MORGAN CHASE CO	COMMON STOCK	46625H100	372,368	2,930,417	SH		DFND	1	2,804,830	0	125,587
J P MORGAN CHASE CO	COMMON STOCK	46625H100	111,604	878,288	SH		OTR	1	803,646	35,955	38,687
JP MORGAN CHASE & CO	COMMON STOCK	46625H100	769	6,045	SH		DFND	5	6,007	0	38
J P MORGAN CHASE CO	COMMON STOCK	46625H100	4	34	SH	Put	DFND	1	34	0	0
J P MORGAN CHASE CO	COMMON STOCK	46625H100	13	50	SH	Put	OTR	1	0	50	0
JPMORGAN ALERIAN MLP INDEX	OTHER	46625H365	13	929	SH		DFND	5	0	0	929
JMP GROUP LLC	COMMON STOCK	46629U107	39	10,000	SH		DFND	1	10,000	0	0
JPMORGAN DIVERSIFIED RETURN ET	OTHER	46641Q209	1	20	SH		DFND	1	20	0	0
JPMORGAN BETABUILDERS DEVELOPE	OTHER	46641Q688	18	682	SH		DFND	1	682	0	0
JPMORGAN BETABUILDERS CANADA E	OTHER	46641Q696	66	2,467	SH		DFND	1	2,467	0	0
JPMORGAN BETABUILDERS JAPAN ET	OTHER	46641Q712	142	5,083	SH		DFND	1	5,083	0	0
JPMORGAN BETABUILDERS EUROPE E	OTHER	46641Q720	98	3,737	SH		DFND	1	3,737	0	0
JPMORGAN ULTRA SHORT INCOME ET	OTHER	46641Q837	299	5,885	SH		DFND	1	885	0	5,000
JPMORGAN ULTRA SHORT INCOME	OTHER	46641Q837	3	54	SH		DFND	5	0	0	54
JAMES HARDIE INDS PLC SPON A D	COMMON STOCK	47030M106	499	16,838	SH		DFND	1	7,801	0	9,037
JAMES HARDIE INDS PLC SPON A D	COMMON STOCK	47030M106	25	846	SH		OTR	1	177	393	276
JAMF HLDG CORP COM	COMMON STOCK	47074L105	2	60	SH		DFND	1	60	0	0
JAMF HLDG CORP COM	COMMON STOCK	47074L105	34	1,141	SH		OTR	1	0	1,141	0
JAPAN SMALLER CAPITALIZATION F	OTHER	47109U104	9	1,000	SH		DFND	1	1,000	0	0
JD COM INC A D R	COMMON STOCK	47215P106	1,960	22,304	SH		DFND	1	18,152	0	4,152
JD COM INC A D R	COMMON STOCK	47215P106	744	8,460	SH		OTR	1	1,916	751	5,793
JEFFERIES FINANCIAL GROUP INC	COMMON STOCK	47233W109	487	19,817	SH		DFND	1	12,901	0	6,916
JEFFERIES FINANCIAL GROUP INC	COMMON STOCK	47233W109	98	3,967	SH		OTR	1	3,888	0	79
JELD WEN HOLDING INC	COMMON STOCK	47580P103	224	8,847	SH		DFND	1	8,699	0	148
JELD WEN HOLDING INC	COMMON STOCK	47580P103	119	4,713	SH		OTR	1	4,669	0	44
JINKOSOLAR HOLDING CO A D R	COMMON STOCK	47759T100	590	9,529	SH		DFND	1	8,418	0	1,111
JINKOSOLAR HOLDING CO A D R	COMMON STOCK	47759T100	4	59	SH		OTR	1	59	0	0
JONES LANG LASALLE INC	COMMON STOCK	48020Q107	93	628	SH		DFND	1	316	0	312
JONES LANG LASALLE INC	COMMON STOCK	48020Q107	3	20	SH		DFND	5	20	0	0
J2 GLOBAL INC	COMMON STOCK	48123V102	665	6,811	SH		DFND	1	6,603	0	208
J2 GLOBAL INC	COMMON STOCK	48123V102	63	645	SH		OTR	1	95	550	0
JUMIA TECHNOLOGIES AG A D R	COMMON STOCK	48138M105	143	3,541	SH		DFND	1	3,541	0	0
JUMIA TECHNOLOGIES AG A D R	COMMON STOCK	48138M105	52	1,282	SH		OTR	1	0	1,282	0
JUNIPER NETWORKS INC	COMMON STOCK	48203R104	511	22,729	SH		DFND	1	19,777	0	2,952
JUNIPER NETWORKS INC	COMMON STOCK	48203R104	75	3,331	SH		OTR	1	2,898	19	414
JUNIPER NETWORKS INC	COMMON STOCK	48203R104	2	72	SH		DFND	5	72	0	0
KAR AUCTION SERVICES INC	COMMON STOCK	48238T109	1,130	60,700	SH		DFND	1	60,500	0	200
KAR AUCTION SERVICES INC	COMMON STOCK	48238T109	34	1,826	SH		OTR	1	1,769	0	57
KB FINANCIAL GROUP A D R	COMMON STOCK	48241A105	803	20,261	SH		DFND	1	9,774	0	10,487
KB FINANCIAL GROUP A D R	COMMON STOCK	48241A105	3	71	SH		OTR	1	0	21	50
KBR INC	COMMON STOCK	48242W106	32	1,047	SH		DFND	1	803	0	244
KKR REAL ESTATE FINANCE TRUS	COMMON STOCK	48251K100	19	1,035	SH		DFND	1	606	0	429
KKR REAL ESTATE FINANCE TRUS	COMMON STOCK	48251K100	1	51	SH		OTR	1	0	51	0
KKR CO INC A	COMMON STOCK	48251W104	87	2,148	SH		DFND	1	2,148	0	0
KKR CO INC A	COMMON STOCK	48251W104	162	4,000	SH		OTR	1	4,000	0	0
KT CORP SP A D R	COMMON STOCK	48268K101	4,841	439,704	SH		DFND	1	439,703	0	1
KT CORP SP A D R	COMMON STOCK	48268K101	1,171	106,317	SH		OTR	1	106,317	0	0
KT CORP	COMMON STOCK	48268K101	11	981	SH		DFND	5	981	0	0
KADANT INC	COMMON STOCK	48282T104	251	1,783	SH		DFND	1	1,745	0	38
KADANT INC	COMMON STOCK	48282T104	88	623	SH		OTR	1	291	332	0
KARUNA THERAPEUTICS INC	COMMON STOCK	48576A100	412	4,056	SH		DFND	1	4,050	0	6
KARUNA THERAPEUTICS INC	COMMON STOCK	48576A100	7	65	SH		OTR	1	65	0	0
KARYOPHARM THERAPEUTICS INC	COMMON STOCK	48576U106	2	144	SH		DFND	1	144	0	0
K B HOME	COMMON STOCK	48666K109	1,396	41,658	SH		DFND	1	40,674	0	984
K B HOME	COMMON STOCK	48666K109	58	1,740	SH		OTR	1	1,046	0	694
KB HOME	COMMON STOCK	48666K109	5	157	SH		DFND	5	157	0	0
KEARNY FINANCIAL CORP MD	COMMON STOCK	48716P108	106	10,067	SH		DFND	1	10,067	0	0
KEURIG DR PEPPER INC	COMMON STOCK	49271V100	1,439	44,977	SH		DFND	1	41,306	0	3,671
KEURIG DR PEPPER INC	COMMON STOCK	49271V100	99	3,098	SH		OTR	1	1,295	43	1,760
KEURIG DR PEPPER INC	COMMON STOCK	49271V100	26	813	SH		DFND	5	813	0	0
KEYSIGHT TECHNOLOGIES INC	COMMON STOCK	49338L103	7,841	59,367	SH		DFND	1	57,609	0	1,758
KEYSIGHT TECHNOLOGIES INC	COMMON STOCK	49338L103	3,198	24,213	SH		OTR	1	19,742	3,975	496
KEYSIGHT TECHNOLOGIES	COMMON STOCK	49338L103	140	1,057	SH		DFND	5	1,039	0	18
KILROY RLTY CORP	COMMON STOCK	49427F108	66	1,147	SH		DFND	1	576	0	571
KILROY RLTY CORP	COMMON STOCK	49427F108	9	157	SH		OTR	1	0	0	157
KIMBALL ELECTRONICS INC	COMMON STOCK	49428J109	11	660	SH		DFND	1	660	0	0
KIMCO REALTY CORP	COMMON STOCK	49446R109	1,625	108,247	SH		DFND	1	105,285	0	2,962
KIMCO REALTY CORP	COMMON STOCK	49446R109	129	8,575	SH		OTR	1	564	0	8,011
KINDER MORGAN INC	COMMON STOCK	49456B101	1,706	124,781	SH		DFND	1	113,478	0	11,303
KINDER MORGAN INC	COMMON STOCK	49456B101	620	45,417	SH		OTR	1	39,017	3,500	2,900
KINDER MORGAN INC	COMMON STOCK	49456B101	10	704	SH		DFND	5	704	0	0
KINSALE CAPITAL GROUP INC	COMMON STOCK	49714P108	392	1,959	SH		DFND	1	182	0	1,777
KINSALE CAPITAL GROUP INC	COMMON STOCK	49714P108	57	285	SH		OTR	1	0	285	0
KINSALE CAPITAL GROUP	COMMON STOCK	49714P108	38	188	SH		DFND	5	188	0	0
KITE REALTY GROUP TRUST	COMMON STOCK	49803T300	242	16,177	SH		DFND	1	14,564	0	1,613
KITE REALTY GROUP TRUST	COMMON STOCK	49803T300	15	1,020	SH		OTR	1	1,020	0	0
KNOWLES CORP	COMMON STOCK	49926D109	507	27,513	SH		DFND	1	26,893	0	620
KNOWLES CORP	COMMON STOCK	49926D109	148	8,053	SH		OTR	1	8,053	0	0
KODIAK SCIENCES INC	COMMON STOCK	50015M109	10	66	SH		DFND	1	66	0	0
KONTOOR BRANDS INC W I	COMMON STOCK	50050N103	130	3,200	SH		DFND	1	3,127	0	73
KONTOOR BRANDS INC W I	COMMON STOCK	50050N103	40	978	SH		OTR	1	772	13	193
KONTOOR BRANDS INC	COMMON STOCK	50050N103	4	103	SH		DFND	5	103	0	0
KOPPERS HLDGS INC	COMMON STOCK	50060P106	1	30	SH		DFND	1	0	0	30
KRATOS DEFENSE & SECURITY	COMMON STOCK	50077B207	23	819	SH		DFND	1	819	0	0
KRATOS DEFENSE & SECURITY	COMMON STOCK	50077B207	69	2,526	SH		OTR	1	204	2,322	0
KRATON CORPORATION	COMMON STOCK	50077C106	6	221	SH		DFND	1	221	0	0
KRONOS WORLDWIDE INC	COMMON STOCK	50105F105	18	1,187	SH		DFND	1	1,099	0	88
KRONOS WORLDWIDE INC	COMMON STOCK	50105F105	2	146	SH		OTR	1	0	146	0
KURA ONCOLOGY INC	COMMON STOCK	50127T109	3	77	SH		DFND	1	77	0	0
LCNB CORPORATION	COMMON STOCK	50181P100	10	650	SH		DFND	1	650	0	0
LCNB CORPORATION	COMMON STOCK	50181P100	15	1,000	SH		OTR	1	1,000	0	0
LG DISPLAY CO LTD A D R	COMMON STOCK	50186V102	4	450	SH		DFND	1	450	0	0
LG DISPLAY CO LTD	COMMON STOCK	50186V102	1	69	SH		DFND	5	69	0	0
LHC GROUP INC	COMMON STOCK	50187A107	2,244	10,518	SH		DFND	1	8,579	0	1,939
LHC GROUP INC	COMMON STOCK	50187A107	1,413	6,625	SH		OTR	1	5,802	823	0
LHC GROUP INC	COMMON STOCK	50187A107	13	59	SH		DFND	5	59	0	0
LGI HOMES INC	COMMON STOCK	50187T106	20	188	SH		DFND	1	46	0	142
LCI INDUSTRIES	COMMON STOCK	50189K103	776	5,985	SH		DFND	1	5,906	0	79
LCI INDUSTRIES	COMMON STOCK	50189K103	6	48	SH		OTR	1	48	0	0
LI AUTO INC A D R	COMMON STOCK	50202M102	2	63	SH		DFND	1	34	0	29
LPL FINANCIAL HOLDINGS INC	COMMON STOCK	50212V100	4,422	42,433	SH		DFND	1	41,379	0	1,054
LPL FINANCIAL HOLDINGS INC	COMMON STOCK	50212V100	698	6,690	SH		OTR	1	6,542	108	40
L S I INDUSTRIES INC	COMMON STOCK	50216C108	13	1,569	SH		DFND	1	1,569	0	0
LABORATORY CORP OF AMERICA HOL	COMMON STOCK	50540R409	2,712	13,326	SH		DFND	1	11,991	0	1,335
LABORATORY CORP OF AMERICA HOL	COMMON STOCK	50540R409	430	2,112	SH		OTR	1	2,090	15	7
LABORATORY CORP OF AMERICA	COMMON STOCK	50540R409	29	141	SH		DFND	5	141	0	0
LANDMARK INFRASTRUCTURE PARTNE	COMMON STOCK	51508J108	21	2,000	SH		OTR	1	0	2,000	0
LANDS END INC	COMMON STOCK	51509F105	3	146	SH		DFND	1	0	0	146
LEAP THERAPEUTICS INC	COMMON STOCK	52187K101	14	6,100	SH		OTR	1	6,100	0	0
WESTERN ASSET TOTAL RETURN ETF	OTHER	52468L810	17	590	SH		DFND	1	590	0	0
LEMONADE INC COM	COMMON STOCK	52567D107	213	1,741	SH		DFND	1	1,741	0	0
LEMONADE INC COM	COMMON STOCK	52567D107	12	100	SH		OTR	1	100	0	0
LEMONADE INC	COMMON STOCK	52567D107	11	87	SH		DFND	5	87	0	0
LENDINGCLUB CORP	COMMON STOCK	52603A208	4	356	SH		DFND	1	204	0	152
LENDINGTREE INC	COMMON STOCK	52603B107	1	5	SH		DFND	1	5	0	0
LENSAR INC W I	COMMON STOCK	52634L108	2	215	SH		DFND	1	0	0	215
LENSAR INC	COMMON STOCK	52634L108	0	1	SH		DFND	5	1	0	0
LEVEL ONE BANCORP INC	COMMON STOCK	52730D208	10	498	SH		OTR	1	498	0	0
LEVI STRAUSS CO CLASS A	COMMON STOCK	52736R102	3	160	SH		DFND	1	0	0	160
LIBERTY OILFIELD SVCS INC	COMMON STOCK	53115L104	4	392	SH		DFND	1	392	0	0
LIGAND PHARMACEUTICALS CL B	COMMON STOCK	53220K504	9	95	SH		DFND	1	78	0	17
LIGAND PHARMACEUTICALS CL B	COMMON STOCK	53220K504	23	231	SH		OTR	1	60	33	138
LIFE STORAGE STOCK	COMMON STOCK	53223X107	1,014	8,491	SH		DFND	1	8,257	0	234
LIFE STORAGE STOCK	COMMON STOCK	53223X107	54	458	SH		OTR	1	400	48	10
LIGHTBRIDGE CORP	COMMON STOCK	53224K302	2	500	SH		DFND	1	500	0	0
LITHIUM AMERICAS CORP	COMMON STOCK	53680Q207	13	1,000	SH		DFND	1	1,000	0	0
LITHIUM AMERICAS CORP	COMMON STOCK	53680Q207	13	1,000	SH		OTR	1	0	0	1,000
LIVE OAK BANCSHARES INC	COMMON STOCK	53803X105	149	3,142	SH		DFND	1	3,113	0	29
LIVENT CORP	COMMON STOCK	53814L108	203	10,797	SH		DFND	1	3,727	0	7,070
LIVENT CORP	COMMON STOCK	53814L108	73	3,872	SH		OTR	1	3,127	273	472
LIVERAMP HOLDINGS INC	COMMON STOCK	53815P108	446	6,096	SH		DFND	1	4,654	0	1,442
LIVERAMP HOLDINGS INC	COMMON STOCK	53815P108	8	112	SH		OTR	1	112	0	0
LOGICBIO THERAPEUTICS INC	COMMON STOCK	54142F102	15	2,000	SH		DFND	1	2,000	0	0
LUMBER LIQUIDATORS HOLDINGS	COMMON STOCK	55003T107	2	80	SH		DFND	1	0	0	80
LUMENTUM HOLDINGS INC W I	COMMON STOCK	55024U109	134	1,419	SH		DFND	1	1,405	0	14
LUMENTUM HOLDINGS INC W I	COMMON STOCK	55024U109	34	362	SH		OTR	1	42	0	320
LUMENTUM HOLDINGS INC	COMMON STOCK	55024U109	33	348	SH		DFND	5	348	0	0
LUMOS PHARMA INC	COMMON STOCK	55028X109	4	111	SH		DFND	1	111	0	0
LYFT INC A	COMMON STOCK	55087P104	611	12,430	SH		DFND	1	10,871	0	1,559
LYFT INC A	COMMON STOCK	55087P104	110	2,239	SH		OTR	1	2,035	115	89
LYFT INC	COMMON STOCK	55087P104	112	2,270	SH		DFND	5	2,270	0	0
M T BANK CORP	COMMON STOCK	55261F104	707	5,555	SH		DFND	1	4,853	0	702
M T BANK CORP	COMMON STOCK	55261F104	402	3,162	SH		OTR	1	20	86	3,056
MFA FINANCIAL INC	COMMON STOCK	55272X102	17	4,450	SH		OTR	1	0	0	4,450
M F S INTER INCOME TR SBI	OTHER	55273C107	23	6,000	SH		DFND	1	6,000	0	0
MGE ENERGY INC	COMMON STOCK	55277P104	2,163	30,886	SH		DFND	1	30,838	0	48
MEI PHARMA INC	COMMON STOCK	55279B202	1	221	SH		DFND	1	0	0	221
MGM GROWTH PROPERTIES LLC A	COMMON STOCK	55303A105	3	84	SH		DFND	1	84	0	0
MGM GROWTH PROPERTIES LLC	COMMON STOCK	55303A105	6	201	SH		DFND	5	201	0	0
MGP INGREDIENTS INC	COMMON STOCK	55303J106	84	1,777	SH		DFND	1	250	0	1,527
MGP INGREDIENTS INC	COMMON STOCK	55303J106	12	250	SH		OTR	1	0	250	0
MGP INGREDIENTS INC	COMMON STOCK	55303J106	10	211	SH		DFND	5	211	0	0
M / I HOMES INC	COMMON STOCK	55305B101	134	3,014	SH		DFND	1	2,403	0	611
M K S INSTRUMENTS INC	COMMON STOCK	55306N104	831	5,520	SH		DFND	1	5,392	0	128
M K S INSTRUMENTS INC	COMMON STOCK	55306N104	45	300	SH		OTR	1	204	85	11
MPLX LP	COMMON STOCK	55336V100	742	34,277	SH		DFND	1	34,277	0	0
MPLX LP	COMMON STOCK	55336V100	278	12,845	SH		OTR	1	6,810	0	6,035
MPLX LP	COMMON STOCK	55336V100	8	359	SH		DFND	5	359	0	0
MRC GLOBAL INC	COMMON STOCK	55345K103	2	353	SH		DFND	1	353	0	0
MSCI INC	COMMON STOCK	55354G100	2,061	4,616	SH		DFND	1	3,796	0	820
MSCI INC	COMMON STOCK	55354G100	532	1,192	SH		OTR	1	768	363	61
MSCI INC	COMMON STOCK	55354G100	493	1,104	SH		DFND	5	1,104	0	0
MYR GROUP INC DELAWARE	COMMON STOCK	55405W104	7	110	SH		DFND	1	110	0	0
MACOM TECHNOLOGY SOLUTIONS H	COMMON STOCK	55405Y100	128	2,333	SH		DFND	1	84	0	2,249
MACQUARIE FIRST TR GLOBAL	OTHER	55607W100	15	1,750	SH		DFND	1	0	0	1,750
MACQUARIE INFRASTRUCTURE	COMMON STOCK	55608B105	121	3,221	SH		DFND	1	2,921	0	300
MACQUARIE INFRASTRUCTURE	COMMON STOCK	55608B105	4	108	SH		OTR	1	108	0	0
MACYS INC	COMMON STOCK	55616P104	127	11,278	SH		DFND	1	6,870	0	4,408
MACYS INC	COMMON STOCK	55616P104	116	10,288	SH		OTR	1	9,485	803	0
MADISON SQUARE GARDEN SPORTS C	COMMON STOCK	55825T103	107	580	SH		DFND	1	466	0	114
MADISON SQUARE GARDEN SPORTS C	COMMON STOCK	55825T103	1	5	SH		OTR	1	5	0	0
MADISON SQUARE GRDN ENTERTNMNT	COMMON STOCK	55826T102	80	766	SH		DFND	1	672	0	94
MADISON SQUARE GRDN ENTERTNMNT	COMMON STOCK	55826T102	1	5	SH		OTR	1	5	0	0
MAGNITE INC COM	COMMON STOCK	55955D100	6	189	SH		DFND	1	189	0	0
MAIN STREET CAPITAL CORP CL C	COMMON STOCK	56035L104	9	287	SH		DFND	1	287	0	0
MAIN ST BANCSHS	COMMON STOCK	56064Y100	42	2,510	SH		OTR	1	2,510	0	0
MALIBU BOATS INC A	COMMON STOCK	56117J100	977	15,655	SH		DFND	1	15,655	0	0
MALIBU BOATS INC A	COMMON STOCK	56117J100	15	239	SH		OTR	1	239	0	0
MANNKIND CORP	COMMON STOCK	56400P706	29	9,345	SH		DFND	1	9,345	0	0
MANPOWERGROUP INC	COMMON STOCK	56418H100	97	1,075	SH		DFND	1	732	0	343
MANPOWERGROUP INC	COMMON STOCK	56418H100	6	66	SH		OTR	1	52	0	14
MANULIFE FINANCIAL CORP	COMMON STOCK	56501R106	1,465	82,254	SH		DFND	1	77,838	0	4,416
MANULIFE FINANCIAL CORP	COMMON STOCK	56501R106	98	5,502	SH		OTR	1	4,574	84	844
MANULIFE FINANCIAL CORP	COMMON STOCK	56501R106	3	157	SH		DFND	5	0	0	157
MARATHON PETROLEUM CORP	COMMON STOCK	56585A102	96,270	2,327,626	SH		DFND	1	2,308,074	0	19,552
MARATHON PETROLEUM CORP	COMMON STOCK	56585A102	17,585	425,165	SH		OTR	1	411,221	9,300	4,644
MARATHON PETROLEUM CORP	COMMON STOCK	56585A102	35	850	SH		DFND	5	850	0	0
MARATHON PATENT GROUP INC COM	COMMON STOCK	56585W401	7	700	SH		DFND	1	700	0	0
MARAVAI LIFESCIENCES HLDGS INC	COMMON STOCK	56600D107	1	31	SH		DFND	1	31	0	0
MARAVAI LIFESCIENCES HOLDINGS	COMMON STOCK	56600D107	0	13	SH		DFND	5	13	0	0
MARKER THERAPEUTICS INC	COMMON STOCK	57055L107	7	5,000	SH		DFND	1	5,000	0	0
MARKER THERAPEUTICS INC	COMMON STOCK	57055L107	12	8,390	SH		OTR	1	8,390	0	0
MARKETAXESS HLDGS INC	COMMON STOCK	57060D108	8,003	14,027	SH		DFND	1	12,162	0	1,865
MARKETAXESS HLDGS INC	COMMON STOCK	57060D108	1,159	2,031	SH		OTR	1	702	1,319	10
MARKETAXESS HOLDINGS INC	COMMON STOCK	57060D108	120	209	SH		DFND	5	209	0	0
MARRIOTT VACATIONS WORLD	COMMON STOCK	57164Y107	1,346	9,814	SH		DFND	1	9,703	0	111
MARRIOTT VACATIONS WORLD	COMMON STOCK	57164Y107	86	626	SH		OTR	1	267	319	40
MASTERCARD INC	COMMON STOCK	57636Q104	282,006	790,062	SH		DFND	1	765,189	0	24,873
MASTERCARD INC	COMMON STOCK	57636Q104	60,450	169,355	SH		OTR	1	147,818	6,520	15,017
MASTERCARD INC	COMMON STOCK	57636Q104	1,187	3,321	SH		DFND	5	3,278	0	43
MATCH GROUP INC NEW	COMMON STOCK	57667L107	75,124	496,882	SH		DFND	1	492,566	0	4,316
MATCH GROUP INC NEW	COMMON STOCK	57667L107	7,362	48,688	SH		OTR	1	46,085	1,340	1,263
MATCH GROUP INC	COMMON STOCK	57667L107	133	879	SH		DFND	5	804	0	75
MATERIALISE NV A D R	COMMON STOCK	57667T100	4	80	SH		DFND	1	80	0	0
MATERIALISE NV A D R	COMMON STOCK	57667T100	25	469	SH		OTR	1	320	149	0
MATSON INC	COMMON STOCK	57686G105	14	239	SH		DFND	1	184	0	55
MATSON INC	COMMON STOCK	57686G105	40	700	SH		OTR	1	0	0	700
MAXIM INTEGRATED PRODS INC	COMMON STOCK	57772K101	18,595	209,753	SH		DFND	1	206,786	0	2,967
MAXIM INTEGRATED PRODS INC	COMMON STOCK	57772K101	3,442	38,821	SH		OTR	1	38,576	180	65
MAXLINEAR INC CLASS A	COMMON STOCK	57776J100	1,431	37,466	SH		DFND	1	37,466	0	0
MAXLINEAR INC CLASS A	COMMON STOCK	57776J100	52	1,354	SH		OTR	1	1,354	0	0
MAXAR TECHNOLOGIES INC	COMMON STOCK	57778K105	39	1,020	SH		DFND	1	723	0	297
MCEWEN MINING INC	COMMON STOCK	58039P107	0	400	SH		DFND	1	400	0	0
MCKESSON CORPORATION	COMMON STOCK	58155Q103	14,320	82,332	SH		DFND	1	81,282	0	1,050
MCKESSON CORPORATION	COMMON STOCK	58155Q103	6,576	37,810	SH		OTR	1	34,306	165	3,339
MCKESSON CORP	COMMON STOCK	58155Q103	45	256	SH		DFND	5	256	0	0
MEDIACO HOLDING INC CL A WI	COMMON STOCK	58450D104	0	13	SH		DFND	1	13	0	0
MEDICAL PROPERTIES TRUST INC	COMMON STOCK	58463J304	659	30,233	SH		DFND	1	29,546	0	687
MEDICAL PROPERTIES TRUST INC	COMMON STOCK	58463J304	33	1,525	SH		OTR	1	1,472	0	53
MEDICAL PROPERTIES TRUST INC	COMMON STOCK	58463J304	36	1,630	SH		DFND	5	1,630	0	0
MEDIFAST INC	COMMON STOCK	58470H101	16	84	SH		DFND	1	50	0	34
MEDNAX INC	COMMON STOCK	58502B106	43	1,730	SH		DFND	1	805	0	925
MEDNAX INC	COMMON STOCK	58502B106	2	74	SH		OTR	1	74	0	0
MEDPACE HLDGS INC	COMMON STOCK	58506Q109	346	2,490	SH		DFND	1	2,282	0	208
MEDPACE HLDGS INC	COMMON STOCK	58506Q109	71	512	SH		OTR	1	512	0	0
MEDPACE HOLDINGS INC	COMMON STOCK	58506Q109	12	88	SH		DFND	5	88	0	0
MERCADOLIBRE INC	COMMON STOCK	58733R102	21,721	12,966	SH		DFND	1	12,942	0	24
MERCADOLIBRE INC	COMMON STOCK	58733R102	1,469	877	SH		OTR	1	675	202	0
MERCADOLIBRE INC	COMMON STOCK	58733R102	1,749	1,044	SH		DFND	5	1,044	0	0
MERCK CO INC	COMMON STOCK	58933Y105	102,061	1,247,693	SH		DFND	1	1,153,432	0	94,261
MERCK CO INC	COMMON STOCK	58933Y105	57,683	705,162	SH		OTR	1	617,111	24,796	63,255
MERCK & COMPANY INC	COMMON STOCK	58933Y105	619	7,566	SH		DFND	5	7,459	0	107
MERIDIAN BANCORP INC MD	COMMON STOCK	58958U103	1	97	SH		DFND	1	97	0	0
MERITAGE HOMES CORPORATION	COMMON STOCK	59001A102	95	1,150	SH		DFND	1	1,081	0	69
MERITOR INC	COMMON STOCK	59001K100	24	858	SH		DFND	1	533	0	325
MERSANA THERAPEUTICS INC	COMMON STOCK	59045L106	2	57	SH		DFND	1	0	0	57
MESA LABS INC	COMMON STOCK	59064R109	2,385	8,322	SH		DFND	1	8,308	0	14
MESA LABS INC	COMMON STOCK	59064R109	449	1,566	SH		OTR	1	766	0	800
METHANEX CORP	COMMON STOCK	59151K108	204	4,423	SH		DFND	1	3,953	0	470
METHANEX CORP	COMMON STOCK	59151K108	11	232	SH		OTR	1	150	26	56
METLIFE INC	COMMON STOCK	59156R108	4,260	90,738	SH		DFND	1	76,340	0	14,398
METLIFE INC	COMMON STOCK	59156R108	723	15,410	SH		OTR	1	10,367	2,257	2,786
METLIFE INC	COMMON STOCK	59156R108	5	95	SH		DFND	5	95	0	0
MICHAELS COS INC THE	COMMON STOCK	59408Q106	12	925	SH		DFND	1	202	0	723
MID-AMER APT CMNTYS INC	COMMON STOCK	59522J103	1,083	8,548	SH		DFND	1	7,533	0	1,015
MID-AMER APT CMNTYS INC	COMMON STOCK	59522J103	168	1,323	SH		OTR	1	1,145	78	100
MILESTONE SCIENTIFIC INC	COMMON STOCK	59935P209	19	9,000	SH		OTR	1	9,000	0	0
MIRATI THERAPEUTICS INC	COMMON STOCK	60468T105	18	81	SH		DFND	1	74	0	7
MIRATI THERAPEUTICS INC	COMMON STOCK	60468T105	1	4	SH		OTR	1	0	0	4
MIZUHO FNL GRP A D R	COMMON STOCK	60687Y109	2,467	967,108	SH		DFND	1	907,260	0	59,848
MIZUHO FNL GRP A D R	COMMON STOCK	60687Y109	447	175,284	SH		OTR	1	173,439	785	1,060
MIZUHO FINANCIAL GROUP	COMMON STOCK	60687Y109	2	668	SH		DFND	5	668	0	0
MODERNA INC	COMMON STOCK	60770K107	610	5,843	SH		DFND	1	4,779	0	1,064
MODERNA INC	COMMON STOCK	60770K107	3,545	33,938	SH		OTR	1	32,858	300	780
MOELIS CO	COMMON STOCK	60786M105	323	6,899	SH		DFND	1	6,854	0	45
MOELIS CO	COMMON STOCK	60786M105	60	1,280	SH		OTR	1	1,280	0	0
MOELIS & CO	COMMON STOCK	60786M105	11	235	SH		DFND	5	235	0	0
MOLINA HEALTHCARE INC	COMMON STOCK	60855R100	258	1,214	SH		DFND	1	959	0	255
MOLINA HEALTHCARE INC	COMMON STOCK	60855R100	215	1,011	SH		OTR	1	0	33	978
MOLINA HEALTHCARE INC	COMMON STOCK	60855R100	51	239	SH		DFND	5	239	0	0
MOLSON COORS BEVERAGE COMPANY	COMMON STOCK	60871R209	96	2,126	SH		DFND	1	1,207	0	919
MOLSON COORS BEVERAGE COMPANY	COMMON STOCK	60871R209	28	627	SH		OTR	1	409	0	218
MOLSON COORS BREWING CO	COMMON STOCK	60871R209	3	71	SH		DFND	5	71	0	0
MOMO INC A D R	COMMON STOCK	60879B107	134	9,638	SH		DFND	1	7,356	0	2,282
MOMO INC A D R	COMMON STOCK	60879B107	0	35	SH		OTR	1	0	0	35
MONGODB INC	COMMON STOCK	60937P106	140	391	SH		DFND	1	211	0	180
MONGODB INC	COMMON STOCK	60937P106	198	551	SH		OTR	1	500	34	17
MONSTER BEVERAGE CORP	COMMON STOCK	61174X109	5,683	61,450	SH		DFND	1	56,893	0	4,557
MONSTER BEVERAGE CORP	COMMON STOCK	61174X109	3,384	36,584	SH		OTR	1	31,919	700	3,965
MONSTER BEVERAGE CORP	COMMON STOCK	61174X109	394	4,258	SH		DFND	5	4,258	0	0
MOSAIC CO THE	COMMON STOCK	61945C103	172	7,481	SH		DFND	1	4,597	0	2,884
MOSAIC CO THE	COMMON STOCK	61945C103	210	9,143	SH		OTR	1	1,460	226	7,457
MR COOPER GROUP INC	COMMON STOCK	62482R107	56	1,792	SH		DFND	1	498	0	1,294
MR COOPER GROUP INC	COMMON STOCK	62482R107	1	42	SH		OTR	1	0	0	42
NGL ENERGY PARTNERS LP	COMMON STOCK	62913M107	14	6,000	SH		OTR	1	6,000	0	0
NIC INC	COMMON STOCK	62914B100	185	7,160	SH		DFND	1	1,160	0	6,000
NIO INC A D R	COMMON STOCK	62914V106	373	7,666	SH		DFND	1	4,713	0	2,953
NIO INC A D R	COMMON STOCK	62914V106	40	818	SH		OTR	1	800	18	0
NGM BIOPHARMACEUTICALS INC	COMMON STOCK	62921N105	8	270	SH		DFND	1	270	0	0
NVR INC	COMMON STOCK	62944T105	294	72	SH		DFND	1	58	0	14
NVR INC	COMMON STOCK	62944T105	28	7	SH		OTR	1	0	3	4
NV5 GLOBAL INC	COMMON STOCK	62945V109	3	39	SH		DFND	1	26	0	13
NANO DIMENSION LTD A D R	COMMON STOCK	63008G203	15	1,700	SH		DFND	1	1,700	0	0
NANOSTRING TECHNOLOGIES INC	COMMON STOCK	63009R109	20	305	SH		OTR	1	305	0	0
NANTKWEST INC	COMMON STOCK	63016Q102	53	3,984	SH		DFND	1	3,660	0	324
NATIONAL HEALTH INVS INC	COMMON STOCK	63633D104	564	8,148	SH		DFND	1	8,066	0	82
NATIONAL HEALTH INVS INC	COMMON STOCK	63633D104	78	1,115	SH		OTR	1	1,085	14	16
NATIONAL VISION HOLDINGS INC	COMMON STOCK	63845R107	444	9,791	SH		DFND	1	9,756	0	35
NATIONAL VISION HOLDINGS INC	COMMON STOCK	63845R107	216	4,762	SH		OTR	1	3,762	1,000	0
NATURA CO HOLDING ADR W I A D	COMMON STOCK	63884N108	138	6,928	SH		DFND	1	3,522	0	3,406
NAVIENT CORP W D	COMMON STOCK	63938C108	95	9,680	SH		DFND	1	3,966	0	5,714
NAVIENT CORP W D	COMMON STOCK	63938C108	1	112	SH		OTR	1	0	0	112
NCINO INC COM	COMMON STOCK	63947U107	1,103	15,234	SH		DFND	1	15,234	0	0
NCINO INC COM	COMMON STOCK	63947U107	262	3,612	SH		OTR	1	3,612	0	0
NELNET INC CL A	COMMON STOCK	64031N108	23	329	SH		DFND	1	218	0	111
NEOGENOMICS INC	COMMON STOCK	64049M209	1,516	28,156	SH		DFND	1	27,997	0	159
NEOGENOMICS INC	COMMON STOCK	64049M209	164	3,041	SH		OTR	1	2,601	440	0
NEOPHOTONICS CORP	COMMON STOCK	64051T100	19	2,099	SH		DFND	1	2,099	0	0
NETAPP INC	COMMON STOCK	64110D104	555	8,385	SH		DFND	1	7,063	0	1,322
NETAPP INC	COMMON STOCK	64110D104	459	6,936	SH		OTR	1	59	216	6,661
NETAPP INC	COMMON STOCK	64110D104	43	654	SH		DFND	5	601	0	53
NETFLIX COM INC	COMMON STOCK	64110L106	25,244	46,684	SH		DFND	1	41,919	0	4,765
NETFLIX COM INC	COMMON STOCK	64110L106	5,562	10,285	SH		OTR	1	9,119	721	445
NETFLIX INC	COMMON STOCK	64110L106	304	562	SH		DFND	5	562	0	0
NETEASE INC A D R	COMMON STOCK	64110W102	1,314	13,720	SH		DFND	1	11,444	0	2,276
NETEASE INC A D R	COMMON STOCK	64110W102	107	1,119	SH		OTR	1	945	0	174
NETEASE INC	COMMON STOCK	64110W102	6	67	SH		DFND	5	67	0	0
NETGEAR INC	COMMON STOCK	64111Q104	143	3,525	SH		DFND	1	3,326	0	199
NETGEAR INC	COMMON STOCK	64111Q104	4	107	SH		OTR	1	0	107	0
NETSCOUT SYS INC	COMMON STOCK	64115T104	377	13,766	SH		DFND	1	13,647	0	119
NETSCOUT SYS INC	COMMON STOCK	64115T104	123	4,458	SH		OTR	1	4,458	0	0
NETSTREIT CORP	COMMON STOCK	64119V303	18	936	SH		OTR	1	0	936	0
NEUBERGER BERMAN INTMD MUNI	OTHER	64124P101	433	28,500	SH		DFND	1	28,500	0	0
NEUROCRINE BIOSCIENCES INC	COMMON STOCK	64125C109	1,424	14,863	SH		DFND	1	14,667	0	196
NEUROCRINE BIOSCIENCES INC	COMMON STOCK	64125C109	598	6,236	SH		OTR	1	3,564	70	2,602
NEUROCRINE BIOSCIENCES INC	COMMON STOCK	64125C109	60	629	SH		DFND	5	615	0	14
NEUBERGER BERMAN HIGH YIELD	OTHER	64128C106	42	3,566	SH		OTR	1	3,566	0	0
NEVRO CORP	COMMON STOCK	64157F103	1	6	SH		OTR	1	0	0	6
NEW RESIDENTIAL INVESTMENT COR	COMMON STOCK	64828T201	55	5,557	SH		DFND	1	3,501	0	2,056
NEW RESIDENTIAL INVESTMENT COR	COMMON STOCK	64828T201	8	852	SH		OTR	1	0	0	852
NEW RELIC INC	COMMON STOCK	64829B100	50	770	SH		DFND	1	708	0	62
NEW RELIC INC	COMMON STOCK	64829B100	8	118	SH		OTR	1	0	96	22
NEWMARK GROUP INC	COMMON STOCK	65158N102	138	18,925	SH		DFND	1	3,515	0	15,410
NEWS CORP NEW CL A W	COMMON STOCK	65249B109	52	2,876	SH		DFND	1	1,324	0	1,552
NEWS CORP NEW CL A W	COMMON STOCK	65249B109	12	671	SH		OTR	1	250	283	138
NEWS CORP CLASS B	COMMON STOCK	65249B208	9	500	SH		DFND	1	500	0	0
NEXTIER OILFIELD SOLUTIONS I	COMMON STOCK	65290C105	5	1,382	SH		DFND	1	1,382	0	0
NEXSTAR BROADCASTING GROUP A	COMMON STOCK	65336K103	961	8,793	SH		DFND	1	8,599	0	194
NEXSTAR BROADCASTING GROUP A	COMMON STOCK	65336K103	163	1,493	SH		OTR	1	1,339	9	145
NEXSTAR BROADCASTING GROUP	COMMON STOCK	65336K103	27	250	SH		DFND	5	250	0	0
NEXTERA ENERGY INC	COMMON STOCK	65339F101	101,471	1,315,252	SH		DFND	1	1,230,256	0	84,996
NEXTERA ENERGY INC	COMMON STOCK	65339F101	47,475	615,358	SH		OTR	1	553,680	13,451	48,227
NEXTERA ENERGY INC	COMMON STOCK	65339F101	266	3,442	SH		DFND	5	3,442	0	0
NEXTERA ENERGY PARTNERS L P	COMMON STOCK	65341B106	550	8,202	SH		DFND	1	7,942	0	260
NEXTERA ENERGY PARTNERS L P	COMMON STOCK	65341B106	222	3,309	SH		OTR	1	700	530	2,079
NEXTERA ENERGY PARTNERS LP	COMMON STOCK	65341B106	4	65	SH		DFND	5	65	0	0
NEXPOINT RESIDENTIAL W I	COMMON STOCK	65341D102	597	14,105	SH		DFND	1	9,210	0	4,895
NEXPOINT RESIDENTIAL W I	COMMON STOCK	65341D102	132	3,121	SH		OTR	1	3,121	0	0
NEXPOINT RESIDENTIAL TRUST INC	COMMON STOCK	65341D102	5	118	SH		DFND	5	118	0	0
NEXTDECADE CORP	COMMON STOCK	65342K105	0	222	SH		DFND	1	0	0	222
NISOURCE INC	COMMON STOCK	65473P105	520	22,669	SH		DFND	1	17,389	0	5,280
NISOURCE INC	COMMON STOCK	65473P105	684	29,828	SH		OTR	1	6,747	0	23,081
NKARTA INC	COMMON STOCK	65487U108	247	4,025	SH		DFND	1	4,025	0	0
NKARTA INC	COMMON STOCK	65487U108	3	50	SH		OTR	1	50	0	0
NOAH HOLDINGS LTD SPON ADS A D	COMMON STOCK	65487X102	204	4,267	SH		DFND	1	2,752	0	1,515
NOAH HOLDINGS LTD SPON ADS A D	COMMON STOCK	65487X102	1	30	SH		OTR	1	0	27	3
NOMURA HOLDINGS INC A D R	COMMON STOCK	65535H208	453	84,952	SH		DFND	1	46,324	0	38,628
NOMURA HOLDINGS INC A D R	COMMON STOCK	65535H208	41	7,752	SH		OTR	1	4,251	2,967	534
NORTHERN DYNASTY MINERALS LTD	COMMON STOCK	66510M204	0	1,000	SH		DFND	1	1,000	0	0
NORTHFIELD BANCORP INC NJ	COMMON STOCK	66611T108	7	595	SH		DFND	1	456	0	139
NORTHWEST NATURAL HOLDING CO	COMMON STOCK	66765N105	571	12,421	SH		DFND	1	8,358	0	4,063
NORTHWEST NATURAL HOLDING CO	COMMON STOCK	66765N105	98	2,123	SH		OTR	1	2,123	0	0
NORTHWEST NATURAL HOLDING CO	COMMON STOCK	66765N105	0	0	SH		DFND	5	0	0	0
NOVARTIS AG A D R	COMMON STOCK	66987V109	19,864	210,356	SH		DFND	1	190,695	0	19,661
NOVARTIS AG A D R	COMMON STOCK	66987V109	7,897	83,630	SH		OTR	1	71,512	3,008	9,110
NOVARTIS AG	COMMON STOCK	66987V109	150	1,585	SH		DFND	5	1,563	0	22
NOVAN INC	COMMON STOCK	66988N106	4	5,000	SH		DFND	1	5,000	0	0
NOVANTA INC	COMMON STOCK	67000B104	480	4,062	SH		DFND	1	140	0	3,922
NOVANTA INC	COMMON STOCK	67000B104	274	2,318	SH		OTR	1	100	478	1,740
NOW INC DE W I	COMMON STOCK	67011P100	2	232	SH		DFND	1	44	0	188
NOVUS CAP CORP	COMMON STOCK	67012U108	31	2,000	SH		DFND	1	2,000	0	0
NU SKIN ENTERPRISES INC - A	COMMON STOCK	67018T105	395	7,240	SH		DFND	1	6,237	0	1,003
NU SKIN ENTERPRISES INC - A	COMMON STOCK	67018T105	17	313	SH		OTR	1	49	170	94
NUANCE COMMUNICATIONS INC	COMMON STOCK	67020Y100	1,026	23,264	SH		DFND	1	21,546	0	1,718
NUANCE COMMUNICATIONS INC	COMMON STOCK	67020Y100	204	4,627	SH		OTR	1	3,630	997	0
NUSTAR ENERGY LP	COMMON STOCK	67058H102	12	775	SH		DFND	1	775	0	0
NUSTAR ENERGY LP	COMMON STOCK	67058H102	39	2,695	SH		OTR	1	2,695	0	0
NUSTAR ENERGY LP	COMMON STOCK	67058H102	0	0	SH		DFND	5	0	0	0
NUTANIX INC	COMMON STOCK	67059N108	94	2,942	SH		DFND	1	2,509	0	433
NUTANIX INC	COMMON STOCK	67059N108	3	95	SH		OTR	1	0	56	39
NUTANIX INC	COMMON STOCK	67059N108	153	4,789	SH		DFND	5	4,789	0	0
NUVEEN CALIFORNIA MUNICIPAL VA	OTHER	67062C107	5	500	SH		DFND	1	500	0	0
NUVEEN MUN INCOME FD INC	OTHER	67062J102	57	5,000	SH		OTR	1	0	5,000	0
NVIDIA CORP	COMMON STOCK	67066G104	30,680	58,751	SH		DFND	1	52,021	0	6,730
NVIDIA CORP	COMMON STOCK	67066G104	6,455	12,361	SH		OTR	1	8,849	989	2,523
NVIDIA CORP	COMMON STOCK	67066G104	818	1,565	SH		DFND	5	1,565	0	0
NUVEEN QUALITY MUNI INCOME FUN	OTHER	67066V101	382	24,865	SH		DFND	1	24,865	0	0
NUVEEN QUALITY MUNI INCOME FUN	OTHER	67066V101	193	12,556	SH		OTR	1	12,556	0	0
NUVEEN CALI QUAL MUN INCOME FD	OTHER	67066Y105	19	1,263	SH		DFND	1	1,263	0	0
NUVEEN CALI QUAL MUN INCOME FD	OTHER	67066Y105	250	16,518	SH		OTR	1	16,518	0	0
NUVEEN SENIOR INCOME FUND	OTHER	67067Y104	23	4,500	SH		DFND	1	4,500	0	0
NUVEEN S P 500 BUY WRITE INCOM	OTHER	6706ER101	10	765	SH		DFND	1	765	0	0
NUVEEN S P 500 BUY WRITE INCOM	OTHER	6706ER101	99	7,725	SH		OTR	1	7,725	0	0
NUVEEN MUNICIPAL CREDIT INCOME	OTHER	67070X101	53	3,275	SH		DFND	1	3,275	0	0
NUVEEN ENHCD AMT-FRE MUN CRDT	OTHER	67071L106	491	29,274	SH		DFND	1	29,274	0	0
NUVEEN ENHCD AMT-FRE MUN CRDT	OTHER	67071L106	86	5,110	SH		OTR	1	5,110	0	0
NUVEEN PREFERRED SECURITIES IN	OTHER	67072C105	19	2,000	SH		DFND	1	0	0	2,000
NUVEEN PREFERRED INCOME	OTHER	67073B106	6	643	SH		DFND	1	643	0	0
NUVEEN PFD CONV INCOME FD 2	OTHER	67073D102	2	241	SH		DFND	1	241	0	0
NUVEEN ENHANCED MUNI VALUE COM	OTHER	67074M101	55	3,400	SH		DFND	1	3,400	0	0
NUVEEN DOW 30 DYNAMIC OVERWRIT	OTHER	67075F105	84	5,535	SH		DFND	1	0	0	5,535
NUVEEN DOW 30 DYNAMIC OVERWRIT	OTHER	67075F105	43	2,825	SH		OTR	1	2,825	0	0
NUTRIEN LTD	COMMON STOCK	67077M108	1,813	37,652	SH		DFND	1	15,398	0	22,254
NUTRIEN LTD	COMMON STOCK	67077M108	517	10,736	SH		OTR	1	7,007	3,700	29
NUTRIEN LTD	COMMON STOCK	67077M108	14	291	SH		DFND	5	291	0	0
NUVEEN ESG LARGE CAP GROWTH ET	OTHER	67092P201	37	650	SH		DFND	1	650	0	0
NUVEEN ESG LARGE CAP GROWTH	OTHER	67092P201	57	1,019	SH		DFND	5	1,019	0	0
NUVEEN ESG LARGE CAP VALUE	OTHER	67092P300	54	1,639	SH		DFND	5	1,639	0	0
NUVEEN ESG SMALL CAP GROWTH	OTHER	67092P607	9	223	SH		DFND	5	223	0	0
NUVEEN SHORT TERM REIT	OTHER	67092P706	47	1,760	SH		DFND	5	0	0	1,760
NUVEEN ESG INTERNATIONAL	OTHER	67092P805	36	1,217	SH		DFND	5	1,217	0	0
NUVEEN ESG HIGH YIELD	OTHER	67092P854	19	750	SH		DFND	5	750	0	0
NUVEEN ESG US AGG BD	OTHER	67092P870	42	1,574	SH		DFND	5	1,574	0	0
NUVEEN ESG INTERNATIONAL	OTHER	67092P888	16	489	SH		DFND	5	489	0	0
O I GLASS INC	COMMON STOCK	67098H104	103	8,646	SH		DFND	1	603	0	8,043
OFS CAPITAL CORP	COMMON STOCK	67103B100	11	1,550	SH		OTR	1	1,550	0	0
O REILLY AUTOMOTIVE INC	COMMON STOCK	67103H107	9,886	21,845	SH		DFND	1	14,761	0	7,084
O REILLY AUTOMOTIVE INC	COMMON STOCK	67103H107	2,322	5,132	SH		OTR	1	4,948	72	112
OREILLY AUTOMOTIVE INC	COMMON STOCK	67103H107	33	74	SH		DFND	5	59	0	15
OFG BANCORP	COMMON STOCK	67103X102	7	385	SH		DFND	1	385	0	0
O SHARES FTSE US QUALITY ETF	OTHER	67110P407	7	175	SH		DFND	1	175	0	0
OAKTREE SPECIALTY LENDING CO	COMMON STOCK	67401P108	2	300	SH		DFND	1	300	0	0
OASIS MIDSTREAM PARTNERS LP	COMMON STOCK	67420T206	12	1,000	SH		OTR	1	1,000	0	0
OCULAR THERAPEUTIX INC	COMMON STOCK	67576A100	10	472	SH		DFND	1	0	0	472
OCULAR THERAPEUTIX INC	COMMON STOCK	67576A100	1,708	82,500	SH		OTR	1	82,500	0	0
OMNICELL INC	COMMON STOCK	68213N109	1,090	9,082	SH		DFND	1	6,436	0	2,646
OMNICELL INC	COMMON STOCK	68213N109	497	4,139	SH		OTR	1	1,001	753	2,385
OMNICELL INC	COMMON STOCK	68213N109	18	148	SH		DFND	5	148	0	0
ONCOCYTE CORP	COMMON STOCK	68235C107	1	500	SH		DFND	1	500	0	0
ONE GAS INC	COMMON STOCK	68235P108	232	3,018	SH		DFND	1	2,624	0	394
ONE GAS INC	COMMON STOCK	68235P108	212	2,767	SH		OTR	1	647	1,720	400
1 800 FLOWERS.COM INC	COMMON STOCK	68243Q106	11	417	SH		DFND	1	392	0	25
ONECONNECT FINANCIAL TECHNO A	COMMON STOCK	68248T105	28	1,428	SH		DFND	1	1,428	0	0
ONEMAIN HOLDINGS INC	COMMON STOCK	68268W103	458	9,514	SH		DFND	1	7,374	0	2,140
ONEMAIN HOLDINGS INC	COMMON STOCK	68268W103	9	188	SH		OTR	1	0	23	165
ONESMART INTERNATIONAL A D R	COMMON STOCK	68276W103	10	2,710	SH		DFND	1	588	0	2,122
ONESPAN INC	COMMON STOCK	68287N100	365	17,663	SH		DFND	1	17,663	0	0
ONESPAN INC	COMMON STOCK	68287N100	93	4,509	SH		OTR	1	4,509	0	0
OPERA LTD ADR	COMMON STOCK	68373M107	4	482	SH		DFND	1	482	0	0
OPERA LTD ADR	COMMON STOCK	68373M107	4	439	SH		OTR	1	0	439	0
OPKO HEALTH INC	COMMON STOCK	68375N103	105	26,523	SH		DFND	1	26,523	0	0
OPORTUN FINANCIAL CORP	COMMON STOCK	68376D104	388	20,045	SH		DFND	1	20,045	0	0
OPORTUN FINANCIAL CORP	COMMON STOCK	68376D104	67	3,465	SH		OTR	1	3,465	0	0
ORACLE CORPORATION	COMMON STOCK	68389X105	58,706	907,507	SH		DFND	1	878,270	0	29,237
ORACLE CORPORATION	COMMON STOCK	68389X105	35,097	542,536	SH		OTR	1	512,021	14,553	15,962
ORACLE CORPORATION	COMMON STOCK	68389X105	103	1,595	SH		DFND	5	1,481	0	114
OPTION CARE HEALTH INC	COMMON STOCK	68404L201	13	837	SH		DFND	1	0	0	837
OPTION CARE HEALTH INC	COMMON STOCK	68404L201	68	4,318	SH		OTR	1	21	4,297	0
ORA SURE TECHNOLOGIES INC	COMMON STOCK	68554V108	9	820	SH		DFND	1	677	0	143
ORBITAL ENERGY	COMMON STOCK	68559A109	128	58,372	SH		DFND	1	58,372	0	0
ORCHID ISLAND CAPITAL INC	COMMON STOCK	68571X103	6	1,093	SH		DFND	1	0	0	1,093
ORGANOVO HLDGS INC	COMMON STOCK	68620A203	6	500	SH		DFND	1	500	0	0
ORIGIN BANCORP INC	COMMON STOCK	68621T102	1	31	SH		DFND	1	0	0	31
ORION GROUP HOLDINGS INC	COMMON STOCK	68628V308	865	174,404	SH		DFND	1	174,404	0	0
ORTHOPEDIATRICS CORP	COMMON STOCK	68752L100	297	7,188	SH		DFND	1	7,188	0	0
ORTHOPEDIATRICS CORP	COMMON STOCK	68752L100	30	733	SH		OTR	1	733	0	0
ORTHOPEDIATRICS CORP	COMMON STOCK	68752L100	89	2,162	SH		DFND	5	2,162	0	0
ORTHOFIX MEDICAL INC	COMMON STOCK	68752M108	1	30	SH		DFND	1	30	0	0
OSISKO GOLD ROYALTIES LTD	COMMON STOCK	68827L101	91	7,163	SH		DFND	1	5,950	0	1,213
OTIS WORLDWIDE CORP	COMMON STOCK	68902V107	8,089	119,743	SH		DFND	1	112,038	0	7,705
OTIS WORLDWIDE CORP	COMMON STOCK	68902V107	5,698	84,354	SH		OTR	1	71,690	9,731	2,933
OTIS WORLDWIDE CORP	COMMON STOCK	68902V107	21	312	SH		DFND	5	312	0	0
OUTFRONT MEDIA INC	COMMON STOCK	69007J106	279	14,295	SH		DFND	1	14,104	0	191
OUTFRONT MEDIA INC	COMMON STOCK	69007J106	12	603	SH		OTR	1	241	51	311
OVINTIV INC	COMMON STOCK	69047Q102	112	7,788	SH		DFND	1	6,356	0	1,432
OVINTIV INC	COMMON STOCK	69047Q102	7	515	SH		OTR	1	0	201	314
PBF ENERGY INC CLASS A	COMMON STOCK	69318G106	16	2,207	SH		DFND	1	1,321	0	886
PBF ENERGY INC CLASS A	COMMON STOCK	69318G106	3	387	SH		OTR	1	0	387	0
PC CONNECTION INC	COMMON STOCK	69318J100	191	4,029	SH		DFND	1	3,637	0	392
PDC ENERGY INC	COMMON STOCK	69327R101	246	11,992	SH		DFND	1	7,805	0	4,187
PDC ENERGY INC	COMMON STOCK	69327R101	127	6,163	SH		OTR	1	5,811	352	0
P G E CORP	COMMON STOCK	69331C108	1,356	108,808	SH		DFND	1	106,901	0	1,907
P G E CORP	COMMON STOCK	69331C108	69	5,534	SH		OTR	1	5,444	0	90
PGT INNOVATIONS INC	COMMON STOCK	69336V101	5	231	SH		DFND	1	200	0	31
PGT INNOVATIONS INC	COMMON STOCK	69336V101	2	92	SH		OTR	1	92	0	0
PJT PARTNERS INC	COMMON STOCK	69343T107	71	949	SH		OTR	1	2	947	0
PLDT INC ADR	COMMON STOCK	69344D408	298	10,679	SH		DFND	1	5,978	0	4,701
PLDT INC ADR	COMMON STOCK	69344D408	15	544	SH		OTR	1	0	0	544
PLDT INC	COMMON STOCK	69344D408	2	64	SH		DFND	5	64	0	0
PNM RESOURCES INC	COMMON STOCK	69349H107	484	9,987	SH		DFND	1	9,774	0	213
PNM RESOURCES INC	COMMON STOCK	69349H107	90	1,850	SH		OTR	1	1,850	0	0
PPL CORPORATION	COMMON STOCK	69351T106	1,922	68,149	SH		DFND	1	58,193	0	9,956
PPL CORPORATION	COMMON STOCK	69351T106	538	19,083	SH		OTR	1	17,217	0	1,866
PPL CORP	COMMON STOCK	69351T106	163	5,786	SH		DFND	5	5,786	0	0
PRA HEALTH SCIENCES INC	COMMON STOCK	69354M108	860	6,852	SH		DFND	1	6,533	0	319
PRA HEALTH SCIENCES INC	COMMON STOCK	69354M108	233	1,857	SH		OTR	1	1,058	655	144
PRA GROUP INC	COMMON STOCK	69354N106	1,591	40,097	SH		DFND	1	40,097	0	0
PRA GROUP INC	COMMON STOCK	69354N106	253	6,375	SH		OTR	1	5,083	1,292	0
PRA GROUP INC	COMMON STOCK	69354N106	98	2,464	SH		DFND	5	2,464	0	0
PPD INC	COMMON STOCK	69355F102	5	136	SH		DFND	1	42	0	94
PS BUSINESS PKS INC	COMMON STOCK	69360J107	758	5,702	SH		DFND	1	5,636	0	66
PS BUSINESS PKS INC	COMMON STOCK	69360J107	132	989	SH		OTR	1	962	0	27
PTC THERAPEUTICS INC	COMMON STOCK	69366J200	18	302	SH		DFND	1	150	0	152
PTC INC	COMMON STOCK	69370C100	876	7,318	SH		DFND	1	7,078	0	240
PTC INC	COMMON STOCK	69370C100	276	2,305	SH		OTR	1	1,823	207	275
PTC INC	COMMON STOCK	69370C100	32	265	SH		DFND	5	265	0	0
PACER TRENDPILOT 100 ETF	OTHER	69374H303	39	754	SH		DFND	1	754	0	0
PACIFIC BIOSCIENCES OF CALIF	COMMON STOCK	69404D108	11	437	SH		DFND	1	400	0	37
PACIFIC PREMIER BANCORP INC	COMMON STOCK	69478X105	615	19,628	SH		DFND	1	16,435	0	3,193
PACIFIC PREMIER BANCORP INC	COMMON STOCK	69478X105	124	3,950	SH		OTR	1	2,987	963	0
PAGERDUTY INC	COMMON STOCK	69553P100	87	2,089	SH		DFND	1	1,687	0	402
PAGERDUTY INC	COMMON STOCK	69553P100	4	102	SH		OTR	1	0	0	102
PALANTIR TECHNOLOGIES INC	COMMON STOCK	69608A108	463	19,675	SH		DFND	1	19,675	0	0
PALANTIR TECHNOLOGIES INC	COMMON STOCK	69608A108	71	3,000	SH		OTR	1	3,000	0	0
PALOMAR HOLDINGS INC	COMMON STOCK	69753M105	90	1,008	SH		DFND	1	0	0	1,008
PALOMAR HOLDINGS INC	COMMON STOCK	69753M105	59	661	SH		OTR	1	0	661	0
PAR PACIFIC HOLDINGS INC	COMMON STOCK	69888T207	12	841	SH		DFND	1	286	0	555
PARAMOUNT GROUP INC	COMMON STOCK	69924R108	21	2,310	SH		DFND	1	1,768	0	542
PARAMOUNT GROUP INC	COMMON STOCK	69924R108	1	156	SH		OTR	1	156	0	0
PARK AEROSPACE CORP	COMMON STOCK	70014A104	1	74	SH		DFND	1	74	0	0
PARSONS CORP	COMMON STOCK	70202L102	534	14,672	SH		DFND	1	14,629	0	43
PARSONS CORP	COMMON STOCK	70202L102	13	348	SH		OTR	1	348	0	0
PAYCOM SOFTWARE INC	COMMON STOCK	70432V102	9,922	21,938	SH		DFND	1	21,607	0	331
PAYCOM SOFTWARE INC	COMMON STOCK	70432V102	1,772	3,918	SH		OTR	1	3,481	27	410
PAYCOM SOFTWARE INC	COMMON STOCK	70432V102	503	1,112	SH		DFND	5	1,097	0	15
PAYLOCITY HOLDING CORP	COMMON STOCK	70438V106	3,134	15,222	SH		DFND	1	15,207	0	15
PAYLOCITY HOLDING CORP	COMMON STOCK	70438V106	393	1,908	SH		OTR	1	713	0	1,195
PAYLOCITY CORP	COMMON STOCK	70438V106	195	948	SH		DFND	5	948	0	0
PAYPAL HOLDINGS INC	COMMON STOCK	70450Y103	162,626	694,392	SH		DFND	1	671,841	0	22,551
PAYPAL HOLDINGS INC	COMMON STOCK	70450Y103	30,548	130,437	SH		OTR	1	118,028	3,463	8,946
PAYPAL HOLDINGS INC	COMMON STOCK	70450Y103	373	1,594	SH		DFND	5	1,495	0	99
PEBBLEBROOK HOTEL TRUST	COMMON STOCK	70509V100	396	21,069	SH		DFND	1	17,808	0	3,261
PEBBLEBROOK HOTEL TRUST	COMMON STOCK	70509V100	15	800	SH		OTR	1	329	309	162
PELOTON INTERACTIVE INC A	COMMON STOCK	70614W100	2,567	16,921	SH		DFND	1	16,785	0	136
PELOTON INTERACTIVE INC A	COMMON STOCK	70614W100	32	212	SH		OTR	1	200	0	12
PELOTON INTERACTIVE INC	COMMON STOCK	70614W100	31	206	SH		DFND	5	206	0	0
PENNANTPARK FLOATING RATE CA	COMMON STOCK	70806A106	42	4,000	SH		DFND	1	4,000	0	0
PENNYMAC MTG INV TR	COMMON STOCK	70931T103	40	2,247	SH		DFND	1	2,083	0	164
PENNYMAC FINANCIAL SERVICES	COMMON STOCK	70932M107	42	634	SH		DFND	1	634	0	0
PENNYMAC FINANCIAL SERVICES	COMMON STOCK	70932M107	46	707	SH		OTR	1	0	707	0
PENSKE AUTOMOTIVE GROUP INC	COMMON STOCK	70959W103	74	1,245	SH		DFND	1	646	0	599
PENSKE AUTOMOTIVE GROUP INC	COMMON STOCK	70959W103	21	346	SH		OTR	1	200	108	38
PENSKE AUTOMOTIVE GROUP INC	COMMON STOCK	70959W103	7	126	SH		DFND	5	126	0	0
PENUMBRA INC	COMMON STOCK	70975L107	914	5,223	SH		DFND	1	5,223	0	0
PENUMBRA INC	COMMON STOCK	70975L107	11	64	SH		OTR	1	18	39	7
PERDOCEO EDUCATION CORP	COMMON STOCK	71363P106	4	328	SH		DFND	1	328	0	0
PERFICIENT INC	COMMON STOCK	71375U101	84	1,759	SH		DFND	1	0	0	1,759
PERFORMANCE FOOD GROUP CO	COMMON STOCK	71377A103	2	43	SH		DFND	1	0	0	43
PERFORMANCE FOOD GROUP CO	COMMON STOCK	71377A103	101	2,125	SH		OTR	1	0	41	2,084
PERFORMANCE FOOD GROUP CO	COMMON STOCK	71377A103	28	590	SH		DFND	5	590	0	0
PERSHING SQUARE TONTINE HLDGS	COMMON STOCK	71531R109	663	23,921	SH		DFND	1	23,921	0	0
PERSHING SQUARE TONTINE HLDGS	COMMON STOCK	71531R109	70	2,536	SH		OTR	1	2,536	0	0
PERSHING SQUARE WT 7	OTHER	71531R117	0	50	SH		DFND	1	50	0	0
PERSHING SQUARE WT 7	OTHER	71531R117	1	156	SH		OTR	1	156	0	0
PETIQ INC	COMMON STOCK	71639T106	478	12,428	SH		DFND	1	8,984	0	3,444
PETIQ INC	COMMON STOCK	71639T106	75	1,938	SH		OTR	1	1,938	0	0
PETROLEO BRASILEIRO SPON A D R	COMMON STOCK	71654V101	20	1,770	SH		DFND	1	1,770	0	0
PETROLEO BRASILEIRO SPON A D R	COMMON STOCK	71654V101	25	2,237	SH		OTR	1	0	0	2,237
PETROLEO BRASILEIRO S A A D R	COMMON STOCK	71654V408	463	41,226	SH		DFND	1	26,786	0	14,440
PETROLEO BRASILEIRO S A A D R	COMMON STOCK	71654V408	35	3,151	SH		OTR	1	2,965	114	72
PETROBRAS PETROLEO BRASILEIRO	COMMON STOCK	71654V408	18	1,563	SH		DFND	5	1,563	0	0
PHIBRO ANIMAL HEALTH CORPORATI	COMMON STOCK	71742Q106	175	9,023	SH		DFND	1	9,023	0	0
PHIBRO ANIMAL HEALTH CORPORATI	COMMON STOCK	71742Q106	46	2,389	SH		OTR	1	2,389	0	0
PHYSICIANS REALTY TRUST	COMMON STOCK	71943U104	408	22,898	SH		DFND	1	22,502	0	396
PHYSICIANS REALTY TRUST	COMMON STOCK	71943U104	150	8,435	SH		OTR	1	8,435	0	0
PHYSICIANS REALTY TRUST	COMMON STOCK	71943U104	41	2,296	SH		DFND	5	2,296	0	0
PHREESIA INC	COMMON STOCK	71944F106	229	4,220	SH		DFND	1	4,220	0	0
PHREESIA INC	COMMON STOCK	71944F106	87	1,602	SH		OTR	1	1,602	0	0
PILGRIMS PRIDE CORP	COMMON STOCK	72147K108	43	2,215	SH		DFND	1	993	0	1,222
PILGRIMS PRIDE CORP	COMMON STOCK	72147K108	1	57	SH		OTR	1	0	0	57
P I M C O MUNICIPAL INCOME FUN	OTHER	72200R107	28	2,045	SH		OTR	1	2,045	0	0
PIMCO CORP AND INCOME STRATEGY	OTHER	72200U100	16	932	SH		DFND	1	932	0	0
PIMCO CORPORATE INCOME OPP FUN	OTHER	72201B101	105	5,900	SH		OTR	1	5,900	0	0
PIMCO FUNDS	OTHER	72201B101	914	51,223	SH		DFND	5	27,348	0	23,875
PIMCO INCOME STRATEGY FUND	OTHER	72201H108	26	2,314	SH		DFND	1	2,314	0	0
PIMCO INCOME STRATEGY FUND II	OTHER	72201J104	59	5,900	SH		DFND	1	3,900	0	2,000
PIMCO INCOME STRATEGY FUND II	OTHER	72201J104	144	14,500	SH		OTR	1	0	0	14,500
PIMCO 1 5 YEAR US TIPS INDEX E	OTHER	72201R205	85	1,578	SH		DFND	1	1,578	0	0
PIMCO 15 YEAR US TIPS INDEX ET	OTHER	72201R304	3	35	SH		DFND	1	35	0	0
PIMCO 15 YR US TIPS INDEX	OTHER	72201R304	22	251	SH		DFND	5	0	0	251
PIMCO ACTIVE BOND ETF	OTHER	72201R775	125	1,104	SH		DFND	1	844	0	260
PIMCO 0 5 YEAR HIGH YIELD CORP	OTHER	72201R783	13	129	SH		DFND	1	129	0	0
PIMCO 0-5 YR HI YLD CORP BOND	OTHER	72201R783	207	2,110	SH		DFND	5	2,033	0	77
PIMCO ENHANCED SHORT MATURITY	OTHER	72201R833	395	3,872	SH		DFND	1	3,872	0	0
PIMCO ENHANCED SHORT MATURITY	OTHER	72201R833	68	667	SH		OTR	1	0	300	367
PIMCO DYNAMIC INCOME FUND	OTHER	72201Y101	61	2,325	SH		DFND	1	2,325	0	0
PIMCO DYN CREDIT & MTGE INCOME	OTHER	72202D106	241	11,360	SH		DFND	1	11,360	0	0
PING IDENTITY HOLDING CORP	COMMON STOCK	72341T103	332	11,600	SH		DFND	1	11,600	0	0
PING IDENTITY HOLDING CORP	COMMON STOCK	72341T103	178	6,223	SH		OTR	1	3,979	2,244	0
PINNACLE FINANCIAL PARTNERS	COMMON STOCK	72346Q104	367	5,707	SH		DFND	1	1,813	0	3,894
PINNACLE FINANCIAL PARTNERS	COMMON STOCK	72346Q104	3	41	SH		OTR	1	0	0	41
PINNACLE FINANCIAL PARTNERS	COMMON STOCK	72346Q104	84	1,298	SH		DFND	5	1,298	0	0
PINTEREST INC CLASS A	COMMON STOCK	72352L106	302	4,577	SH		DFND	1	3,456	0	1,121
PINTEREST INC CLASS A	COMMON STOCK	72352L106	237	3,608	SH		OTR	1	3,406	185	17
PIXELWORKS INC	COMMON STOCK	72581M305	1	500	SH		DFND	1	500	0	0
PLAINS GP HOLDINGS L P CL A	COMMON STOCK	72651A207	885	104,776	SH		DFND	1	49,071	0	55,705
PLAINS GP HOLDINGS L P CL A	COMMON STOCK	72651A207	101	12,000	SH		OTR	1	0	12,000	0
PLAINS GP HOLDINGS LP	COMMON STOCK	72651A207	5	599	SH		DFND	5	599	0	0
PLANET FITNESS INC CL A	COMMON STOCK	72703H101	39	503	SH		DFND	1	281	0	222
PLANET FITNESS INC CL A	COMMON STOCK	72703H101	804	10,359	SH		OTR	1	0	10,321	38
PLUG POWER INC	COMMON STOCK	72919P202	1,280	37,750	SH		DFND	1	37,579	0	171
PLUG POWER INC	COMMON STOCK	72919P202	49	1,458	SH		OTR	1	1,415	0	43
PLURALSIGHT INC A	COMMON STOCK	72941B106	665	31,744	SH		DFND	1	31,657	0	87
PLURALSIGHT INC A	COMMON STOCK	72941B106	71	3,410	SH		OTR	1	3,244	0	166
PLURASIGHT INC CL A	COMMON STOCK	72941B106	162	7,749	SH		DFND	5	7,749	0	0
POOL CORPORATION	COMMON STOCK	73278L105	5,564	14,936	SH		DFND	1	14,137	0	799
POOL CORPORATION	COMMON STOCK	73278L105	842	2,262	SH		OTR	1	2,216	41	5
POOL CORP	COMMON STOCK	73278L105	131	351	SH		DFND	5	344	0	7
POSTAL REALTY TRUST INC A	COMMON STOCK	73757R102	94	5,592	SH		DFND	1	5,592	0	0
POSTAL REALTY TRUST INC A	COMMON STOCK	73757R102	57	3,400	SH		OTR	1	3,400	0	0
POSTAL REALTY TRUST INC	COMMON STOCK	73757R102	1	38	SH		DFND	5	38	0	0
PQ GROUP HOLDINGS INC	COMMON STOCK	73943T103	11	773	SH		DFND	1	0	0	773
PRECIGEN INC	COMMON STOCK	74017N105	9	862	SH		DFND	1	862	0	0
PRECISION BIOSCIENCES INC	COMMON STOCK	74019P108	22	2,669	SH		DFND	1	2,000	0	669
PREFERRED APATMENT COMMUN A	COMMON STOCK	74039L103	4	526	SH		DFND	1	526	0	0
PREDICTIVE ONCOLOGY INC	COMMON STOCK	74039M200	0	1	SH		DFND	1	1	0	0
PREMIER INC CLASS A	COMMON STOCK	74051N102	97	2,758	SH		DFND	1	1,647	0	1,111
PREMIER INC CLASS A	COMMON STOCK	74051N102	3	103	SH		OTR	1	10	0	93
PREMIER FINANCIAL CORP	COMMON STOCK	74052F108	3	138	SH		DFND	1	138	0	0
PRELUDE THERAPEUTICS INC COM	COMMON STOCK	74065P101	1	18	SH		DFND	1	18	0	0
PRESTIGE BRANDS HLDGS INC	COMMON STOCK	74112D101	385	11,035	SH		DFND	1	7,608	0	3,427
PRESTIGE BRANDS HLDGS INC	COMMON STOCK	74112D101	6	160	SH		OTR	1	160	0	0
T ROWE PRICE GROUP INC	COMMON STOCK	74144T108	9,478	62,609	SH		DFND	1	59,465	0	3,144
T ROWE PRICE GROUP INC	COMMON STOCK	74144T108	3,719	24,566	SH		OTR	1	23,619	833	114
T ROWE PRICE GROUP INC	COMMON STOCK	74144T108	56	372	SH		DFND	5	372	0	0
PRIMORIS SERVICES CORP	COMMON STOCK	74164F103	14	511	SH		DFND	1	383	0	128
PRIMERICA INC	COMMON STOCK	74164M108	14	106	SH		DFND	1	74	0	32
PRIMERICA INC	COMMON STOCK	74164M108	9	66	SH		OTR	1	0	53	13
PRIMO WATER CORP	COMMON STOCK	74167P108	21	1,321	SH		DFND	1	1,321	0	0
PRINCIPAL FINANCIAL GROUP INC	COMMON STOCK	74251V102	1,507	30,381	SH		DFND	1	25,618	0	4,763
PRINCIPAL FINANCIAL GROUP INC	COMMON STOCK	74251V102	859	17,309	SH		OTR	1	14,386	856	2,067
PRINCIPAL FINANCIAL GROUP	COMMON STOCK	74251V102	0	2	SH		DFND	5	2	0	0
PROASSURANCE CORPORATION	COMMON STOCK	74267C106	3	159	SH		DFND	1	159	0	0
PROCURE SPACE ETF	OTHER	74280R205	25	983	SH		DFND	1	983	0	0
PROG HOLDINGS INC COM	COMMON STOCK	74319R101	964	17,903	SH		DFND	1	17,550	0	353
PROG HOLDINGS INC COM	COMMON STOCK	74319R101	208	3,847	SH		OTR	1	485	3,310	52
PROG HOLDINGS INC	COMMON STOCK	74319R101	14	264	SH		DFND	5	225	0	39
PROLOGIS INC	COMMON STOCK	74340W103	12,922	129,668	SH		DFND	1	121,263	0	8,405
PROLOGIS INC	COMMON STOCK	74340W103	4,082	40,961	SH		OTR	1	31,033	342	9,586
PROLOGIS INC	COMMON STOCK	74340W103	149	1,498	SH		DFND	5	1,498	0	0
PROS HOLDINGS INC	COMMON STOCK	74346Y103	1,172	23,084	SH		DFND	1	23,076	0	8
PROS HOLDINGS INC	COMMON STOCK	74346Y103	289	5,700	SH		OTR	1	1,630	0	4,070
PROS HOLDINGS INC	COMMON STOCK	74346Y103	177	3,495	SH		DFND	5	3,495	0	0
PROSHARES TRUST ETF	OTHER	74347B201	82	5,000	SH		OTR	1	5,000	0	0
PROSHARES SHORT DOW30 ETF	OTHER	74347B235	0	6	SH		OTR	1	6	0	0
PROSHARES SHORT SP500 ETF	OTHER	74347B425	12	640	SH		DFND	1	640	0	0
PROSHARES SHORT SP500 ETF	OTHER	74347B425	9	490	SH		OTR	1	0	0	490
PROSHARES SHORT QQQ ETF	OTHER	74347B714	10	700	SH		DFND	1	700	0	0
PROPETRO HLDG CORP	COMMON STOCK	74347M108	12	1,587	SH		DFND	1	1,406	0	181
PROSHARES ULTRA SP500 ETF	OTHER	74347R107	195	2,140	SH		DFND	1	2,140	0	0
PROSHARES ULTRA SP500 ETF	OTHER	74347R107	14	158	SH		OTR	1	158	0	0
PROSHARES SHORT HIGH YIELD ETF	OTHER	74347R131	6	330	SH		DFND	1	330	0	0
ULTRA QQQ PROSHARES ETF	OTHER	74347R206	357	3,100	SH		DFND	1	3,100	0	0
PROSHARES ULTRA NASDAQ ETF	OTHER	74347R214	28	323	SH		DFND	1	323	0	0
PROSHARES ULTRA NASDAQ ETF	OTHER	74347R214	15	177	SH		OTR	1	177	0	0
PROSHARES LARGE CAP CORE PLUS	OTHER	74347R248	35	420	SH		DFND	1	0	0	420
PROSHARES ULTRASHORT 710 YEAR	OTHER	74347R313	1	40	SH		OTR	1	40	0	0
PROSHARES ULTRA SILVER ETF	OTHER	74347W353	845	16,485	SH		DFND	1	16,485	0	0
PROSHARES ULTRA FINANCIALS ETF	OTHER	74347X633	25	600	SH		DFND	1	600	0	0
PROSHARES ULTRAPRO RUSSELL2000	OTHER	74347X799	0	5	SH		DFND	1	5	0	0
PROSHARES SHORT 20 YEAR TREASU	OTHER	74347X849	19	1,175	SH		OTR	1	50	0	1,125
PROSHARES ULTRA BLOOMBERG CRUD	OTHER	74347Y888	13	360	SH		OTR	1	360	0	0
PROSHARES PET CARE ETF	OTHER	74348A145	36	500	SH		DFND	1	500	0	0
PROSHARES SHORT RUSSELL2000 ET	OTHER	74348A210	9	355	SH		DFND	1	355	0	0
PROSHARES SHORT RUSSELL2000 ET	OTHER	74348A210	13	500	SH		OTR	1	0	0	500
PROSHARES S P 500 DIVIDEND ETF	OTHER	74348A467	3,247	40,643	SH		DFND	1	40,643	0	0
PROSHARES S&P 500 ARISTOCRATS	OTHER	74348A467	54	675	SH		DFND	5	0	0	675
PROSPECT CAPITAL CORP	COMMON STOCK	74348T102	994	183,642	SH		DFND	1	180,837	0	2,805
PROSPECT CAPITAL CORP	COMMON STOCK	74348T102	1,302	240,600	SH		OTR	1	240,600	0	0
PROVENTION BIO INC	COMMON STOCK	74374N102	51	3,000	SH		DFND	1	3,000	0	0
PROVIDENT FINANCIAL SERVICES	COMMON STOCK	74386T105	2	135	SH		DFND	1	0	0	135
PRUDENTIAL P L C A D R	COMMON STOCK	74435K204	444	12,030	SH		DFND	1	7,181	0	4,849
PRUDENTIAL P L C A D R	COMMON STOCK	74435K204	47	1,253	SH		OTR	1	460	750	43
PUBLIC STORAGE INC	COMMON STOCK	74460D109	7,349	31,823	SH		DFND	1	28,948	0	2,875
PUBLIC STORAGE INC	COMMON STOCK	74460D109	2,157	9,338	SH		OTR	1	5,254	1,784	2,300
PUBLIC STORAGE INC	COMMON STOCK	74460D109	7	29	SH		DFND	5	29	0	0
PUMA BIOTECHNOLOGY INC	COMMON STOCK	74587V107	2	200	SH		DFND	1	200	0	0
PUMA BIOTECHNOLOGY INC	COMMON STOCK	74587V107	3	310	SH		OTR	1	0	310	0
PURE STORAGE INC CLASS A	COMMON STOCK	74624M102	446	19,712	SH		DFND	1	19,712	0	0
PURE STORAGE INC CLASS A	COMMON STOCK	74624M102	153	6,766	SH		OTR	1	6,766	0	0
PURPLE INNOVATION INC	COMMON STOCK	74640Y106	61	1,850	SH		DFND	1	1,850	0	0
PURPLE INNOVATION INC	COMMON STOCK	74640Y106	2	50	SH		OTR	1	50	0	0
QCR HOLDINGS INC	COMMON STOCK	74727A104	114	2,881	SH		DFND	1	2,881	0	0
QAD INC A	COMMON STOCK	74727D306	13	205	SH		DFND	1	25	0	180
QIWI PLCSPONSORED A D R	COMMON STOCK	74735M108	18	1,739	SH		DFND	1	1,739	0	0
QTS REALTY TRUST INC CL A	COMMON STOCK	74736A103	1,862	30,085	SH		DFND	1	26,950	0	3,135
QTS REALTY TRUST INC CL A	COMMON STOCK	74736A103	163	2,630	SH		OTR	1	861	1,769	0
QTS REALTY TRUST INC	COMMON STOCK	74736A103	11	180	SH		DFND	5	180	0	0
QORVO INC	COMMON STOCK	74736K101	859	5,169	SH		DFND	1	4,741	0	428
QORVO INC	COMMON STOCK	74736K101	228	1,373	SH		OTR	1	696	644	33
QORVO INC	COMMON STOCK	74736K101	51	309	SH		DFND	5	309	0	0
Q2 HOLDINGS INC	COMMON STOCK	74736L109	1,480	11,697	SH		DFND	1	9,872	0	1,825
Q2 HOLDINGS INC	COMMON STOCK	74736L109	170	1,342	SH		OTR	1	272	0	1,070
QUALIGEN THERAPEUTICS INC COM	COMMON STOCK	74754R103	2	804	SH		DFND	1	804	0	0
QUALIGEN THERAPEUTICS INC COM	COMMON STOCK	74754R103	27	9,047	SH		OTR	1	9,047	0	0
QUALYS INC	COMMON STOCK	74758T303	4,215	34,585	SH		DFND	1	34,571	0	14
QUALYS INC	COMMON STOCK	74758T303	1,130	9,268	SH		OTR	1	7,669	0	1,599
QUALYS INC	COMMON STOCK	74758T303	47	384	SH		DFND	5	366	0	18
QUANTERIX CORP	COMMON STOCK	74766Q101	667	14,339	SH		DFND	1	14,302	0	37
QUANTERIX CORP	COMMON STOCK	74766Q101	10	210	SH		OTR	1	210	0	0
QUANTUMSCAPE CORP	COMMON STOCK	74767V109	104	1,235	SH		DFND	1	1,235	0	0
QUEST DIAGNOSTICS INC	COMMON STOCK	74834L100	1,619	13,582	SH		DFND	1	12,935	0	647
QUEST DIAGNOSTICS INC	COMMON STOCK	74834L100	335	2,814	SH		OTR	1	2,279	123	412
QUEST DIAGNOSTICS INC	COMMON STOCK	74834L100	65	548	SH		DFND	5	531	0	17
QUIDEL CORP	COMMON STOCK	74838J101	60	334	SH		DFND	1	242	0	92
QUIDEL CORP	COMMON STOCK	74838J101	7	37	SH		OTR	1	0	37	0
QUINSTREET INC	COMMON STOCK	74874Q100	3	119	SH		DFND	1	0	0	119
QUTOUTIAO INC A D R	COMMON STOCK	74915J107	18	11,565	SH		DFND	1	11,565	0	0
QURATE RETAIL INC SERIES A	COMMON STOCK	74915M100	72	6,534	SH		DFND	1	3,697	0	2,837
QURATE RETAIL INC SERIES A	COMMON STOCK	74915M100	1	121	SH		OTR	1	0	0	121
MOTLEY FOOL 100 INDEX ETF	OTHER	74933W601	26	738	SH		DFND	1	738	0	0
MFAM SMALL CAP GROWTH ETF	OTHER	74933W874	16	413	SH		DFND	1	413	0	0
RLJ LODGING TRUST	COMMON STOCK	74965L101	27	1,882	SH		DFND	1	1,367	0	515
RLJ LODGING TRUST	COMMON STOCK	74965L101	10	679	SH		OTR	1	0	679	0
RMR GROUP INC THE A	COMMON STOCK	74967R106	14	363	SH		DFND	1	240	0	123
RMR GROUP INC THE A	COMMON STOCK	74967R106	1	36	SH		OTR	1	0	36	0
RH	COMMON STOCK	74967X103	117	261	SH		DFND	1	238	0	23
RH	COMMON STOCK	74967X103	1	3	SH		OTR	1	0	0	3
RPT REALTY	COMMON STOCK	74971D101	1	161	SH		DFND	1	161	0	0
RANGE RESOURCES CORP	COMMON STOCK	75281A109	114	17,109	SH		DFND	1	8,019	0	9,090
RANGE RESOURCES CORP	COMMON STOCK	75281A109	8	1,089	SH		OTR	1	400	0	689
RAYONIER ADVANCED MATERIALS	COMMON STOCK	75508B104	113	17,300	SH		DFND	1	17,300	0	0
RBC BEARINGS INC	COMMON STOCK	75524B104	1,015	5,669	SH		DFND	1	4,486	0	1,183
RBC BEARINGS INC	COMMON STOCK	75524B104	272	1,522	SH		OTR	1	81	266	1,175
RBC BEARINGS INC	COMMON STOCK	75524B104	78	438	SH		DFND	5	438	0	0
RE MAX HOLDINGS INC CL A	COMMON STOCK	75524W108	2	57	SH		DFND	1	41	0	16
READY CAPITAL CORP	COMMON STOCK	75574U101	5	436	SH		DFND	1	161	0	275
REALOGY HOLDINGS CORP	COMMON STOCK	75605Y106	43	3,287	SH		DFND	1	380	0	2,907
REALPAGE INC	COMMON STOCK	75606N109	1,103	12,642	SH		DFND	1	12,284	0	358
REALPAGE INC	COMMON STOCK	75606N109	879	10,079	SH		OTR	1	8,087	0	1,992
REATA PHARMACEUTICALS INC	COMMON STOCK	75615P103	127	1,028	SH		DFND	1	827	0	201
RED ROBIN GOURMET BURGERS	COMMON STOCK	75689M101	5	245	SH		DFND	1	245	0	0
RED ROCK RESORTS INC	COMMON STOCK	75700L108	6	256	SH		DFND	1	256	0	0
REDFIN CORP	COMMON STOCK	75737F108	10	141	SH		DFND	1	141	0	0
REDFIN CORP	COMMON STOCK	75737F108	2	23	SH		OTR	1	0	0	23
REGENERON PHARMACEUTICALS INC	COMMON STOCK	75886F107	1,857	3,844	SH		DFND	1	3,088	0	756
REGENERON PHARMACEUTICALS INC	COMMON STOCK	75886F107	735	1,521	SH		OTR	1	730	67	724
REGENRON PHARMACEUTICALS	COMMON STOCK	75886F107	13	27	SH		DFND	5	27	0	0
REGENXBIO INC	COMMON STOCK	75901B107	8	185	SH		DFND	1	0	0	185
REGIONS FINL CORP	COMMON STOCK	7591EP100	2,015	124,990	SH		DFND	1	113,979	0	11,011
REGIONS FINL CORP	COMMON STOCK	7591EP100	1,041	64,565	SH		OTR	1	62,277	994	1,294
REGIONS FINANCIAL CORP	COMMON STOCK	7591EP100	185	11,499	SH		DFND	5	11,499	0	0
RENEWABLE ENERGY GROUP INC	COMMON STOCK	75972A301	1,151	16,249	SH		DFND	1	13,845	0	2,404
RENEWABLE ENERGY GROUP INC	COMMON STOCK	75972A301	30	429	SH		OTR	1	429	0	0
RENT A CTR INC	COMMON STOCK	76009N100	48	1,246	SH		DFND	1	302	0	944
REPLIMUNE GROUP INC	COMMON STOCK	76029N106	9	245	SH		DFND	1	169	0	76
RESIDEO TECHNOLOGIES INC	COMMON STOCK	76118Y104	63	2,998	SH		DFND	1	2,654	0	344
RESIDEO TECHNOLOGIES INC	COMMON STOCK	76118Y104	152	7,168	SH		OTR	1	6,969	172	27
RESIDEO TECHNOLOGIES INC	COMMON STOCK	76118Y104	0	10	SH		DFND	5	10	0	0
RESOURCES CONNECTION INC	COMMON STOCK	76122Q105	3	231	SH		DFND	1	88	0	143
RETRACTABLE TECHNOLOGIES INC	COMMON STOCK	76129W105	1	100	SH		DFND	1	100	0	0
RESTAURANT BRANDS INTERN	COMMON STOCK	76131D103	608	9,954	SH		DFND	1	8,940	0	1,014
RESTAURANT BRANDS INTERN	COMMON STOCK	76131D103	180	2,943	SH		OTR	1	2,872	16	55
RETAIL OPPORTUNITY INVESTMENTS	COMMON STOCK	76131N101	27	2,007	SH		DFND	1	1,738	0	269
RETAIL OPPORTUNITY INVESTMENTS	COMMON STOCK	76131N101	1	73	SH		OTR	1	0	73	0
RETAIL PROPERTIES OF AME A	COMMON STOCK	76131V202	128	14,949	SH		DFND	1	11,266	0	3,683
RETAIL PROPERTIES OF AME A	COMMON STOCK	76131V202	9	1,063	SH		OTR	1	1,063	0	0
REVOLVE GROUP INC	COMMON STOCK	76156B107	9	300	SH		DFND	1	300	0	0
REVOLVE GROUP INC	COMMON STOCK	76156B107	540	17,320	SH		OTR	1	17,000	0	320
REXNORD CORP	COMMON STOCK	76169B102	1,663	42,102	SH		DFND	1	42,102	0	0
REXNORD CORP	COMMON STOCK	76169B102	147	3,725	SH		OTR	1	3,174	204	347
REXNORD CORP	COMMON STOCK	76169B102	0	6	SH		DFND	5	6	0	0
REXFORD INDUSTRIAL REALTY IN	COMMON STOCK	76169C100	1,592	32,411	SH		DFND	1	32,384	0	27
REXFORD INDUSTRIAL REALTY IN	COMMON STOCK	76169C100	57	1,165	SH		OTR	1	1,165	0	0
REYNOLDS CONSUMER PRODS I	COMMON STOCK	76171L106	943	31,378	SH		DFND	1	31,360	0	18
REYNOLDS CONSUMER PRODS I	COMMON STOCK	76171L106	311	10,365	SH		OTR	1	10,330	0	35
REYNOLDS CONSUMER PRODUCTS INC	COMMON STOCK	76171L106	2	62	SH		DFND	5	62	0	0
RHYTHM PHARMACEUTICALS INC	COMMON STOCK	76243J105	15	494	SH		DFND	1	46	0	448
RHYTHM PHARMACEUTICALS INC	COMMON STOCK	76243J105	10	335	SH		OTR	1	335	0	0
RINGCENTRAL INC CLASS A	COMMON STOCK	76680R206	74,653	196,988	SH		DFND	1	195,202	0	1,786
RINGCENTRAL INC CLASS A	COMMON STOCK	76680R206	8,419	22,214	SH		OTR	1	21,258	578	378
RINGCENTRAL INC	COMMON STOCK	76680R206	53	141	SH		DFND	5	129	0	12
RMR MORTGAGE TRUST	OTHER	76970B101	2	200	SH		DFND	1	200	0	0
ROCKET COMPANIES INC	COMMON STOCK	77311W101	2	83	SH		DFND	5	83	0	0
ROCKET PHARMACEUTICALS INC	COMMON STOCK	77313F106	14	255	SH		DFND	1	164	0	91
ROCKY MOUNTAIN CHOC FACT INC	COMMON STOCK	77467X101	2	442	SH		DFND	1	442	0	0
ROKU INC	COMMON STOCK	77543R102	1,745	5,256	SH		DFND	1	4,886	0	370
ROKU INC	COMMON STOCK	77543R102	388	1,168	SH		OTR	1	335	63	770
ROKU INC	COMMON STOCK	77543R102	5	15	SH		DFND	5	15	0	0
ROYCE GLOBAL VALUE TRUST	OTHER	78081T104	7	498	SH		DFND	1	0	0	498
ROYCE GLOBAL VALUE TRUST	OTHER	78081T104	20	1,496	SH		OTR	1	0	1,496	0
RUBIUS THERAPEUTICS INC	COMMON STOCK	78116T103	10	1,286	SH		DFND	1	0	0	1,286
RYMAN HOSPITALITY PROPERTIES	COMMON STOCK	78377T107	84	1,236	SH		DFND	1	1,202	0	34
RYMAN HOSPITALITY PROPERTIES	COMMON STOCK	78377T107	1	18	SH		OTR	1	0	0	18
S P GLOBAL INC	COMMON STOCK	78409V104	65,613	199,594	SH		DFND	1	195,468	0	4,126
S P GLOBAL INC	COMMON STOCK	78409V104	12,341	37,538	SH		OTR	1	30,847	3,020	3,671
S&P GLOBAL INC	COMMON STOCK	78409V104	220	669	SH		DFND	5	659	0	10
SBA COMMUNICATIONS CORP	COMMON STOCK	78410G104	6,228	22,075	SH		DFND	1	21,166	0	909
SBA COMMUNICATIONS CORP	COMMON STOCK	78410G104	1,129	4,000	SH		OTR	1	3,294	140	566
SBA COMMUNICATIONS CORP	COMMON STOCK	78410G104	234	831	SH		DFND	5	777	0	54
SEACOR MARINE HOLDINGS INC	COMMON STOCK	78413P101	6	2,292	SH		OTR	1	0	2,292	0
SK TELECOM CO LTD A D R	COMMON STOCK	78440P108	534	21,824	SH		DFND	1	10,504	0	11,320
SK TELECOM CO LTD A D R	COMMON STOCK	78440P108	126	5,142	SH		OTR	1	0	108	5,034
SL GREEN RLTY CORP	COMMON STOCK	78440X101	320	5,366	SH		DFND	1	4,313	0	1,053
SL GREEN RLTY CORP	COMMON STOCK	78440X101	98	1,653	SH		OTR	1	1,653	0	0
SLM CORP	COMMON STOCK	78442P106	40	3,283	SH		DFND	1	3,126	0	157
SLM CORP	COMMON STOCK	78442P106	2	206	SH		OTR	1	0	0	206
SM ENERGY CO	COMMON STOCK	78454L100	274	44,810	SH		DFND	1	44,810	0	0
SM ENERGY CO	COMMON STOCK	78454L100	27	4,400	SH		OTR	1	4,400	0	0
SPDR S P 500 ETF	OTHER	78462F103	1,496,941	4,003,800	SH		DFND	1	3,986,924	0	16,876
SPDR S P 500 ETF	OTHER	78462F103	495,216	1,324,533	SH		OTR	1	1,296,722	7,602	20,209
SPDR S&P 500 ETF TRUST	OTHER	78462F103	24,032	64,276	SH		DFND	5	50,256	0	14,020
SPDR S P 500 ETF	OTHER	78462F103	157	16	SH	Call	DFND	1	16	0	0
SPDR S P 500 ETF	OTHER	78462F103	755	1,570	SH	Put	DFND	1	1,570	0	0
SPDR S P 500 ETF	OTHER	78462F103	196	20	SH	Call	OTR	1	20	0	0
SPDR S P 500 ETF	OTHER	78462F103	261	870	SH	Put	OTR	1	820	0	50
SPS COMMERCE INC	COMMON STOCK	78463M107	3,546	32,660	SH		DFND	1	32,348	0	312
SPS COMMERCE INC	COMMON STOCK	78463M107	662	6,093	SH		OTR	1	2,801	497	2,795
SPS COMMERCE INC	COMMON STOCK	78463M107	7	63	SH		DFND	5	63	0	0
SPDR GOLD SHARES ETF	OTHER	78463V107	109,611	614,547	SH		DFND	1	610,397	0	4,150
SPDR GOLD SHARES ETF	OTHER	78463V107	14,206	79,646	SH		OTR	1	76,065	1,186	2,395
SPDR GOLD TRUST	OTHER	78463V107	3,051	17,108	SH		DFND	5	730	0	16,378
SPDR EURO STOXX 50 ETF	OTHER	78463X202	93	2,225	SH		OTR	1	2,225	0	0
SPDR S P EMERGING ASIA PACIFIC	OTHER	78463X301	72	569	SH		DFND	1	569	0	0
SPDR S P EMERGING ASIA PACIFIC	OTHER	78463X301	12	95	SH		OTR	1	0	40	55
SPDR S&P EMERGING ASIA PACIFIC	OTHER	78463X301	22	173	SH		DFND	5	173	0	0
SPDR EURO STOXX SMALL CAP	OTHER	78463X392	2	26	SH		DFND	5	26	0	0
SPDR S P CHINA ETF	OTHER	78463X400	79	605	SH		OTR	1	605	0	0
SPDR PORTFOLIO EMERGING MARKET	OTHER	78463X509	48	1,132	SH		DFND	1	1,132	0	0
SPDR S&P EMERGING MARKETS	OTHER	78463X509	18	438	SH		DFND	5	122	0	316
SPDR DOW JONES GLOBAL REAL ETF	OTHER	78463X749	224	5,044	SH		DFND	1	5,044	0	0
SPDR DJ WHILSHRE GLBL REAL EST	OTHER	78463X749	1,933	43,471	SH		DFND	5	0	0	43,471
SPDR S P EMERGING MARKETS SMAL	OTHER	78463X756	13	260	SH		DFND	1	44	0	216
SPDR S P INTERNATIONAL DIVIDEN	OTHER	78463X772	84	2,294	SH		DFND	1	300	0	1,994
SPDR MSCI ACWI EX US ETF	OTHER	78463X848	18	637	SH		DFND	1	637	0	0
SPDR DOW JONES INTERNATIONAL E	OTHER	78463X863	8,095	237,471	SH		DFND	1	236,354	0	1,117
SPDR DOW JONES INTERNATIONAL E	OTHER	78463X863	1,915	56,186	SH		OTR	1	54,595	316	1,275
SPDR S P INTERNATIONAL SMALL C	OTHER	78463X871	41	1,150	SH		DFND	1	1,150	0	0
SPDR S&P INTL SMALL CAP	OTHER	78463X871	2	57	SH		DFND	5	57	0	0
SPDR PORTFOLIO DEVELOPED WLD E	OTHER	78463X889	4,660	138,079	SH		DFND	1	138,079	0	0
SPDR PORTFOLIO DEVELOPED WLD E	OTHER	78463X889	361	10,694	SH		OTR	1	10,694	0	0
SPDR S&P WORLD (EX-US)	OTHER	78463X889	14	400	SH		DFND	5	400	0	0
SPDR NYSE TECHNOLOGY ETF	OTHER	78464A102	79	559	SH		DFND	1	559	0	0
SPDR S&P 1500 VALUE TILT	OTHER	78464A128	62	513	SH		DFND	5	0	0	513
SPDR BARCLAYS CAP INTL CORP BD	OTHER	78464A151	1	35	SH		DFND	5	0	0	35
SPDR S P 600 SMALL CAP GROWTH	OTHER	78464A201	11	140	SH		DFND	1	140	0	0
SPDR S&P 600 SMALL CAP GROWTH	OTHER	78464A201	785	10,278	SH		DFND	5	9,984	0	294
SPDR WELLS FARGO PREFERRED STO	OTHER	78464A292	113	2,545	SH		DFND	1	2,545	0	0
SPDR WELLS FARGO PREFERRED STO	OTHER	78464A292	98	2,200	SH		OTR	1	2,200	0	0
SPDR WELLS FARGO PFD STOCK	OTHER	78464A292	15	341	SH		DFND	5	341	0	0
SPDR S&P 600 SMALL CAP VALUE E	OTHER	78464A300	99	1,490	SH		DFND	1	1,390	0	100
SPDR S&P 600 SMALL CAP VALUE	OTHER	78464A300	2	27	SH		DFND	5	0	0	27
SPDR BLMBG BARCLAYS CONVERT SE	OTHER	78464A359	1,144	13,824	SH		DFND	1	13,824	0	0
SPDR BLMBG BARCLAYS CONVERT SE	OTHER	78464A359	68	825	SH		OTR	1	825	0	0
SPDR BARCLAYS CAP CONVERTIBLE	OTHER	78464A359	149	1,798	SH		DFND	5	0	0	1,798
SPDR SERIES TRUST	OTHER	78464A367	26	800	SH		DFND	1	800	0	0
SPDR PORTFOLIO INTERMEDIATE TE	OTHER	78464A375	37	982	SH		DFND	1	49	0	933
SPDR PORTFOLIO INTERMEDIATE TE	OTHER	78464A375	81	2,179	SH		OTR	1	0	0	2,179
SPDR BARCLAYS CAP INTER CREDIT	OTHER	78464A375	694	18,672	SH		DFND	5	18,672	0	0
SPDR PORTFOLIO SP 500 GROWTH E	OTHER	78464A409	1,486	26,882	SH		DFND	1	26,882	0	0
SPDR S&P 500 GROWTH	OTHER	78464A409	624	11,279	SH		DFND	5	8,872	0	2,407
SPDR PORTFOLIO SHORT TERM ETF	OTHER	78464A474	3,038	96,750	SH		DFND	1	91,078	0	5,672
SPDR FTSE INTERNATIONAL ETF	OTHER	78464A490	252	4,284	SH		DFND	1	4,284	0	0
SPDR PORTFOLIO S P 500 VALUE E	OTHER	78464A508	125	3,635	SH		DFND	1	3,635	0	0
SPDR S&P 500 VALUE	OTHER	78464A508	5	145	SH		DFND	5	145	0	0
SPDR BLMBG BARCLAYS INTL TRS B	OTHER	78464A516	81	2,604	SH		DFND	1	2,604	0	0
SPDR BLMBG BARCLAYS INTL TRS B	OTHER	78464A516	140	4,498	SH		OTR	1	4,498	0	0
SPDR BARCLAYS INTL TREASURY BD	OTHER	78464A516	79	2,521	SH		DFND	5	623	0	1,898
SPDR S&P TRANSPORTATION	OTHER	78464A532	51	710	SH		DFND	5	710	0	0
SPDR S&P HEALTH CARE EQUIPMENT	OTHER	78464A581	92	805	SH		DFND	1	740	0	65
SPDR DOW JONES REIT ETF	OTHER	78464A607	46	533	SH		DFND	1	200	0	333
SPDR DOW JONES REIT ETF	OTHER	78464A607	711	8,206	SH		OTR	1	8,206	0	0
SPDR S&P AEROSPACE DEFENSE ETF	OTHER	78464A631	33	289	SH		DFND	1	289	0	0
SPDR PORTOLIO AGGREGATE BOND E	OTHER	78464A649	37	1,192	SH		DFND	1	1,192	0	0
SPDR BARCLAYS CAPITAL TIPS	OTHER	78464A656	30	965	SH		DFND	5	965	0	0
SPDR S&P REGIONAL BANKING ETF	OTHER	78464A698	13	254	SH		OTR	1	254	0	0
SPDR S&P RETAIL	OTHER	78464A714	32	498	SH		DFND	5	498	0	0
SPDR S&P PHARMACEUTICALS ETF	OTHER	78464A722	29	556	SH		DFND	1	556	0	0
SPDR S&P METALS AND MINING ETF	OTHER	78464A755	67	2,000	SH		DFND	1	2,000	0	0
SPDR S&P METALS AND MINING ETF	OTHER	78464A755	34	1,030	SH		OTR	1	1,000	0	30
SPDR METALS & MINING	OTHER	78464A755	31	921	SH		DFND	5	921	0	0
SPDR S&P DIVIDEND ETF	OTHER	78464A763	1,967	18,569	SH		DFND	1	18,496	0	73
SPDR S&P DIVIDEND ETF	OTHER	78464A763	4,595	43,382	SH		OTR	1	42,102	0	1,280
SPDR S&P DIVIDEND	OTHER	78464A763	527	4,973	SH		DFND	5	0	0	4,973
SPDR S&P CAPITAL MARKETS ETF	OTHER	78464A771	1	17	SH		DFND	1	17	0	0
SPDR S&P INSURANCE ETF	OTHER	78464A789	41	1,215	SH		DFND	1	1,215	0	0
SPDR S&P INSURANCE ETF	OTHER	78464A789	36	1,080	SH		OTR	1	1,080	0	0
SPDR S P BANK ETF	OTHER	78464A797	4	95	SH		DFND	1	95	0	0
SPDR DOW JONES TOTAL MARKET	OTHER	78464A805	8	173	SH		DFND	5	173	0	0
SPDR S&P 600 SMALL CAP ETF	OTHER	78464A813	74	930	SH		DFND	1	930	0	0
SPDR S&P 400 MID CAP GROWTH ET	OTHER	78464A821	11	153	SH		DFND	1	153	0	0
SPDR S&P 400 MID CAP GROWTH	OTHER	78464A821	422	6,082	SH		DFND	5	6,082	0	0
SPDR S&P 400 MID CAP VALUE	OTHER	78464A839	321	5,788	SH		DFND	5	5,788	0	0
SPDR PORTFOLIO S P 400 MID CAP	OTHER	78464A847	49	1,217	SH		DFND	1	1,217	0	0
SPDR PORTFOLIO S P 400 MID CAP	OTHER	78464A847	174	4,303	SH		OTR	1	4,303	0	0
SPDR PORTIFOLIO S P 500 ETF	OTHER	78464A854	33	753	SH		DFND	1	740	0	13
SPDR PORTIFOLIO S P 500 ETF	OTHER	78464A854	88	2,000	SH		OTR	1	2,000	0	0
SPDR S&P SEMICONDUCTOR ETF	OTHER	78464A862	2	14	SH		DFND	1	14	0	0
SPDR S&P BIOTECH ETF	OTHER	78464A870	181	1,287	SH		DFND	1	1,287	0	0
SPDR S&P BIOTECH ETF	OTHER	78464A870	198	1,409	SH		OTR	1	209	0	1,200
SS C TECHNOLOGIES HOLDINGS	COMMON STOCK	78467J100	1,247	17,131	SH		DFND	1	16,806	0	325
SS C TECHNOLOGIES HOLDINGS	COMMON STOCK	78467J100	160	2,210	SH		OTR	1	1,124	1,046	40
SS&C TECHNOLOGIES HOLDINGS	COMMON STOCK	78467J100	51	702	SH		DFND	5	702	0	0
SPDR BLACKSTONE GSO SR LOAN	OTHER	78467V608	291	6,377	SH		DFND	5	6,377	0	0
SPDR DOW JONES INDUSTRIAL ETF	OTHER	78467X109	2,881	9,423	SH		DFND	1	9,423	0	0
SPDR DOW JONES INDUSTRIAL ETF	OTHER	78467X109	890	2,911	SH		OTR	1	2,911	0	0
SPDR S P MIDCAP 400 ETF	OTHER	78467Y107	447,749	1,066,271	SH		DFND	1	1,057,363	0	8,908
SPDR S P MIDCAP 400 ETF	OTHER	78467Y107	77,932	185,589	SH		OTR	1	181,850	1,571	2,168
SPDR S&P MIDCAP 400 ETF TRUST	OTHER	78467Y107	2,401	5,717	SH		DFND	5	10	0	5,707
SPDR S P MIDCAP 400 ETF	OTHER	78467Y107	29	13	SH	Put	DFND	1	13	0	0
SPDR BLOOMBERG BARCLAYS SHORT	OTHER	78468R101	5	165	SH		DFND	1	165	0	0
SPDR BARCLAYS CAP INVESTMENT	OTHER	78468R200	0	10	SH		DFND	5	10	0	0
SPDR BARCLAYS CAPITAL SHORT	OTHER	78468R408	191	7,095	SH		DFND	5	7,095	0	0
SPDR S P OIL GAS EXPLOR PRODTN	OTHER	78468R556	117	1,996	SH		DFND	1	1,996	0	0
SPDR S P OIL GAS EXPLOR PRODTN	OTHER	78468R556	7	125	SH		OTR	1	125	0	0
SPDR PORTFOLIO HIGH YIELD BOND	OTHER	78468R606	3	100	SH		DFND	1	100	0	0
SPDR BLOOMBERG BARCLAYS HIGH E	OTHER	78468R622	16,064	147,453	SH		DFND	1	147,453	0	0
SPDR BLOOMBERG BARCLAYS HIGH E	OTHER	78468R622	210	1,929	SH		OTR	1	1,929	0	0
BLOOMBERG BARCLAYS HIGH YLD BD	OTHER	78468R622	539	4,946	SH		DFND	5	4,946	0	0
SPDR KENSHO CLEAN POWER ETF	OTHER	78468R655	76	700	SH		DFND	1	700	0	0
SPDR BLOOMBERG BARCLAYS 1 3 ET	OTHER	78468R663	3,966	43,336	SH		DFND	1	43,336	0	0
SPDR BLOOMBERG BARCLAYS 1 3 ET	OTHER	78468R663	19	213	SH		OTR	1	63	0	150
SPDR BLOOMBERG BARCLAYS 1-3	OTHER	78468R663	6,390	69,825	SH		DFND	5	0	0	69,825
SPDR S&P 1500 MOMENTUM TILT ET	OTHER	78468R705	86	533	SH		DFND	1	533	0	0
SPDR NUVEEN BLOOMBERG BARCLAYS	OTHER	78468R721	14	276	SH		DFND	1	276	0	0
SPDR NUVEEN BARCLAYS MUNI BOND	OTHER	78468R721	26	493	SH		DFND	5	0	0	493
SPDR NUVEEN BLOOMBERG BARCLAYS	OTHER	78468R739	5	95	SH		DFND	1	95	0	0
SPDR NUVEEN BLOOMBERG BARCLAYS	OTHER	78468R739	16	312	SH		OTR	1	312	0	0
SPDR NUVEEN BARCLAYS SHORT	OTHER	78468R739	8	152	SH		DFND	5	0	0	152
SPDR S&P 500 HIGH DIVDEND	OTHER	78468R788	10	297	SH		DFND	5	297	0	0
SPDR S P 500 FOSSIL FUEL ETF	OTHER	78468R796	49	525	SH		DFND	1	525	0	0
SPDR SSGA US LARGE CAP LOW ETF	OTHER	78468R804	15	123	SH		DFND	1	123	0	0
SPDR MSCI USA STRATEGICFACTORS	OTHER	78468R812	5	52	SH		DFND	1	52	0	0
SP PLUS CORP	COMMON STOCK	78469C103	2	72	SH		OTR	1	0	34	38
SPX FLOW INC W I	COMMON STOCK	78469X107	98	1,686	SH		DFND	1	578	0	1,108
SPX FLOW INC W I	COMMON STOCK	78469X107	242	4,180	SH		OTR	1	4,180	0	0
SVB FINL GROUP	COMMON STOCK	78486Q101	1,755	4,526	SH		DFND	1	4,325	0	201
SVB FINL GROUP	COMMON STOCK	78486Q101	61	158	SH		OTR	1	101	7	50
SVB FINANCIAL GROUP	COMMON STOCK	78486Q101	5	14	SH		DFND	5	14	0	0
SVMK INC	COMMON STOCK	78489X103	2	71	SH		DFND	1	0	0	71
SABRA HEALTH CARE REIT INC	COMMON STOCK	78573L106	54	3,085	SH		DFND	1	2,910	0	175
SABRA HEALTH CARE REIT INC	COMMON STOCK	78573L106	32	1,864	SH		OTR	1	1,800	0	64
SABRE CORP	COMMON STOCK	78573M104	63	5,255	SH		DFND	1	4,967	0	288
SABRE CORP	COMMON STOCK	78573M104	11	875	SH		OTR	1	0	0	875
SABRE CORP	COMMON STOCK	78573M104	1	96	SH		DFND	5	96	0	0
SAFEHOLD INC	COMMON STOCK	78645L100	657	9,061	SH		DFND	1	9,061	0	0
SAFEHOLD INC	COMMON STOCK	78645L100	153	2,116	SH		OTR	1	2,116	0	0
SAFEHOLD INC	COMMON STOCK	78645L100	1	13	SH		DFND	5	13	0	0
SAFETY INSURANCE GROUP INC	COMMON STOCK	78648T100	144	1,841	SH		DFND	1	1,841	0	0
SAGE THERAPEUTICS INC	COMMON STOCK	78667J108	123	1,427	SH		DFND	1	808	0	619
SAGE THERAPEUTICS INC	COMMON STOCK	78667J108	144	1,668	SH		OTR	1	1,519	68	81
SAIA INC	COMMON STOCK	78709Y105	1,084	5,997	SH		DFND	1	5,782	0	215
SAIA INC	COMMON STOCK	78709Y105	139	769	SH		OTR	1	769	0	0
SAILPOINT TECHNOLOGIES HOLDI	COMMON STOCK	78781P105	754	14,163	SH		DFND	1	14,105	0	58
SAILPOINT TECHNOLOGIES HOLDI	COMMON STOCK	78781P105	287	5,388	SH		OTR	1	5,388	0	0
SALESFORCE COM INC	COMMON STOCK	79466L302	67,828	304,806	SH		DFND	1	290,919	0	13,887
SALESFORCE COM INC	COMMON STOCK	79466L302	18,319	82,318	SH		OTR	1	73,634	3,413	5,271
SALESFORCE.COM INC	COMMON STOCK	79466L302	705	3,169	SH		DFND	5	3,169	0	0
SANDSTORM GOLD LTD	COMMON STOCK	80013R206	41	5,740	SH		DFND	1	5,740	0	0
SANOFI A D R	COMMON STOCK	80105N105	1,706	35,096	SH		DFND	1	23,357	0	11,739
SANOFI A D R	COMMON STOCK	80105N105	656	13,488	SH		OTR	1	12,051	805	632
SANOFI AVENTIS	COMMON STOCK	80105N105	32	652	SH		DFND	5	652	0	0
SANTANDER CONSUMER USA HOLDI	COMMON STOCK	80283M101	125	5,713	SH		DFND	1	4,235	0	1,478
SANTANDER CONSUMER USA HOLDI	COMMON STOCK	80283M101	3	144	SH		OTR	1	0	0	144
SCHNEIDER NATIONAL INC CL B	COMMON STOCK	80689H102	2,379	114,931	SH		DFND	1	114,243	0	688
SCHNEIDER NATIONAL INC CL B	COMMON STOCK	80689H102	40	1,904	SH		OTR	1	103	351	1,450
SCHNEIDER NATIONAL INC	COMMON STOCK	80689H102	25	1,186	SH		DFND	5	1,186	0	0
SCHOLAR ROCK HOLDING CORP	COMMON STOCK	80706P103	16	326	SH		DFND	1	65	0	261
SCHRODINGER INC	COMMON STOCK	80810D103	62	780	SH		DFND	1	780	0	0
SCIENTIFIC GAMES CORPORATION C	COMMON STOCK	80874P109	9	213	SH		DFND	1	152	0	61
SCIENTIFIC GAMES CORPORATION C	COMMON STOCK	80874P109	8	196	SH		OTR	1	0	0	196
SCIENTIFIC GAMES CORP	COMMON STOCK	80874P109	0	2	SH		DFND	5	2	0	0
SEA LTD ADR	COMMON STOCK	81141R100	1,845	9,272	SH		DFND	1	9,057	0	215
SEA LTD ADR	COMMON STOCK	81141R100	454	2,283	SH		OTR	1	2,238	45	0
SEA LTD	COMMON STOCK	81141R100	13	64	SH		DFND	5	64	0	0
SEAGEN INC	COMMON STOCK	81181C104	2,233	12,749	SH		DFND	1	11,413	0	1,336
SEAGEN INC	COMMON STOCK	81181C104	161	917	SH		OTR	1	655	161	101
SEAGEN INC	COMMON STOCK	81181C104	10	58	SH		DFND	5	58	0	0
SEALED AIR CORP	COMMON STOCK	81211K100	1,233	26,917	SH		DFND	1	24,733	0	2,184
SEALED AIR CORP	COMMON STOCK	81211K100	656	14,325	SH		OTR	1	13,916	400	9
SEALED AIR CORP	COMMON STOCK	81211K100	24	529	SH		DFND	5	529	0	0
SEASPINE HOLDINGS CORP	COMMON STOCK	81255T108	20	1,138	SH		DFND	1	1,138	0	0
SEAWORLD ENTERTAINMENT INC	COMMON STOCK	81282V100	3	83	SH		DFND	1	0	0	83
SEAWORLD ENTERTAINMENT INC	COMMON STOCK	81282V100	148	4,681	SH		OTR	1	0	0	4,681
SEAWORLD ENTERTAINMENT INC	COMMON STOCK	81282V100	6	196	SH		DFND	5	196	0	0
SECOND SIGHT MED PRODS INC	COMMON STOCK	81362J209	2	1,000	SH		DFND	1	1,000	0	0
MATERIALS SELECT SECTOR SPDR E	OTHER	81369Y100	4,585	63,336	SH		DFND	1	62,872	0	464
MATERIALS SELECT SECTOR SPDR E	OTHER	81369Y100	3,618	49,975	SH		OTR	1	49,330	645	0
SPDR MATERIALS SELECT SECTOR	OTHER	81369Y100	1	15	SH		DFND	5	15	0	0
HEALTH CARE SELECT SECTOR SPDR	OTHER	81369Y209	52,769	465,171	SH		DFND	1	464,044	0	1,127
HEALTH CARE SELECT SECTOR SPDR	OTHER	81369Y209	17,132	151,016	SH		OTR	1	143,383	7,433	200
SPDR HEALTH CARE SELECT SECTOR	OTHER	81369Y209	171	1,510	SH		DFND	5	1,044	0	466
CONSUMER STAPLES SELECT SECTOR	OTHER	81369Y308	17,439	258,540	SH		DFND	1	256,667	0	1,873
CONSUMER STAPLES SELECT SECTOR	OTHER	81369Y308	5,093	75,510	SH		OTR	1	63,953	11,020	537
CONSUMER STAPLES SELECT SECTOR	OTHER	81369Y308	54	806	SH		DFND	5	338	0	468
CONSUMER DISCRETIONARY SELECT	OTHER	81369Y407	31,163	193,826	SH		DFND	1	192,350	0	1,476
CONSUMER DISCRETIONARY SELECT	OTHER	81369Y407	22,555	140,284	SH		OTR	1	126,419	7,825	6,040
CONSUMER DISCRETIONARY SELECT	OTHER	81369Y407	2,386	14,842	SH		DFND	5	16	0	14,826
ENERGY SELECT SECTOR SPDR FUND	OTHER	81369Y506	14,549	383,890	SH		DFND	1	383,121	0	769
ENERGY SELECT SECTOR SPDR FUND	OTHER	81369Y506	3,589	94,707	SH		OTR	1	90,953	200	3,554
ENERGY SELECT SECTOR SPDR	OTHER	81369Y506	9	225	SH		DFND	5	0	0	225
ENERGY SELECT SECTOR SPDR FUND ETF	OTHER	81369Y506	101	318	SH	Call	DFND	1	318	0	0
ENERGY SELECT SECTOR SPDR FUND ETF	OTHER	81369Y506	114	318	SH	Put	DFND	1	318	0	0
FINANCIAL SELECT SECTOR SPDR E	OTHER	81369Y605	25,112	851,830	SH		DFND	1	846,723	0	5,107
FINANCIAL SELECT SECTOR SPDR E	OTHER	81369Y605	12,234	415,020	SH		OTR	1	412,415	1,000	1,605
SPDR FINANCIAL SELECT SECTOR	OTHER	81369Y605	488	16,555	SH		DFND	5	0	0	16,555
INDUSTRIAL SELECT SECTOR SPDR	OTHER	81369Y704	15,460	174,582	SH		DFND	1	171,850	0	2,732
INDUSTRIAL SELECT SECTOR SPDR	OTHER	81369Y704	4,536	51,223	SH		OTR	1	51,066	157	0
INDUSTRIAL SELECT SECTOR SPDR	OTHER	81369Y704	66	748	SH		DFND	5	748	0	0
TECHNOLOGY SELECT SECTOR SPDR	OTHER	81369Y803	153,755	1,182,556	SH		DFND	1	1,170,785	0	11,771
TECHNOLOGY SELECT SECTOR SPDR	OTHER	81369Y803	61,343	471,800	SH		OTR	1	466,352	5,338	110
SPDR TECHNOLOGY SELECT SECTOR	OTHER	81369Y803	7,041	54,150	SH		DFND	5	3,414	0	50,736
COMMUNICATION SERVICES SELECT	OTHER	81369Y852	6,497	96,274	SH		DFND	1	95,914	0	360
COMMUNICATION SERVICES SELECT	OTHER	81369Y852	411	6,091	SH		OTR	1	6,091	0	0
COMMUNICATION SERVICES SELECT	OTHER	81369Y852	40	594	SH		DFND	5	594	0	0
THE REAL ESTATE SELECT SECTOR	OTHER	81369Y860	812	22,222	SH		DFND	1	22,222	0	0
THE REAL ESTATE SELECT SECTOR	OTHER	81369Y860	315	8,617	SH		OTR	1	8,617	0	0
UTILITIES SELECT SECTOR SPDR E	OTHER	81369Y886	25,331	403,996	SH		DFND	1	403,321	0	675
UTILITIES SELECT SECTOR SPDR E	OTHER	81369Y886	10,513	167,679	SH		OTR	1	160,971	6,308	400
SPDR UTILITIES SELECT SECTOR	OTHER	81369Y886	184	2,930	SH		DFND	5	191	0	2,739
SECUREWORKS CORP A	COMMON STOCK	81374A105	324	22,754	SH		DFND	1	22,754	0	0
SECUREWORKS CORP A	COMMON STOCK	81374A105	94	6,570	SH		OTR	1	6,570	0	0
SELECT ENERGY SERVICES INC A	COMMON STOCK	81617J301	15	3,694	SH		DFND	1	0	0	3,694
SELECT MEDICAL HLDGS CORP	COMMON STOCK	81619Q105	16	574	SH		DFND	1	574	0	0
SELECT MEDICAL HLDGS CORP	COMMON STOCK	81619Q105	2	80	SH		OTR	1	0	38	42
SENSIENT TECHNOLOGIES CORP	COMMON STOCK	81725T100	484	6,552	SH		DFND	1	5,594	0	958
SENSEONICS HLDGS INC	COMMON STOCK	81727U105	17	20,000	SH		DFND	1	20,000	0	0
SERITAGE GROWTH PROP A REIT	COMMON STOCK	81752R100	29	1,988	SH		DFND	1	1,395	0	593
SERITAGE GROWTH PROP A REIT	COMMON STOCK	81752R100	1	76	SH		OTR	1	0	0	76
SERVICE PROPERTIES TRUST	COMMON STOCK	81761L102	191	16,580	SH		DFND	1	13,794	0	2,786
SERVICE PROPERTIES TRUST	COMMON STOCK	81761L102	24	2,100	SH		OTR	1	500	681	919
SERVICENOW INC	COMMON STOCK	81762P102	75,950	137,983	SH		DFND	1	134,462	0	3,521
SERVICENOW INC	COMMON STOCK	81762P102	19,685	35,764	SH		OTR	1	30,971	3,602	1,191
SERVICENOW INC	COMMON STOCK	81762P102	598	1,087	SH		DFND	5	1,059	0	28
SEVERN BANCORP INC	COMMON STOCK	81811M100	14	2,000	SH		OTR	1	2,000	0	0
SHAW COMMUNICATIONS INC CL B	COMMON STOCK	82028K200	91	5,216	SH		DFND	1	2,857	0	2,359
SHAW COMMUNICATIONS INC CL B	COMMON STOCK	82028K200	42	2,386	SH		OTR	1	2,112	0	274
SHENANDOAH TELECOMMUN COM	COMMON STOCK	82312B106	640	14,798	SH		DFND	1	14,704	0	94
SHIFT TECHNOLOGIES INC	COMMON STOCK	82452T107	4	500	SH		DFND	1	500	0	0
SHOCKWAVE MEDICAL INC	COMMON STOCK	82489T104	76	728	SH		DFND	1	706	0	22
SHOCKWAVE MEDICAL INC	COMMON STOCK	82489T104	65	628	SH		OTR	1	33	0	595
SHOPIFY INC A	COMMON STOCK	82509L107	13,954	12,327	SH		DFND	1	12,237	0	90
SHOPIFY INC A	COMMON STOCK	82509L107	872	770	SH		OTR	1	745	0	25
SHOPIFY INC	COMMON STOCK	82509L107	1,341	1,185	SH		DFND	5	1,170	0	15
SHOTSPOTTER INC	COMMON STOCK	82536T107	12	329	SH		DFND	1	329	0	0
SIBANYE STILLWATER LTD ADR A D	COMMON STOCK	82575P107	30	1,923	SH		DFND	1	1,645	0	278
SIBANYE STILLWATER LTD ADR A D	COMMON STOCK	82575P107	117	7,334	SH		OTR	1	7,334	0	0
SIERRA BANCORP	COMMON STOCK	82620P102	10	432	SH		DFND	1	432	0	0
SIGNATURE BK	COMMON STOCK	82669G104	218	1,607	SH		DFND	1	1,029	0	578
SIGNATURE BK	COMMON STOCK	82669G104	12	87	SH		OTR	1	87	0	0
SIGNATURE BANK	COMMON STOCK	82669G104	100	740	SH		DFND	5	740	0	0
SILICON MOTION TECHNOLOGY CORP	COMMON STOCK	82706C108	6,008	124,785	SH		DFND	1	124,388	0	397
SILICON MOTION TECHNOLOGY CORP	COMMON STOCK	82706C108	1,557	32,344	SH		OTR	1	32,344	0	0
SILICON MOTION TECHNOLOGY CORP	COMMON STOCK	82706C108	20	424	SH		DFND	5	424	0	0
SILK ROAD MEDICAL INC	COMMON STOCK	82710M100	20	310	SH		DFND	1	284	0	26
SILK ROAD MEDICAL INC	COMMON STOCK	82710M100	1	15	SH		OTR	1	15	0	0
SILVERCORP METALS INC	COMMON STOCK	82835P103	10	1,563	SH		DFND	1	0	0	1,563
SIMPLY GOOD FOODS COMPANY	COMMON STOCK	82900L102	69	2,186	SH		OTR	1	0	2,186	0
SIRIUS XM HOLDINGS	COMMON STOCK	82968B103	364	57,133	SH		DFND	1	53,790	0	3,343
SIRIUS XM HOLDINGS	COMMON STOCK	82968B103	329	51,636	SH		OTR	1	7,256	0	44,380
SIRIUS XM HOLDINGS INC	COMMON STOCK	82968B103	0	0	SH		DFND	5	0	0	0
SITE CENTERS CORP	COMMON STOCK	82981J109	28	2,727	SH		DFND	1	1,577	0	1,150
SITE CTRS CORP	COMMON STOCK	82981J109	1	105	SH		DFND	5	105	0	0
SITEONE LANDSCAPE SUPPLY INC	COMMON STOCK	82982L103	1,459	9,199	SH		DFND	1	7,880	0	1,319
SITEONE LANDSCAPE SUPPLY INC	COMMON STOCK	82982L103	310	1,955	SH		OTR	1	210	0	1,745
SIX FLAGS ENTERTAINMENT	COMMON STOCK	83001A102	88	2,584	SH		DFND	1	1,205	0	1,379
SIX FLAGS ENTERTAINMENT	COMMON STOCK	83001A102	29	849	SH		OTR	1	288	366	195
SIX FLAGS ENTERTAINMENT CORP	COMMON STOCK	83001A102	5	155	SH		DFND	5	155	0	0
SIXTH STREET SPECIALTY LENDING	COMMON STOCK	83012A109	710	34,226	SH		DFND	1	34,120	0	106
SKILLZ INC	COMMON STOCK	83067L109	40	2,000	SH		DFND	1	2,000	0	0
SKYWORKS SOLUTIONS INC	COMMON STOCK	83088M102	14,184	92,777	SH		DFND	1	90,388	0	2,389
SKYWORKS SOLUTIONS INC	COMMON STOCK	83088M102	2,993	19,582	SH		OTR	1	17,019	282	2,281
SKYWORKS SOLUTIONS INC	COMMON STOCK	83088M102	105	690	SH		DFND	5	674	0	16
SLACK TECHNOLOGIES INC CL A	COMMON STOCK	83088V102	154	3,651	SH		DFND	1	2,971	0	680
SLACK TECHNOLOGIES INC CL A	COMMON STOCK	83088V102	88	2,090	SH		OTR	1	2,090	0	0
SLEEP NUMBER CORP	COMMON STOCK	83125X103	4,994	61,001	SH		DFND	1	61,001	0	0
SMITH NEPHEW PLC SPON A D R	COMMON STOCK	83175M205	2,291	54,337	SH		DFND	1	52,308	0	2,029
SMITH NEPHEW PLC SPON A D R	COMMON STOCK	83175M205	379	9,004	SH		OTR	1	8,213	731	60
SMITH & NEPHEW PLC	COMMON STOCK	83175M205	241	5,714	SH		DFND	5	5,714	0	0
SMARTSHEET INC CLASS A	COMMON STOCK	83200N103	36	521	SH		DFND	1	502	0	19
SMARTSHEET INC CLASS A	COMMON STOCK	83200N103	199	2,870	SH		OTR	1	2,175	307	388
SNAP INC	COMMON STOCK	83304A106	1,470	29,365	SH		DFND	1	28,169	0	1,196
SNAP INC	COMMON STOCK	83304A106	346	6,901	SH		OTR	1	6,901	0	0
SNAP INC	COMMON STOCK	83304A106	29	584	SH		DFND	5	521	0	63
SOGOU INC A D R	COMMON STOCK	83409V104	11	1,393	SH		DFND	1	620	0	773
SOHU COM LTD A D R	COMMON STOCK	83410S108	133	8,371	SH		DFND	1	7,473	0	898
SOHU COM LTD A D R	COMMON STOCK	83410S108	40	2,499	SH		OTR	1	2,499	0	0
SOHU.COM LTD	COMMON STOCK	83410S108	0	4	SH		DFND	5	4	0	0
SOLAR CAPITAL LTD	COMMON STOCK	83413U100	9	500	SH		OTR	1	500	0	0
SOLAREDGE TECHNOLOGIES INC	COMMON STOCK	83417M104	1,345	4,215	SH		DFND	1	1,612	0	2,603
SOLAREDGE TECHNOLOGIES INC	COMMON STOCK	83417M104	229	718	SH		OTR	1	676	38	4
SOLAREDGE TECHNOLOGIES INC	COMMON STOCK	83417M104	9	29	SH		DFND	5	25	0	4
SOLARWINDS CORP	COMMON STOCK	83417Q105	19	1,281	SH		DFND	1	1,197	0	84
SONIC AUTOMOTIVE INC CLASS A	COMMON STOCK	83545G102	36	945	SH		DFND	1	146	0	799
SONOMA PHARMACEUTICALS INC	COMMON STOCK	83558L204	4	500	SH		DFND	1	500	0	0
SONOS INC	COMMON STOCK	83570H108	16	689	SH		DFND	1	100	0	589
SONOS INC	COMMON STOCK	83570H108	10	447	SH		DFND	5	447	0	0
SORRENTO THERAPEUTICS INC COM	COMMON STOCK	83587F202	79	11,627	SH		DFND	1	10,654	0	973
SORRENTO THERAPEUTICS INC COM	COMMON STOCK	83587F202	41	6,000	SH		OTR	1	6,000	0	0
SOUTHERN COPPER CORP DEL	COMMON STOCK	84265V105	272	4,172	SH		DFND	1	2,547	0	1,625
SOUTHERN COPPER CORP DEL	COMMON STOCK	84265V105	61	939	SH		OTR	1	878	0	61
SOUTHSIDE BANCSHARES INC	COMMON STOCK	84470P109	136	4,393	SH		DFND	1	3,316	0	1,077
SPECIAL OPPORTUNITIES FD INC	OTHER	84741T104	3	231	SH		DFND	1	231	0	0
SPECTRUM BRANDS HOLDINGS INC	COMMON STOCK	84790A105	578	7,328	SH		DFND	1	6,125	0	1,203
SPECTRUM BRANDS HOLDINGS INC	COMMON STOCK	84790A105	11	135	SH		OTR	1	135	0	0
SPERO THERAPEUTICS INC	COMMON STOCK	84833T103	58	3,000	SH		DFND	1	3,000	0	0
SPIRE INC	COMMON STOCK	84857L101	805	12,568	SH		DFND	1	9,390	0	3,178
SPIRE INC	COMMON STOCK	84857L101	1,033	16,125	SH		OTR	1	14,051	574	1,500
SPIRIT REALTY CAPTIAL INC	COMMON STOCK	84860W300	254	6,321	SH		DFND	1	3,177	0	3,144
SPIRIT REALTY CAPTIAL INC	COMMON STOCK	84860W300	55	1,380	SH		OTR	1	1,000	0	380
SPIRIT REALTY CAPITAL INC	COMMON STOCK	84860W300	4	110	SH		DFND	5	110	0	0
SPORTSMANS WAREHOUSE HOLDIN	COMMON STOCK	84920Y106	1	36	SH		DFND	1	0	0	36
SPROTT PHYSICAL GOLD TRUST	OTHER	85207H104	20	1,335	SH		DFND	1	1,335	0	0
SPROTT PHYSICAL GOLD TRUST	OTHER	85207H104	45	3,000	SH		OTR	1	3,000	0	0
SPROTT PHYSICAL SILVER TRUST	OTHER	85207K107	711	76,160	SH		DFND	1	76,160	0	0
SPROUTS FARMERS MARKETS INC	COMMON STOCK	85208M102	90	4,450	SH		DFND	1	3,085	0	1,365
SPROUTS FARMERS MARKETS INC	COMMON STOCK	85208M102	77	3,806	SH		OTR	1	2,962	422	422
SPROTT PHYSICAL GOLD SILVER TR	OTHER	85208R101	699	36,119	SH		DFND	1	36,119	0	0
SPROTT PHYSICAL GOLD SILVER TR	OTHER	85208R101	65	3,367	SH		OTR	1	3,367	0	0
SPROUT SOCIAL INC CLASS A	COMMON STOCK	85209W109	891	19,628	SH		DFND	1	19,628	0	0
SPROUT SOCIAL INC CLASS A	COMMON STOCK	85209W109	13	281	SH		OTR	1	281	0	0
SPROUT SOCIAL INC	COMMON STOCK	85209W109	3	67	SH		DFND	5	67	0	0
STABLE ROAD ACQUISITION CL A	COMMON STOCK	85236Q109	9	500	SH		DFND	1	500	0	0
STAG INDUSTRIAL INC	COMMON STOCK	85254J102	777	24,804	SH		DFND	1	24,228	0	576
STAG INDUSTRIAL INC	COMMON STOCK	85254J102	283	9,038	SH		OTR	1	9,038	0	0
STANTEC INC	COMMON STOCK	85472N109	3	107	SH		DFND	1	0	0	107
STAR GROUP L P	COMMON STOCK	85512C105	85	9,000	SH		DFND	1	9,000	0	0
STARWOOD PROPERTY TRUST INC	COMMON STOCK	85571B105	664	34,376	SH		DFND	1	31,508	0	2,868
STARWOOD PROPERTY TRUST INC	COMMON STOCK	85571B105	3	148	SH		OTR	1	0	0	148
STARWOOD PROPERTY TRUST INC	COMMON STOCK	85571B105	1	64	SH		DFND	5	64	0	0
STERLING BANCORP	COMMON STOCK	85917A100	369	20,509	SH		DFND	1	8,040	0	12,469
STERLING BANCORP	COMMON STOCK	85917A100	19	1,109	SH		OTR	1	694	59	356
STONERIDGE INC	COMMON STOCK	86183P102	2	64	SH		DFND	1	64	0	0
STONERIDGE INC	COMMON STOCK	86183P102	67	2,216	SH		OTR	1	0	2,216	0
STRATEGIC EDUCATION INC	COMMON STOCK	86272C103	6	61	SH		DFND	1	61	0	0
STRIDE INC	COMMON STOCK	86333M108	1	53	SH		DFND	1	53	0	0
SUMITOMO MITSUI FINL GROUP A D	COMMON STOCK	86562M209	7,107	1,152,016	SH		DFND	1	1,058,963	0	93,053
SUMITOMO MITSUI FINL GROUP A D	COMMON STOCK	86562M209	1,680	272,449	SH		OTR	1	246,192	10,372	15,885
SUMITOMO MITSUI	COMMON STOCK	86562M209	0	0	SH		DFND	5	0	0	0
SUMMIT MATERIALS INC CL A	COMMON STOCK	86614U100	23	1,160	SH		DFND	1	1,089	0	71
SUMMIT MATERIALS INC CL A	COMMON STOCK	86614U100	55	2,729	SH		OTR	1	0	2,729	0
SUMMIT THERAPEUTICS INC	COMMON STOCK	86627T108	1	216	SH		DFND	1	216	0	0
SUNNOVA ENERGY INTERNATIONAL	COMMON STOCK	86745K104	59	1,300	SH		OTR	1	1,300	0	0
SUNOCO L P	COMMON STOCK	86765K109	14	500	SH		DFND	1	500	0	0
SUNRUN INC	COMMON STOCK	86771W105	46	664	SH		DFND	1	114	0	550
SUNRUN INC	COMMON STOCK	86771W105	57	829	SH		OTR	1	650	0	179
SUPER MICRO COMPUTER INC	COMMON STOCK	86800U104	13	400	SH		OTR	1	400	0	0
SURFACE ONCOLOGY INC	COMMON STOCK	86877M209	7	798	SH		DFND	1	0	0	798
SURGERY PARTNERS INC	COMMON STOCK	86881A100	6	214	SH		DFND	1	45	0	169
SURGERY PARTNERS INC	COMMON STOCK	86881A100	1	44	SH		OTR	1	0	0	44
SUZANO SA A D R	COMMON STOCK	86959K105	93	8,343	SH		DFND	1	2,389	0	5,954
SUZANO SA A D R	COMMON STOCK	86959K105	1	110	SH		OTR	1	0	99	11
SWITCH INC A	COMMON STOCK	87105L104	66,012	4,032,521	SH		DFND	1	4,032,173	0	348
SWITCH INC A	COMMON STOCK	87105L104	36	2,190	SH		OTR	1	1,995	0	195
SWITCHBACK ENERGY ACQUISIT A	COMMON STOCK	87105M102	1	25	SH		DFND	1	25	0	0
SYNAPTICS INC	COMMON STOCK	87157D109	604	6,270	SH		DFND	1	6,134	0	136
SYNAPTICS INC	COMMON STOCK	87157D109	46	471	SH		OTR	1	161	310	0
SYNOVUS FINANCIAL CORP	COMMON STOCK	87161C501	562	17,371	SH		DFND	1	15,059	0	2,312
SYNOVUS FINANCIAL CORP	COMMON STOCK	87161C501	79	2,436	SH		OTR	1	1,748	311	377
SYNNEX CORP	COMMON STOCK	87162W100	330	4,058	SH		DFND	1	3,710	0	348
SYNNEX CORP	COMMON STOCK	87162W100	210	2,577	SH		OTR	1	2,310	252	15
SYNDAX PHARMACEUTICALS INC	COMMON STOCK	87164F105	96	4,330	SH		OTR	1	4,330	0	0
SYNCHRONY FINANCIAL	COMMON STOCK	87165B103	407	11,729	SH		DFND	1	6,668	0	5,061
SYNCHRONY FINANCIAL	COMMON STOCK	87165B103	367	10,568	SH		OTR	1	2,949	31	7,588
SYNEOS HEALTH INC	COMMON STOCK	87166B102	1,531	22,461	SH		DFND	1	22,178	0	283
SYNEOS HEALTH INC	COMMON STOCK	87166B102	378	5,556	SH		OTR	1	5,463	0	93
SYNEOS HEALTH INC	COMMON STOCK	87166B102	86	1,268	SH		DFND	5	1,268	0	0
SYROS PHARMACEUTICALS INC	COMMON STOCK	87184Q107	8	728	SH		DFND	1	570	0	158
TC PIPELINES LP	COMMON STOCK	87233Q108	15	500	SH		DFND	1	500	0	0
TFS FINL CORP	COMMON STOCK	87240R107	40	2,255	SH		DFND	1	1,103	0	1,152
TFS FINL CORP	COMMON STOCK	87240R107	3	160	SH		OTR	1	0	0	160
TFI INTERNATIONAL INC	COMMON STOCK	87241L109	1	20	SH		DFND	1	20	0	0
TFI INTERNATIONAL INC	COMMON STOCK	87241L109	57	1,099	SH		OTR	1	0	1,099	0
TRI POINTE GROUP INC	COMMON STOCK	87265H109	651	37,742	SH		DFND	1	36,139	0	1,603
TRI POINTE GROUP INC	COMMON STOCK	87265H109	75	4,368	SH		OTR	1	4,224	0	144
TPI COMPOSITES INC	COMMON STOCK	87266J104	1,099	20,817	SH		DFND	1	15,168	0	5,649
TPI COMPOSITES INC	COMMON STOCK	87266J104	30	576	SH		OTR	1	576	0	0
TPG RE FINANCE TRUST INC	COMMON STOCK	87266M107	88	8,304	SH		DFND	1	1,258	0	7,046
TTM TECHNOLOGIES	COMMON STOCK	87305R109	351	25,491	SH		DFND	1	25,292	0	199
TTM TECHNOLOGIES	COMMON STOCK	87305R109	7	473	SH		OTR	1	473	0	0
TACTILE SYSTEMS TECHNOLOGY INC	COMMON STOCK	87357P100	185	4,109	SH		DFND	1	362	0	3,747
TACTILE SYSTEMS TECHNOLOGY INC	COMMON STOCK	87357P100	1	18	SH		OTR	1	18	0	0
TALOS ENERGY INC	COMMON STOCK	87484T108	11	1,332	SH		DFND	1	1,053	0	279
TARGA RESOURCES CORP	COMMON STOCK	87612G101	287	10,871	SH		DFND	1	7,080	0	3,791
TARGA RESOURCES CORP	COMMON STOCK	87612G101	70	2,639	SH		OTR	1	1,970	45	624
TARGA RESOURCES CORP	COMMON STOCK	87612G101	8	305	SH		DFND	5	305	0	0
TAYLOR MORRISON HOME CORP A	COMMON STOCK	87724P106	105	4,084	SH		DFND	1	2,425	0	1,659
TAYLOR MORRISON HOME CORP A	COMMON STOCK	87724P106	3	130	SH		OTR	1	0	0	130
TC ENERGY CORP	COMMON STOCK	87807B107	14,727	361,663	SH		DFND	1	357,045	0	4,618
TC ENERGY CORP	COMMON STOCK	87807B107	4,286	105,235	SH		OTR	1	95,577	365	9,293
TC ENERGY CORP	COMMON STOCK	87807B107	160	3,922	SH		DFND	5	3,922	0	0
TCR2 THERAPEUTICS INC	COMMON STOCK	87808K106	636	20,549	SH		DFND	1	20,549	0	0
TCR2 THERAPEUTICS INC	COMMON STOCK	87808K106	27	866	SH		OTR	1	866	0	0
TECHTARGET INC	COMMON STOCK	87874R100	9	156	SH		DFND	1	0	0	156
TEGNA INC	COMMON STOCK	87901J105	155	11,140	SH		DFND	1	1,995	0	9,145
TEGNA INC	COMMON STOCK	87901J105	4	310	SH		OTR	1	310	0	0
TELADOC INC	COMMON STOCK	87918A105	42,321	211,649	SH		DFND	1	210,398	0	1,251
TELADOC INC	COMMON STOCK	87918A105	6,887	34,442	SH		OTR	1	29,659	3,918	865
TELADOC INC	COMMON STOCK	87918A105	294	1,472	SH		DFND	5	1,472	0	0
TELEFONICA BRASIL SA A D R	COMMON STOCK	87936R205	93	10,493	SH		DFND	1	4,565	0	5,928
TELEFONICA BRASIL SA	COMMON STOCK	87936R205	2	183	SH		DFND	5	183	0	0
TELOS CORP MD	COMMON STOCK	87969B101	11	322	SH		OTR	1	0	322	0
TELUS CORP	COMMON STOCK	87971M103	113	5,722	SH		DFND	1	5,722	0	0
TELUS CORP	COMMON STOCK	87971M103	0	19	SH		DFND	5	19	0	0
TEMPLETON DRAGON FUND INC	OTHER	88018T101	23	987	SH		OTR	1	987	0	0
TEMPUR SEALY INTERNATIONAL INC	COMMON STOCK	88023U101	9	340	SH		DFND	1	220	0	120
TEMPUR SEALY INTERNATIONAL INC	COMMON STOCK	88023U101	1	44	SH		OTR	1	0	0	44
TENABLE HOLDINGS INC	COMMON STOCK	88025T102	1,072	20,520	SH		DFND	1	20,506	0	14
TENABLE HOLDINGS INC	COMMON STOCK	88025T102	504	9,642	SH		OTR	1	8,489	1,153	0
10X GENOMICS INC CLASS A	COMMON STOCK	88025U109	28	200	SH		DFND	1	200	0	0
TENARIS SA A D R	COMMON STOCK	88031M109	133	8,327	SH		DFND	1	2,054	0	6,273
TENARIS SA A D R	COMMON STOCK	88031M109	17	1,051	SH		OTR	1	339	634	78
TENET HEALTHCARE CORP	COMMON STOCK	88033G407	42	1,045	SH		DFND	1	486	0	559
TENET HEALTHCARE CORP	COMMON STOCK	88033G407	6	159	SH		OTR	1	0	0	159
TENCENT MUSIC ENTERTAINM A D R	COMMON STOCK	88034P109	586	30,440	SH		DFND	1	25,122	0	5,318
TENCENT MUSIC ENTERTAINM A D R	COMMON STOCK	88034P109	71	3,717	SH		OTR	1	2,856	0	861
TERADATA CORP	COMMON STOCK	88076W103	50	2,241	SH		DFND	1	2,209	0	32
TERADATA CORP	COMMON STOCK	88076W103	21	946	SH		OTR	1	249	0	697
TERRITORIAL BANCORP INC	COMMON STOCK	88145X108	281	11,700	SH		DFND	1	11,700	0	0
TERRENO REALTY CORP	COMMON STOCK	88146M101	2,489	42,543	SH		DFND	1	42,478	0	65
TERRENO REALTY CORP	COMMON STOCK	88146M101	443	7,575	SH		OTR	1	6,441	1,134	0
TERRENO REALTY CORP	COMMON STOCK	88146M101	2	32	SH		DFND	5	32	0	0
TESLA INC	COMMON STOCK	88160R101	33,881	48,013	SH		DFND	1	42,733	0	5,280
TESLA INC	COMMON STOCK	88160R101	11,281	15,986	SH		OTR	1	14,900	671	415
TESLA MOTORS INC	COMMON STOCK	88160R101	104	147	SH		DFND	5	147	0	0
TESLA INC	COMMON STOCK	88160R101	160	5	SH	Call	DFND	1	5	0	0
TETRA TECH INC	COMMON STOCK	88162G103	444	3,832	SH		DFND	1	3,717	0	115
TETRA TECH INC	COMMON STOCK	88162G103	97	839	SH		OTR	1	839	0	0
TETRA TECH INC	COMMON STOCK	88162G103	22	194	SH		DFND	5	194	0	0
TEXAS CAPITAL BANCSHARES INC	COMMON STOCK	88224Q107	85	1,433	SH		DFND	1	148	0	1,285
TEXAS CAPITAL BANCSHARES INC	COMMON STOCK	88224Q107	19	312	SH		OTR	1	0	202	110
TG THERAPEUTICS INC	COMMON STOCK	88322Q108	644	12,374	SH		DFND	1	12,234	0	140
TG THERAPEUTICS INC	COMMON STOCK	88322Q108	20	387	SH		OTR	1	387	0	0
THE ODP CORP COM	COMMON STOCK	88337F105	52	1,777	SH		DFND	1	220	0	1,557
THERAPEUTICS MD INC	COMMON STOCK	88338N107	10	7,976	SH		DFND	1	7,976	0	0
THERAPEUTICS MD INC	COMMON STOCK	88338N107	15	12,300	SH		OTR	1	12,300	0	0
TRADE DESK INC THE CLASS A	COMMON STOCK	88339J105	68,888	86,002	SH		DFND	1	85,097	0	905
TRADE DESK INC THE CLASS A	COMMON STOCK	88339J105	9,733	12,151	SH		OTR	1	10,702	1,425	24
TRADE DESK INC (THE)	COMMON STOCK	88339J105	54	68	SH		DFND	5	68	0	0
REALREAL INC THE	COMMON STOCK	88339P101	27	1,386	SH		DFND	1	1,386	0	0
3D SYS CORP DEL	COMMON STOCK	88554D205	65	6,181	SH		DFND	1	6,181	0	0
3D SYS CORP DEL	COMMON STOCK	88554D205	48	4,550	SH		OTR	1	3,050	0	1,500
360 DIGITECH INC	COMMON STOCK	88557W101	43	3,671	SH		DFND	1	2,688	0	983
360 FINANCE INC	COMMON STOCK	88557W101	1	43	SH		DFND	5	43	0	0
3M CO	COMMON STOCK	88579Y101	485,796	2,779,316	SH		DFND	1	2,704,048	0	75,268
3M CO	COMMON STOCK	88579Y101	272,583	1,559,491	SH		OTR	1	1,514,742	22,943	21,806
3M CO	COMMON STOCK	88579Y101	132	754	SH		DFND	5	734	0	20
3M CO	COMMON STOCK	88579Y101	1	4	SH	Put	DFND	1	4	0	0
3M CO	COMMON STOCK	88579Y101	0	10	SH	Put	OTR	1	0	10	0
TIDEWATER INC NEW WT 11	OTHER	88642R174	0	1	SH		DFND	1	1	0	0
TIDEWATER INC NEW WT 11	OTHER	88642R174	0	2	SH		OTR	1	2	0	0
TILRAY INC CLASS 2 COMMON	COMMON STOCK	88688T100	1	113	SH		DFND	1	113	0	0
TILRAY INC CLASS 2 COMMON	COMMON STOCK	88688T100	2	300	SH		OTR	1	300	0	0
TIM S A SPNDR A D R	COMMON STOCK	88706T108	32	2,325	SH		DFND	1	565	0	1,760
TITAN INTL INC ILL	COMMON STOCK	88830M102	755	155,417	SH		DFND	1	155,417	0	0
TITAN MACHY INC	COMMON STOCK	88830R101	21	1,070	SH		DFND	1	1,070	0	0
TITAN MEDICAL INC	COMMON STOCK	88830X819	1	500	SH		DFND	1	500	0	0
TIVITY HEALTH INC	COMMON STOCK	88870R102	39	2,016	SH		DFND	1	1,460	0	556
TIVITY HEALTH INC	COMMON STOCK	88870R102	66	3,351	SH		OTR	1	0	3,351	0
TOPBUILD CORP	COMMON STOCK	89055F103	297	1,612	SH		DFND	1	1,603	0	9
TOPBUILD CORP	COMMON STOCK	89055F103	129	703	SH		OTR	1	48	0	655
TORTOISE ENERGY INFRASTRUCTURE	OTHER	89147L886	1	60	SH		DFND	1	60	0	0
TOWNEBANK PORTSMOUTH VA	COMMON STOCK	89214P109	478	20,372	SH		DFND	1	20,201	0	171
TOWNEBANK PORTSMOUTH VA	COMMON STOCK	89214P109	55	2,337	SH		OTR	1	2,337	0	0
TRANSALTA CORP	COMMON STOCK	89346D107	1	139	SH		OTR	1	0	139	0
TRANSLATE BIO INC	COMMON STOCK	89374L104	8	454	SH		DFND	1	0	0	454
TRANSUNION	COMMON STOCK	89400J107	37,667	379,630	SH		DFND	1	375,979	0	3,651
TRANSUNION	COMMON STOCK	89400J107	3,240	32,659	SH		OTR	1	31,027	1,186	446
TRANSUNION	COMMON STOCK	89400J107	35	351	SH		DFND	5	351	0	0
TRAVERE THERAPEUTICS INC COM	COMMON STOCK	89422G107	3	101	SH		DFND	1	101	0	0
TREAN INS GROUP INC COM	COMMON STOCK	89457R101	41	3,139	SH		OTR	1	0	3,139	0
TREEHOUSE FOODS INC	COMMON STOCK	89469A104	349	8,216	SH		DFND	1	1,825	0	6,391
TREEHOUSE FOODS INC	COMMON STOCK	89469A104	31	718	SH		OTR	1	563	0	155
TREX COMPANY INC	COMMON STOCK	89531P105	761	9,090	SH		DFND	1	3,942	0	5,148
TREX COMPANY INC	COMMON STOCK	89531P105	260	3,103	SH		OTR	1	233	0	2,870
TREX COMPANY INC	COMMON STOCK	89531P105	37	439	SH		DFND	5	439	0	0
TRICIDA INC	COMMON STOCK	89610F101	7	1,006	SH		DFND	1	300	0	706
TRILOGY METALS INC NEW	COMMON STOCK	89621C105	0	6	SH		DFND	1	6	0	0
TRIMTABS US FREE CASH FLOW ETF	OTHER	89628W302	134	2,883	SH		OTR	1	2,883	0	0
TRIP COM GROUP LTD A D R	COMMON STOCK	89677Q107	512	15,179	SH		DFND	1	12,391	0	2,788
TRIP COM GROUP LTD A D R	COMMON STOCK	89677Q107	32	964	SH		OTR	1	964	0	0
TRIP.COM GROUP LTD	COMMON STOCK	89677Q107	2	61	SH		DFND	5	61	0	0
TRISTATE CAPITAL HLDGS INC	COMMON STOCK	89678F100	4	227	SH		DFND	1	0	0	227
TRISTATE CAPITAL HLDGS INC	COMMON STOCK	89678F100	48	2,750	SH		OTR	1	0	2,750	0
TRUEBLUE INC	COMMON STOCK	89785X101	346	18,495	SH		DFND	1	18,020	0	475
TRUEBLUE INC	COMMON STOCK	89785X101	109	5,807	SH		OTR	1	5,807	0	0
TRUIST FINANCIAL CORP	COMMON STOCK	89832Q109	18,999	396,374	SH		DFND	1	340,730	0	55,644
TRUIST FINANCIAL CORP	COMMON STOCK	89832Q109	5,612	117,095	SH		OTR	1	100,024	11,244	5,827
TRUIST FINL CORP	COMMON STOCK	89832Q109	114	2,369	SH		DFND	5	2,142	0	227
T2 BIOSYSTEMS INC	COMMON STOCK	89853L104	1	500	SH		DFND	1	500	0	0
T2 BIOSYSTEMS INC	COMMON STOCK	89853L104	21	16,835	SH		OTR	1	16,835	0	0
TTEC HOLDINGS INC	COMMON STOCK	89854H102	118	1,622	SH		DFND	1	1,622	0	0
TTEC HOLDINGS INC	COMMON STOCK	89854H102	22	300	SH		OTR	1	300	0	0
TURNING PT BRANDS INC	COMMON STOCK	90041L105	2	53	SH		DFND	1	53	0	0
TURNING POINT THERAPEUTICS I	COMMON STOCK	90041T108	14	116	SH		DFND	1	95	0	21
22ND CENTURY GROUP INC	COMMON STOCK	90137F103	11	5,000	SH		DFND	1	5,000	0	0
21VIANET GROUP INC A D R	COMMON STOCK	90138A103	75	2,171	SH		DFND	1	1,744	0	427
TWILIO INC A	COMMON STOCK	90138F102	709	2,093	SH		DFND	1	1,164	0	929
TWILIO INC A	COMMON STOCK	90138F102	305	902	SH		OTR	1	340	457	105
TWILIO INC	COMMON STOCK	90138F102	4	11	SH		DFND	5	0	0	11
TWIST BIOSCIENCE CORP	COMMON STOCK	90184D100	8	60	SH		DFND	1	16	0	44
TWIST BIOSCIENCE CORP	COMMON STOCK	90184D100	1	9	SH		OTR	1	0	0	9
TWITTER INC	COMMON STOCK	90184L102	510	9,424	SH		DFND	1	5,923	0	3,501
TWITTER INC	COMMON STOCK	90184L102	20	373	SH		OTR	1	278	21	74
TWITTER INC	COMMON STOCK	90184L102	0	4	SH		DFND	5	4	0	0
TWO HARBORS INVESTMENT CORP	COMMON STOCK	90187B408	50	7,793	SH		DFND	1	0	0	7,793
2U INC	COMMON STOCK	90214J101	867	21,672	SH		DFND	1	21,672	0	0
2U INC	COMMON STOCK	90214J101	47	1,165	SH		OTR	1	1,165	0	0
2U INC	COMMON STOCK	90214J101	99	2,469	SH		DFND	5	2,469	0	0
UBS ETRACS ALERIAN MLPINFRS ET	OTHER	90274D382	1	73	SH		DFND	1	73	0	0
UFP INDUSTRIES INC COM	COMMON STOCK	90278Q108	314	5,658	SH		DFND	1	5,545	0	113
USANA HEALTH SCIENCES INC	COMMON STOCK	90328M107	14	180	SH		DFND	1	16	0	164
US CONCRETE INC	COMMON STOCK	90333L201	40	1,000	SH		DFND	1	1,000	0	0
US CONCRETE INC	COMMON STOCK	90333L201	4	110	SH		OTR	1	110	0	0
U S PHYSICAL THERAPY INC	COMMON STOCK	90337L108	1,116	9,276	SH		DFND	1	9,159	0	117
U S PHYSICAL THERAPY INC	COMMON STOCK	90337L108	184	1,534	SH		OTR	1	1,534	0	0
UBER TECHNOLOGIES INC	COMMON STOCK	90353T100	1,188	23,289	SH		DFND	1	19,115	0	4,174
UBER TECHNOLOGIES INC	COMMON STOCK	90353T100	45	892	SH		OTR	1	120	614	158
UBER TECHNOLOGIES INC	COMMON STOCK	90353T100	752	14,741	SH		DFND	5	14,741	0	0
UBIQUITI INC	COMMON STOCK	90353W103	226	812	SH		DFND	1	671	0	141
UBIQUITI INC	COMMON STOCK	90353W103	11	38	SH		OTR	1	0	31	7
ULTA BEAUTY, INC	COMMON STOCK	90384S303	1,321	4,601	SH		DFND	1	3,173	0	1,428
ULTA BEAUTY, INC	COMMON STOCK	90384S303	699	2,436	SH		OTR	1	700	1,736	0
ULTA SALON COSMETICS & FRAGRNC	COMMON STOCK	90384S303	426	1,483	SH		DFND	5	1,483	0	0
ULTRA CLEAN HOLDINGS	COMMON STOCK	90385V107	2	56	SH		DFND	1	56	0	0
ULTRAGENYX PHARMACEUTICAL IN	COMMON STOCK	90400D108	670	4,843	SH		DFND	1	4,790	0	53
ULTRAGENYX PHARMACEUTICAL IN	COMMON STOCK	90400D108	1,397	10,092	SH		OTR	1	10,068	24	0
ULTRAPAR PARTICIPACOES A D R	COMMON STOCK	90400P101	31	6,818	SH		DFND	1	2,378	0	4,440
ULTRAPAR PARTICIPACOES A D R	COMMON STOCK	90400P101	0	97	SH		OTR	1	0	0	97
UNITED COMMUNITY BANKS INC	COMMON STOCK	90984P303	1,047	36,805	SH		DFND	1	36,176	0	629
UNITED COMMUNITY BANKS INC	COMMON STOCK	90984P303	83	2,923	SH		OTR	1	2,244	679	0
UNITED STATES OIL L P ETF	OTHER	91232N207	15	444	SH		DFND	1	444	0	0
UNITED STATES OIL L P ETF	OTHER	91232N207	5	137	SH		OTR	1	137	0	0
UNITED STATES OIL FUND LP	OTHER	91232N207	1	26	SH		DFND	5	0	0	26
UNITED THERAPEUTICS CORP	COMMON STOCK	91307C102	341	2,246	SH		DFND	1	1,823	0	423
UNITED THERAPEUTICS CORP	COMMON STOCK	91307C102	27	174	SH		OTR	1	164	0	10
UNITEDHEALTH GROUP INC	COMMON STOCK	91324P102	274,225	781,981	SH		DFND	1	721,056	0	60,925
UNITEDHEALTH GROUP INC	COMMON STOCK	91324P102	96,434	274,992	SH		OTR	1	162,830	6,792	105,370
UNITEDHEALTH GROUP INC	COMMON STOCK	91324P102	1,472	4,189	SH		DFND	5	4,189	0	0
UNITI GROUP INC	COMMON STOCK	91325V108	20	1,678	SH		DFND	1	913	0	765
UNITI GROUP INC	COMMON STOCK	91325V108	0	20	SH		OTR	1	20	0	0
UNITY SOFTWARE INC	COMMON STOCK	91332U101	411	2,680	SH		DFND	1	2,680	0	0
UNITY SOFTWARE INC	COMMON STOCK	91332U101	9	56	SH		OTR	1	56	0	0
UNITY SOFTWARE INC	COMMON STOCK	91332U101	16	104	SH		DFND	5	104	0	0
UNIVAR SOLUTIONS INC	COMMON STOCK	91336L107	271	14,283	SH		DFND	1	13,268	0	1,015
UNIVAR SOLUTIONS INC	COMMON STOCK	91336L107	9	461	SH		OTR	1	346	0	115
UNIVERSAL DISPLAY CORPORATION	COMMON STOCK	91347P105	104	453	SH		DFND	1	347	0	106
UNIVERSAL DISPLAY CORPORATION	COMMON STOCK	91347P105	156	677	SH		OTR	1	0	39	638
UNIVERSAL DISPLAY CORP	COMMON STOCK	91347P105	13	57	SH		DFND	5	57	0	0
UNIVERSAL INSURANCE HOLDINGS	COMMON STOCK	91359V107	95	6,271	SH		DFND	1	6,162	0	109
UNUM GROUP	COMMON STOCK	91529Y106	2,812	122,577	SH		DFND	1	114,077	0	8,500
UNUM GROUP	COMMON STOCK	91529Y106	24	1,034	SH		OTR	1	75	479	480
UPLAND SOFTWARE INC	COMMON STOCK	91544A109	76	1,663	SH		OTR	1	0	1,663	0
UPWORK INC	COMMON STOCK	91688F104	1,485	43,039	SH		DFND	1	43,039	0	0
UPWORK INC	COMMON STOCK	91688F104	663	19,208	SH		OTR	1	19,208	0	0
UPWORK INC	COMMON STOCK	91688F104	0	3	SH		DFND	5	3	0	0
UR ENERGY INC	COMMON STOCK	91688R108	1	1,500	SH		DFND	1	1,500	0	0
URBAN EDGE PROPERTIES W I	COMMON STOCK	91704F104	21	1,644	SH		DFND	1	1,202	0	442
URBAN EDGE PROPERTIES W I	COMMON STOCK	91704F104	1	79	SH		OTR	1	0	76	3
US ECOLOGY INC	COMMON STOCK	91734M103	108	2,980	SH		DFND	1	2,847	0	133
US ECOLOGY INC	COMMON STOCK	91734M103	10	276	SH		OTR	1	276	0	0
VBI VACCINES INC	COMMON STOCK	91822J103	0	80	SH		OTR	1	80	0	0
VAIL RESORTS INC	COMMON STOCK	91879Q109	883	3,166	SH		DFND	1	3,000	0	166
VAIL RESORTS INC	COMMON STOCK	91879Q109	133	476	SH		OTR	1	459	10	7
VAIL RESORTS INC	COMMON STOCK	91879Q109	36	129	SH		DFND	5	129	0	0
VALERO ENERGY CORP	COMMON STOCK	91913Y100	9,797	173,188	SH		DFND	1	168,471	0	4,717
VALERO ENERGY CORP	COMMON STOCK	91913Y100	1,966	34,753	SH		OTR	1	27,330	1,460	5,963
VALERO ENERGY CORP	COMMON STOCK	91913Y100	3	53	SH		DFND	5	0	0	53
VALVOLINE INC	COMMON STOCK	92047W101	265	11,445	SH		DFND	1	6,189	0	5,256
VALVOLINE INC	COMMON STOCK	92047W101	19	823	SH		OTR	1	823	0	0
VANECK VECTORS GOLD MINERS ETF	OTHER	92189F106	1,551	43,061	SH		DFND	1	17,311	0	25,750
VANECK VECTORS GOLD MINERS ETF	OTHER	92189F106	278	7,729	SH		OTR	1	5,129	2,600	0
VANECK VECTORS GOLD MINERS	OTHER	92189F106	52	1,441	SH		DFND	5	0	0	1,441
VANECK VECTORS VDO GAMING AND	OTHER	92189F114	9	125	SH		DFND	1	125	0	0
VANECK VECTORS FALLEN ANGEL	OTHER	92189F437	29	913	SH		DFND	5	913	0	0
VANECK VECTORS INVESTMENT GRAD	OTHER	92189F486	35	1,403	SH		DFND	1	1,403	0	0
VANECK VECTORS LOW CARBON EN E	OTHER	92189F502	109	658	SH		DFND	1	658	0	0
VANECK VECTORS LONG MUNI ETF	OTHER	92189F536	43	1,969	SH		DFND	1	1,969	0	0
VANECK VECTORS MORNINGSTAR WID	OTHER	92189F643	1,926	31,069	SH		DFND	1	31,069	0	0
VANECK VECTORS SEMICONDUCTOR E	OTHER	92189F676	105	482	SH		DFND	1	482	0	0
VANECK VECTORS SEMICONDUCTOR	OTHER	92189F676	17	78	SH		DFND	5	0	0	78
VANECK VECTORS AGRIBUSINESS ET	OTHER	92189F700	60	764	SH		DFND	1	764	0	0
VANECK VECTORS AGRIBUSINESS ET	OTHER	92189F700	1,875	24,075	SH		OTR	1	23,815	0	260
VANECK VECTORS BIOTECH ETF	OTHER	92189F726	25	150	SH		DFND	1	150	0	0
VANECK VECTORS JUNIOR GOLD ETF	OTHER	92189F791	420	7,738	SH		DFND	1	7,738	0	0
VANECK VECTORS JUNIOR GOLD ETF	OTHER	92189F791	14	259	SH		OTR	1	259	0	0
VANECK VECTORS MORNINGSTAR DUR	OTHER	92189H102	32	1,102	SH		DFND	1	1,102	0	0
VANECK VECTORS INTERMEDIATE MU	OTHER	92189H201	207	3,972	SH		OTR	1	0	3,972	0
VANECK VECTORS AMT FREE	OTHER	92189H201	352	6,763	SH		DFND	5	6,545	0	218
VANECK VECTORS JP MORGAN EM LC	OTHER	92189H300	4	121	SH		DFND	1	121	0	0
VANECK VECTORS HIGH YIELD MUNI	OTHER	92189H409	82	1,336	SH		DFND	1	1,336	0	0
VANGUARD TOTAL INTERNATIONAL E	OTHER	92203J407	6,702	114,463	SH		DFND	1	114,051	0	412
VANGUARD TOTAL INTERNATIONAL E	OTHER	92203J407	2,595	44,317	SH		OTR	1	43,723	594	0
VANGUARD TOTAL INTL BOND	OTHER	92203J407	1,626	27,768	SH		DFND	5	25,952	0	1,816
VANGUARD CONSUMER DISCRETIONAR	OTHER	92204A108	100	364	SH		DFND	1	364	0	0
VANGUARD CONSUMER DISCRETIONAR	OTHER	92204A108	23	85	SH		OTR	1	85	0	0
VANGUARD CONSUMER STAPLES INDE	OTHER	92204A207	352	2,025	SH		DFND	1	2,025	0	0
VANGUARD CONSUMER STAPLES	OTHER	92204A207	21	118	SH		DFND	5	118	0	0
VANGUARD ENERGY INDEX FUND ETF	OTHER	92204A306	54	1,046	SH		DFND	1	1,046	0	0
VANGUARD ENERGY INDEX FUND ETF	OTHER	92204A306	3	55	SH		OTR	1	55	0	0
VANGUARD GROUP INC	OTHER	92204A306	0	0	SH		DFND	5	0	0	0
VANGUARD FINANCIALS INDEX FUND	OTHER	92204A405	400	5,491	SH		DFND	1	5,491	0	0
VANGUARD FINANCIALS INDEX FUND	OTHER	92204A405	7	100	SH		OTR	1	100	0	0
VANGUARD FINANCIALS	OTHER	92204A405	222	3,044	SH		DFND	5	2,824	0	220
VANGUARD HEALTH CARE INDEX FUN	OTHER	92204A504	2,068	9,243	SH		DFND	1	9,243	0	0
VANGUARD HEALTH CARE INDEX FUN	OTHER	92204A504	149	665	SH		OTR	1	665	0	0
VANGUARD HEALTH CARE VIPER	OTHER	92204A504	2,660	11,890	SH		DFND	5	313	0	11,577
VANGUARD INDUSTRIALS INDEX FUN	OTHER	92204A603	133	782	SH		DFND	1	782	0	0
VANGUARD INDUSTRIALS	OTHER	92204A603	21	121	SH		DFND	5	0	0	121
VANGUARD INFORMATION ETF	OTHER	92204A702	5,097	14,408	SH		DFND	1	14,208	0	200
VANGUARD INFORMATION ETF	OTHER	92204A702	639	1,805	SH		OTR	1	1,190	0	615
VANGUARD INFO TECH INDEX	OTHER	92204A702	562	1,589	SH		DFND	5	91	0	1,498
VANGUARD MATERIALS INDEX FUND	OTHER	92204A801	219	1,394	SH		DFND	1	1,394	0	0
VANGUARD UTILITIES INDEX FUND	OTHER	92204A876	129	941	SH		DFND	1	941	0	0
VANGUARD UTILITIES	OTHER	92204A876	40	292	SH		DFND	5	0	0	292
VANGUARD COMMUNICATION SERVICE	OTHER	92204A884	45	371	SH		DFND	1	371	0	0
VANGUARD COMMUNICATION SERVICE	OTHER	92204A884	43	354	SH		OTR	1	354	0	0
VANGUARD SHORT TERM TREASURY E	OTHER	92206C102	223	3,626	SH		OTR	1	3,626	0	0
VANGUARD SHORT TERM GOVT BOND	OTHER	92206C102	2	40	SH		DFND	5	0	0	40
VANGUARD SHORT TERM CORPORATE	OTHER	92206C409	1,330	15,972	SH		DFND	1	8,273	0	7,699
VANGUARD SHORT TERM CORPORATE	OTHER	92206C409	3,220	38,676	SH		OTR	1	20,406	18,195	75
VANGUARD SHORT TERM CORP BOND	OTHER	92206C409	530	6,371	SH		DFND	5	6,324	0	47
VANGUARD RUSSELL 2000 INDEX FU	OTHER	92206C664	42	267	SH		DFND	1	267	0	0
VANGUARD RUSSELL 1000 GROWTH E	OTHER	92206C680	136	548	SH		DFND	1	548	0	0
VANGUARD INTERMEDIATE TERM ETF	OTHER	92206C706	298	4,288	SH		OTR	1	4,288	0	0
VANGUARD INTM TERM GOVT BOND	OTHER	92206C706	10	138	SH		DFND	5	138	0	0
VANGUARD RUSSELL 1000 VALUE ET	OTHER	92206C714	67	555	SH		DFND	1	450	0	105
VANGUARD RUSSELL 1000 VALUE ET	OTHER	92206C714	3	25	SH		OTR	1	25	0	0
VANGUARD RUSSELL 1000 INDEX FU	OTHER	92206C730	9	54	SH		DFND	1	54	0	0
VANGUARD MORTGAGE BACKED ETF	OTHER	92206C771	142	2,635	SH		DFND	1	2,487	0	148
VANGUARD MORTGAGE BACKED ETF	OTHER	92206C771	300	5,543	SH		OTR	1	5,543	0	0
VANGUARD MTGE BUSINESS SEC	OTHER	92206C771	775	14,340	SH		DFND	5	11,609	0	2,731
VANGUARD LONG TERM TREASURY ET	OTHER	92206C847	3	33	SH		DFND	1	33	0	0
VANGUARD LONG TERM TREASURY ET	OTHER	92206C847	146	1,519	SH		OTR	1	1,519	0	0
VANGUARD INTERMEDIATE TERM ETF	OTHER	92206C870	1,233	12,693	SH		DFND	1	9,251	0	3,442
VANGUARD INTERMEDIATE TERM ETF	OTHER	92206C870	229	2,361	SH		OTR	1	1,937	0	424
VANGUARD INTM TERM CORP BOND	OTHER	92206C870	55	565	SH		DFND	5	565	0	0
VAREX IMAGING CORP	COMMON STOCK	92214X106	56	3,368	SH		DFND	1	3,160	0	208
VAREX IMAGING CORP	COMMON STOCK	92214X106	5	320	SH		OTR	1	320	0	0
VARIAN MED SYS INC	COMMON STOCK	92220P105	5,098	29,133	SH		DFND	1	28,865	0	268
VARIAN MED SYS INC	COMMON STOCK	92220P105	2,328	13,302	SH		OTR	1	12,074	783	445
VECTOR GROUP LTD	COMMON STOCK	92240M108	45	3,903	SH		DFND	1	3,903	0	0
VECTRUS INC	COMMON STOCK	92242T101	12	235	SH		DFND	1	212	0	23
VECTRUS INC	COMMON STOCK	92242T101	0	6	SH		OTR	1	0	0	6
VEDANTA LTD A D R	COMMON STOCK	92242Y100	471	53,608	SH		DFND	1	19,309	0	34,299
VEDANTA LTD A D R	COMMON STOCK	92242Y100	9	995	SH		OTR	1	0	261	734
VELODYNE LIDAR INC	COMMON STOCK	92259F101	273	11,959	SH		DFND	1	11,959	0	0
VELODYNE LIDAR INC	COMMON STOCK	92259F101	40	1,740	SH		OTR	1	1,740	0	0
VENTAS INC	COMMON STOCK	92276F100	833	16,988	SH		DFND	1	14,141	0	2,847
VENTAS INC	COMMON STOCK	92276F100	111	2,258	SH		OTR	1	1,765	376	117
VENTAS INC	COMMON STOCK	92276F100	47	958	SH		DFND	5	958	0	0
VEONEER INC	COMMON STOCK	92336X109	13	605	SH		DFND	1	605	0	0
VEONEER INC	COMMON STOCK	92336X109	12	582	SH		OTR	1	582	0	0
VERACYTE INC	COMMON STOCK	92337F107	36	737	SH		DFND	1	737	0	0
VERACYTE INC	COMMON STOCK	92337F107	135	2,752	SH		OTR	1	371	0	2,381
VEREIT INC	COMMON STOCK	92339V308	405	10,706	SH		DFND	1	9,021	0	1,685
VEREIT INC	COMMON STOCK	92339V308	4	95	SH		OTR	1	95	0	0
VEREIT INC	COMMON STOCK	92339V308	10	277	SH		DFND	5	277	0	0
VERIZON COMMUNICATIONS INC	COMMON STOCK	92343V104	133,725	2,276,169	SH		DFND	1	2,171,157	0	105,012
VERIZON COMMUNICATIONS INC	COMMON STOCK	92343V104	56,374	959,554	SH		OTR	1	881,070	26,814	51,670
VERIZON COMMUNICATIONS	COMMON STOCK	92343V104	675	11,487	SH		DFND	5	11,082	0	405
VERINT SYSTEMS INC	COMMON STOCK	92343X100	1,698	25,277	SH		DFND	1	25,027	0	250
VERINT SYSTEMS INC	COMMON STOCK	92343X100	334	4,971	SH		OTR	1	4,971	0	0
VERISK ANALYTICS INC CL A	COMMON STOCK	92345Y106	13,382	64,464	SH		DFND	1	62,240	0	2,224
VERISK ANALYTICS INC CL A	COMMON STOCK	92345Y106	1,049	5,052	SH		OTR	1	4,511	138	403
VERISK ANALYTICS INC	COMMON STOCK	92345Y106	886	4,264	SH		DFND	5	4,264	0	0
VERICEL CORP	COMMON STOCK	92346J108	17	555	SH		OTR	1	0	555	0
VERITONE INC	COMMON STOCK	92347M100	3	92	SH		DFND	1	0	0	92
VERRA MOBILITY CORP	COMMON STOCK	92511U102	2	134	SH		DFND	1	0	0	134
VERRA MOBILITY CORP	COMMON STOCK	92511U102	24	1,767	SH		OTR	1	0	1,767	0
VERTEX PHARMACEUTICALS INC	COMMON STOCK	92532F100	1,802	7,624	SH		DFND	1	5,601	0	2,023
VERTEX PHARMACEUTICALS INC	COMMON STOCK	92532F100	1,480	6,262	SH		OTR	1	2,707	2,943	612
VERTEX PHARMACEUTICALS INC	COMMON STOCK	92532F100	52	222	SH		DFND	5	222	0	0
VERTEX ENERGY INC	COMMON STOCK	92534K107	290	407,781	SH		DFND	1	407,781	0	0
VERTIV HOLDINGS LLC	COMMON STOCK	92537N108	2	99	SH		DFND	5	99	0	0
VERTEX INC CL A	COMMON STOCK	92538J106	1,388	39,840	SH		DFND	1	39,840	0	0
VERTEX INC CL A	COMMON STOCK	92538J106	191	5,468	SH		OTR	1	1,143	0	4,325
VIAD CORP	COMMON STOCK	92552R406	26	730	SH		DFND	1	730	0	0
VIASAT INC	COMMON STOCK	92552V100	39	1,200	SH		DFND	1	1,200	0	0
VIASAT INC	COMMON STOCK	92552V100	3	98	SH		OTR	1	68	0	30
VIACOMCBS INC CLASS A	COMMON STOCK	92556H107	25	669	SH		DFND	1	669	0	0
VIACOMCBS INC CLASS A	COMMON STOCK	92556H107	9	239	SH		OTR	1	239	0	0
VIACOMCBS INC CLASS B	COMMON STOCK	92556H206	3,225	86,568	SH		DFND	1	76,197	0	10,371
VIACOMCBS INC CLASS B	COMMON STOCK	92556H206	190	5,095	SH		OTR	1	3,969	947	179
VIATRIS INC	COMMON STOCK	92556V106	67,314	3,592,013	SH		DFND	1	3,559,521	0	32,492
VIATRIS INC	COMMON STOCK	92556V106	11,007	587,401	SH		OTR	1	560,598	17,555	9,248
VIATRIS INC	COMMON STOCK	92556V106	14	760	SH		DFND	5	739	0	21
VICTORYSHARES US 500 ENHANCED	OTHER	92647N782	9	142	SH		DFND	1	142	0	0
VICTORYSHARES US EQ INCOME ETF	OTHER	92647N824	2	37	SH		DFND	1	37	0	0
VICTORYSHARES US LARGE CAP HIG	OTHER	92647N865	295	6,265	SH		DFND	1	6,265	0	0
VICTORYSHARES US SMALL CAP HIG	OTHER	92647N873	66	1,331	SH		DFND	1	1,331	0	0
VIEWRAY INC	COMMON STOCK	92672L107	2	637	SH		DFND	1	0	0	637
VIKING THERAPEUTICS INC	COMMON STOCK	92686J106	7	1,193	SH		DFND	1	0	0	1,193
VIKING THERAPEUTICS INC	COMMON STOCK	92686J106	1	266	SH		OTR	1	0	0	266
VIPER ENERGY PARTNERS LP	COMMON STOCK	92763M105	12	1,070	SH		DFND	1	70	0	1,000
VIPSHOP HOLDINGS LTD A D R	COMMON STOCK	92763W103	1,102	39,172	SH		DFND	1	35,363	0	3,809
VIPSHOP HOLDINGS LTD A D R	COMMON STOCK	92763W103	72	2,583	SH		OTR	1	2,140	0	443
VIR BIOTECHNOLOGY INC	COMMON STOCK	92764N102	40	1,500	SH		DFND	1	1,500	0	0
VIRGIN GALACTIC HOLDINGS INC	COMMON STOCK	92766K106	24	1,000	SH		DFND	1	1,000	0	0
VIRNETX HOLDING CORPORATION	COMMON STOCK	92823T108	54	10,750	SH		DFND	1	10,750	0	0
VIRNETX HOLDING CORPORATION	COMMON STOCK	92823T108	31	6,068	SH		OTR	1	6,068	0	0
VISA INC CLASS A SHARES	COMMON STOCK	92826C839	182,820	835,824	SH		DFND	1	806,279	0	29,545
VISA INC CLASS A SHARES	COMMON STOCK	92826C839	59,790	273,350	SH		OTR	1	189,387	16,363	67,600
VISA INC	COMMON STOCK	92826C839	2,371	10,831	SH		DFND	5	10,739	0	92
VISA INC CLASS A SHARES	COMMON STOCK	92826C839	0	9	SH	Put	DFND	1	9	0	0
VIRTUSA CORP	COMMON STOCK	92827P102	6	111	SH		DFND	1	111	0	0
VIRTUS INVESTMENT PARTNERS	COMMON STOCK	92828Q109	9	42	SH		DFND	1	31	0	11
VISTEON CORP	COMMON STOCK	92839U206	1,482	11,810	SH		DFND	1	11,709	0	101
VISTEON CORP	COMMON STOCK	92839U206	40	320	SH		OTR	1	320	0	0
VISTRA ENERGY CORP	COMMON STOCK	92840M102	336	17,089	SH		DFND	1	13,908	0	3,181
VISTRA ENERGY CORP	COMMON STOCK	92840M102	42	2,114	SH		OTR	1	2,114	0	0
VISTRA ENERGY CORP	COMMON STOCK	92840M102	64	3,247	SH		DFND	5	3,247	0	0
VISTRA ENERGY CORP WT 2	OTHER	92840M128	0	1,215	SH		DFND	1	1,215	0	0
VOCERA COMMUNICATIONS INC	COMMON STOCK	92857F107	637	15,312	SH		DFND	1	10,294	0	5,018
VOCERA COMMUNICATIONS INC	COMMON STOCK	92857F107	221	5,318	SH		OTR	1	438	0	4,880
VODAFONE GROUP PLC A D R	COMMON STOCK	92857W308	547	33,197	SH		DFND	1	24,681	0	8,516
VODAFONE GROUP PLC A D R	COMMON STOCK	92857W308	373	22,609	SH		OTR	1	21,264	1,047	298
VODAFONE GROUP PLC	COMMON STOCK	92857W308	296	17,983	SH		DFND	5	17,983	0	0
VONAGE HOLDINGS CORP	COMMON STOCK	92886T201	567	44,115	SH		DFND	1	44,024	0	91
VONAGE HOLDINGS CORP	COMMON STOCK	92886T201	8	585	SH		OTR	1	585	0	0
VOYAGER THERAPEUTICS INC	COMMON STOCK	92915B106	2	284	SH		OTR	1	0	284	0
VROOM INC COM	COMMON STOCK	92918V109	12	300	SH		DFND	1	300	0	0
VUZIX CORP	COMMON STOCK	92921W300	91	10,000	SH		DFND	1	10,000	0	0
WNS HOLDINGS LTD A D R	COMMON STOCK	92932M101	3,994	55,427	SH		DFND	1	53,183	0	2,244
WNS HOLDINGS LTD A D R	COMMON STOCK	92932M101	1,097	15,222	SH		OTR	1	13,835	1,315	72
WNS HOLDINGS LTD	COMMON STOCK	92932M101	10	145	SH		DFND	5	145	0	0
WP CAREY INC	COMMON STOCK	92936U109	1,032	14,623	SH		DFND	1	7,577	0	7,046
WP CAREY INC	COMMON STOCK	92936U109	905	12,813	SH		OTR	1	6,598	6,200	15
W P CAREY INC	COMMON STOCK	92936U109	1	15	SH		DFND	5	15	0	0
WPP PLC SPON A D R	COMMON STOCK	92937A102	439	8,116	SH		DFND	1	5,329	0	2,787
WPP PLC SPON A D R	COMMON STOCK	92937A102	14	250	SH		OTR	1	163	18	69
WPP PLC	COMMON STOCK	92937A102	9	164	SH		DFND	5	164	0	0
WEC ENERGY GROUP INC	COMMON STOCK	92939U106	14,398	156,452	SH		DFND	1	146,560	0	9,892
WEC ENERGY GROUP INC	COMMON STOCK	92939U106	8,201	89,112	SH		OTR	1	87,952	49	1,111
WEC ENERGY GROUP INC	COMMON STOCK	92939U106	78	843	SH		DFND	5	843	0	0
WALKER & DUNLOP INC	COMMON STOCK	93148P102	2,385	25,918	SH		DFND	1	25,918	0	0
WALKER & DUNLOP INC	COMMON STOCK	93148P102	80	865	SH		OTR	1	865	0	0
WARRIOR MET COAL INC	COMMON STOCK	93627C101	20	950	SH		DFND	1	262	0	688
WASHINGTON PRIME GROUP	COMMON STOCK	93964W405	138	21,142	SH		DFND	1	21,115	0	27
WASTE CONNECTIONS INC	COMMON STOCK	94106B101	422	4,114	SH		DFND	1	1,193	0	2,921
WASTE CONNECTIONS INC	COMMON STOCK	94106B101	1,357	13,227	SH		OTR	1	1,500	3,427	8,300
WASTE CONNECTIONS INC	COMMON STOCK	94106B101	1	9	SH		DFND	5	9	0	0
WASTE MANAGEMENT INC	COMMON STOCK	94106L109	16,644	141,131	SH		DFND	1	135,379	0	5,752
WASTE MANAGEMENT INC	COMMON STOCK	94106L109	6,983	59,215	SH		OTR	1	53,371	1,741	4,103
WASTE MANAGEMENT INC	COMMON STOCK	94106L109	218	1,849	SH		DFND	5	1,849	0	0
WATERSTONE FINANCIAL INC	COMMON STOCK	94188P101	15	795	SH		DFND	1	795	0	0
WAYFAIR INC CLASS A	COMMON STOCK	94419L101	321	1,422	SH		DFND	1	1,097	0	325
WAYFAIR INC CLASS A	COMMON STOCK	94419L101	46	201	SH		OTR	1	121	50	30
WAYFAIR INC	COMMON STOCK	94419L101	0	1	SH		DFND	5	1	0	0
WELLTOWER INC	COMMON STOCK	95040Q104	1,968	30,464	SH		DFND	1	28,056	0	2,408
WELLTOWER INC	COMMON STOCK	95040Q104	542	8,393	SH		OTR	1	8,183	83	127
WELLTOWER INC	COMMON STOCK	95040Q104	17	270	SH		DFND	5	270	0	0
WENDYS CO THE	COMMON STOCK	95058W100	910	41,535	SH		DFND	1	40,675	0	860
WENDYS CO THE	COMMON STOCK	95058W100	201	9,185	SH		OTR	1	8,978	135	72
WESCO INTERNATIONAL INC	COMMON STOCK	95082P105	774	9,876	SH		DFND	1	4,727	0	5,149
WESCO INTERNATIONAL INC	COMMON STOCK	95082P105	48	613	SH		OTR	1	385	228	0
WEST BANCORPORATION INC	COMMON STOCK	95123P106	466	24,155	SH		DFND	1	24,155	0	0
WESTERN ASSET HIGH INCOME FD I	OTHER	95766J102	8	1,225	SH		DFND	1	1,225	0	0
WESTERN ASSET HIGH INCOME OP	OTHER	95766K109	3	533	SH		OTR	1	533	0	0
WESTERN ASSET MUN HI INCM FDCO	OTHER	95766N103	5	713	SH		DFND	1	713	0	0
WESTERN ASSET INFLATION LINKED	OTHER	95766R104	195	15,743	SH		DFND	1	15,743	0	0
WESTERN COPPER CORPORATION	COMMON STOCK	95805V108	0	25	SH		DFND	1	25	0	0
WESTROCK CO	COMMON STOCK	96145D105	724	16,618	SH		DFND	1	14,179	0	2,439
WESTROCK CO	COMMON STOCK	96145D105	535	12,304	SH		OTR	1	12,260	34	10
WESTROCK CO	COMMON STOCK	96145D105	14	310	SH		DFND	5	310	0	0
WEX INC	COMMON STOCK	96208T104	1,317	6,469	SH		DFND	1	6,250	0	219
WEX INC	COMMON STOCK	96208T104	140	688	SH		OTR	1	596	6	86
WIDEOPENWEST INC	COMMON STOCK	96758W101	3	239	SH		DFND	1	239	0	0
WINTRUST FINANCIAL CORP	COMMON STOCK	97650W108	3,331	54,543	SH		DFND	1	53,075	0	1,468
WINTRUST FINANCIAL CORP	COMMON STOCK	97650W108	123	2,012	SH		OTR	1	1,007	979	26
WIPRO LIMITED A D R	COMMON STOCK	97651M109	746	132,036	SH		DFND	1	113,015	0	19,021
WIPRO LIMITED A D R	COMMON STOCK	97651M109	142	25,286	SH		OTR	1	24,596	690	0
WISDOMTREE US HGH DIV FD ETF	OTHER	97717W208	30	429	SH		DFND	1	429	0	0
WISDOMTREE EMERGING MARKETS SM	OTHER	97717W281	22	457	SH		DFND	1	457	0	0
WISDOMTREE EMERGING MARKETS SM	OTHER	97717W281	24	500	SH		OTR	1	500	0	0
WISDOMTREE EMERGING MKTS SML	OTHER	97717W281	21	439	SH		DFND	5	0	0	439
WISDOMTREE US LC DIV FD ETF	OTHER	97717W307	135	1,258	SH		DFND	1	1,258	0	0
WISDOMTREE US LC DIV FD ETF	OTHER	97717W307	684	6,365	SH		OTR	1	6,365	0	0
WISDOMTREE LARGECAP DIVIDEND	OTHER	97717W307	0	0	SH		DFND	5	0	0	0
WISDOMTREE EMERGING MARKETS HI	OTHER	97717W315	167	4,058	SH		DFND	1	4,058	0	0
WISDOMTREE EMERGING MKTS EQTY	OTHER	97717W315	1	19	SH		DFND	5	19	0	0
WISDOMTREE US DIV EX FNL FD ET	OTHER	97717W406	40	476	SH		DFND	1	476	0	0
WISDOMTREE US DIV EX FNL FD ET	OTHER	97717W406	643	7,680	SH		OTR	1	7,680	0	0
WISDOMTREE INTEREST RATE HEDGE	OTHER	97717W430	30	1,389	SH		DFND	1	0	0	1,389
WISDOMTREE BOFA MERRILL LYNCH	OTHER	97717W430	170	7,774	SH		DFND	5	7,774	0	0
WISDOMTREE US MC DIV FD ETF	OTHER	97717W505	44	1,260	SH		DFND	1	1,260	0	0
WISDOMTREE MIDCAP DIVIDEND	OTHER	97717W505	4	121	SH		DFND	5	115	0	6
WISDOMTREE US QUALITY SHAREHLD	OTHER	97717W547	28	286	SH		DFND	1	286	0	0
WISDOMTREE U S SMALLCAP FUND E	OTHER	97717W562	27	690	SH		DFND	1	690	0	0
WISDOMTREE MIDCAP EARNINGS	OTHER	97717W570	7	168	SH		DFND	5	0	0	168
WISDOMTREE U S LARGECAP FUND E	OTHER	97717W588	12	295	SH		DFND	1	295	0	0
WISDOMTREE INTERNATIONAL ETF	OTHER	97717W760	136	2,004	SH		DFND	1	2,004	0	0
WISDOMTREE INTERNATIONAL ETF	OTHER	97717W760	196	2,880	SH		OTR	1	2,880	0	0
WISDOMTREE INTL SMALL CAP	OTHER	97717W760	1	10	SH		DFND	5	0	0	10
WISDOMTREE INTERNATIONAL MIDCA	OTHER	97717W778	35	553	SH		DFND	1	553	0	0
WISDOMTREE INTERNATIONAL ETF	OTHER	97717W794	11	247	SH		DFND	1	247	0	0
WISDOMTREE INTL LARGECAP DVD	OTHER	97717W794	19	424	SH		DFND	5	0	0	424
WISDOMTREE GLOBAL EX US QUAL D	OTHER	97717W844	435	5,710	SH		DFND	1	5,710	0	0
WISDOMTREE JAPAN HEDGED EQUITY	OTHER	97717W851	49	892	SH		DFND	1	892	0	0
WISDOMTREE EUROPE SMALLCAP ETF	OTHER	97717W869	1	8	SH		DFND	1	8	0	0
WISDOMTREE EMMKTS EX STT OWND	OTHER	97717X578	19	489	SH		OTR	1	0	0	489
WISDOMTREE FLOATING RATE	OTHER	97717X628	44	1,754	SH		DFND	5	0	0	1,754
WISDOMTREE US QUALITY DIVIDEND	OTHER	97717X669	33	609	SH		DFND	1	609	0	0
WISDOMTREE EUROPE HEDGED EQUIT	OTHER	97717X701	33	504	SH		DFND	1	104	0	400
WISDOMTREE CONTINUOUS COMMODIT	OTHER	97717Y683	34	1,795	SH		DFND	1	1,795	0	0
WISDOMTREE ENHANCED COMMODITY	OTHER	97717Y683	11	571	SH		DFND	5	0	0	571
WORKDAY INC	COMMON STOCK	98138H101	2,129	8,885	SH		DFND	1	8,611	0	274
WORKDAY INC	COMMON STOCK	98138H101	1,934	8,070	SH		OTR	1	969	4,195	2,906
WORKDAY INC	COMMON STOCK	98138H101	165	689	SH		DFND	5	689	0	0
WORKHORSE GROUP INC	COMMON STOCK	98138J206	334	16,905	SH		DFND	1	16,905	0	0
WORKHORSE GROUP INC	COMMON STOCK	98138J206	6	326	SH		OTR	1	326	0	0
WORKIVA INC	COMMON STOCK	98139A105	1,398	15,259	SH		DFND	1	14,429	0	830
WORKIVA INC	COMMON STOCK	98139A105	226	2,466	SH		OTR	1	1,485	981	0
WORKIVA INC	COMMON STOCK	98139A105	276	3,007	SH		DFND	5	3,007	0	0
WORLD WRESTLING ENTERTAINMENT	COMMON STOCK	98156Q108	8	169	SH		DFND	1	169	0	0
WORLD WRESTLING ENTERTAINMENT	COMMON STOCK	98156Q108	1	30	SH		OTR	1	0	0	30
WORLD WRESTLING ENTMNT INC	COMMON STOCK	98156Q108	0	10	SH		DFND	5	10	0	0
WPX ENERGY INC	COMMON STOCK	98212B103	480	58,896	SH		DFND	1	38,313	0	20,583
WPX ENERGY INC	COMMON STOCK	98212B103	20	2,459	SH		OTR	1	1,446	166	847
WRAP TECHNOLOGIES INC	COMMON STOCK	98212N107	2,415	500,000	SH		DFND	1	500,000	0	0
WRAP TECHNOLOGIES INC	COMMON STOCK	98212N107	10	2,000	SH		OTR	1	2,000	0	0
WYNDHAM DESTINATIONS INC	COMMON STOCK	98310W108	1,272	28,353	SH		DFND	1	25,989	0	2,364
WYNDHAM DESTINATIONS INC	COMMON STOCK	98310W108	835	18,603	SH		OTR	1	18,481	0	122
WYNDAM HOTELS RESORTS INC	COMMON STOCK	98311A105	440	7,398	SH		DFND	1	6,902	0	496
WYNDAM HOTELS RESORTS INC	COMMON STOCK	98311A105	912	15,341	SH		OTR	1	15,296	0	45
WYNDHAM HOTELS & RESORTS	COMMON STOCK	98311A105	15	254	SH		DFND	5	254	0	0
XPEL INC	COMMON STOCK	98379L100	7	135	SH		OTR	1	0	135	0
XCEL ENERGY INC	COMMON STOCK	98389B100	22,162	332,408	SH		DFND	1	319,022	0	13,386
XCEL ENERGY INC	COMMON STOCK	98389B100	10,182	152,723	SH		OTR	1	123,343	18,161	11,219
XCEL ENERGY INC	COMMON STOCK	98389B100	170	2,551	SH		DFND	5	2,551	0	0
XPERI HOLDING CORP COM	COMMON STOCK	98390M103	25	1,187	SH		DFND	1	1,187	0	0
XBIOTECH INC	COMMON STOCK	98400H102	8	520	SH		DFND	1	354	0	166
XAI OCTAGON FLOATING RATE	OTHER	98400T106	375	50,000	SH		DFND	1	50,000	0	0
XENCOR INC	COMMON STOCK	98401F105	1	17	SH		DFND	1	5	0	12
XYLEM INC	COMMON STOCK	98419M100	32,400	318,298	SH		DFND	1	312,988	0	5,310
XYLEM INC	COMMON STOCK	98419M100	2,751	27,019	SH		OTR	1	26,014	151	854
XYLEM INC	COMMON STOCK	98419M100	2	21	SH		DFND	5	21	0	0
XEROX HOLDINGS CORP	COMMON STOCK	98421M106	47	2,043	SH		DFND	1	1,615	0	428
XEROX HOLDINGS CORP	COMMON STOCK	98421M106	19	810	SH		OTR	1	810	0	0
XPENG INC A D R	COMMON STOCK	98422D105	158	3,706	SH		DFND	1	3,471	0	235
YAMANA GOLD INC	COMMON STOCK	98462Y100	331	58,000	SH		DFND	1	50,861	0	7,139
YAMANA GOLD INC	COMMON STOCK	98462Y100	52	9,201	SH		OTR	1	9,201	0	0
YAMANA GOLD INC	COMMON STOCK	98462Y100	16	2,889	SH		DFND	5	2,889	0	0
YEXT INC	COMMON STOCK	98585N106	9	570	SH		DFND	1	570	0	0
YETI HOLDINGS INC	COMMON STOCK	98585X104	4	61	SH		DFND	1	29	0	32
YETI HOLDINGS INC	COMMON STOCK	98585X104	245	3,581	SH		OTR	1	0	3,581	0
YOUDAO INC A D R	COMMON STOCK	98741T104	22	812	SH		DFND	1	812	0	0
YUM CHINA HOLDINGS INC	COMMON STOCK	98850P109	2,917	51,104	SH		DFND	1	46,913	0	4,191
YUM CHINA HOLDINGS INC	COMMON STOCK	98850P109	1,366	23,914	SH		OTR	1	19,793	2,951	1,170
YUM CHINA HOLDINGS INC	COMMON STOCK	98850P109	12	216	SH		DFND	5	216	0	0
ZAI LAB LTD A D R	COMMON STOCK	98887Q104	275	2,031	SH		DFND	1	1,171	0	860
ZAI LAB LTD A D R	COMMON STOCK	98887Q104	1	6	SH		OTR	1	0	0	6
ZEALAND PHARMA A S A D R	COMMON STOCK	98920Y304	3	94	SH		DFND	1	27	0	67
ZEALAND PHARMA A S A D R	COMMON STOCK	98920Y304	4	120	SH		OTR	1	0	120	0
ZENDESK INC	COMMON STOCK	98936J101	187	1,304	SH		DFND	1	625	0	679
ZENDESK INC	COMMON STOCK	98936J101	266	1,856	SH		OTR	1	120	698	1,038
ZILLOW GROUP INC	COMMON STOCK	98954M101	1,784	13,123	SH		DFND	1	12,632	0	491
ZILLOW GROUP INC	COMMON STOCK	98954M101	341	2,510	SH		OTR	1	2,493	8	9
ZILLOW GROUP INC C	COMMON STOCK	98954M200	7,289	56,158	SH		DFND	1	56,158	0	0
ZILLOW GROUP INC C	COMMON STOCK	98954M200	13	104	SH		OTR	1	104	0	0
ZILLOW GROUP INC	COMMON STOCK	98954M200	51	393	SH		DFND	5	393	0	0
ZIMMER BIOMET HOLDINGS INC	COMMON STOCK	98956P102	15,449	100,260	SH		DFND	1	96,710	0	3,550
ZIMMER BIOMET HOLDINGS INC	COMMON STOCK	98956P102	7,452	48,357	SH		OTR	1	45,908	1,070	1,379
ZIMMER BIOMET HOLDINGS INC	COMMON STOCK	98956P102	93	605	SH		DFND	5	605	0	0
ZIX CORPORATION	COMMON STOCK	98974P100	119	13,835	SH		DFND	1	13,835	0	0
ZIX CORPORATION	COMMON STOCK	98974P100	40	4,595	SH		OTR	1	4,595	0	0
ZOGENIX INC	COMMON STOCK	98978L204	6	300	SH		DFND	1	55	0	245
ZOETIS INC	COMMON STOCK	98978V103	11,660	70,457	SH		DFND	1	63,220	0	7,237
ZOETIS INC	COMMON STOCK	98978V103	4,626	27,950	SH		OTR	1	19,623	5,489	2,838
ZOETIS INC	COMMON STOCK	98978V103	1,103	6,653	SH		DFND	5	6,569	0	84
ZTO EXPRESS CAYMAN INC A D R	COMMON STOCK	98980A105	185	6,348	SH		DFND	1	3,451	0	2,897
ZTO EXPRESS CAYMAN INC A D R	COMMON STOCK	98980A105	105	3,583	SH		OTR	1	0	0	3,583
ZSCALER INC	COMMON STOCK	98980G102	2,638	13,210	SH		DFND	1	12,913	0	297
ZSCALER INC	COMMON STOCK	98980G102	1,039	5,205	SH		OTR	1	5,013	87	105
ZOOM VIDEO COMMUNICATIONS A	COMMON STOCK	98980L101	1,538	4,559	SH		DFND	1	4,018	0	541
ZOOM VIDEO COMMUNICATIONS A	COMMON STOCK	98980L101	561	1,663	SH		OTR	1	948	176	539
ZOOM VIDEO COMMUNICATIONS	COMMON STOCK	98980L101	11	32	SH		DFND	5	32	0	0
ZUORA INC CLASS A	COMMON STOCK	98983V106	1,293	92,854	SH		DFND	1	92,808	0	46
ZUORA INC CLASS A	COMMON STOCK	98983V106	81	5,822	SH		OTR	1	5,822	0	0
ZYMEWORKS INC	COMMON STOCK	98985W102	316	6,683	SH		DFND	1	6,683	0	0
ZYMEWORKS INC	COMMON STOCK	98985W102	5	100	SH		OTR	1	100	0	0
ZYNGA INC	COMMON STOCK	98986T108	138	14,011	SH		DFND	1	12,611	0	1,400
ZYNGA INC	COMMON STOCK	98986T108	280	28,302	SH		OTR	1	10,000	901	17,401
ZYNGA INC	COMMON STOCK	98986T108	0	31	SH		DFND	5	31	0	0