investment in the first quarter of 2023. This segment was also unfavorably impacted by $415,000 of additional total borrowings interest expense due to the higher average balance of FHLB term borrowings. Favorably impacting net income for this segment were lower expenses by $344,000 for legal and professional services costs and an increased level of interest income from investment securities by $150,000 as the higher interest rates and a portfolio restructuring in late 2023 resulted in an improved overall total securities yield.
…..BALANCE SHEET…..The Company’s total consolidated assets were $1.4 billion at March 31, 2024, which declined by $5.1 million, or 0.4%, from the December 31, 2023 asset level. This change was related, primarily, to reduced total loans which were partially offset by increased levels of cash and cash equivalents, investment securities, leases, and other assets. Specifically, total loans decreased $11.8 million, or 1.1%, as payoff activity so far in 2024 has surpassed loan originations. During 2023 and into 2024, the spread between overnight borrowings and the yield on new investment securities ranged from negative to only marginally positive causing a slowdown in purchasing activity. Thus, the $729,000, or 0.3%, increase in investment securities was used to replace cash flow from maturing securities to maintain appropriate balances for pledging purposes related to public funds deposits. The slowdown in loan originations and investment security purchases along with an increase in total deposits led to a $3.4 million, or 24.4%, increase in cash and cash equivalents. The $1.2 million, or 38.1%, increase in the right-of-use asset for operating and financing leases resulted from the execution of two new leases for office locations and one new lease of equipment during the first quarter of 2024. Finally, other assets increased $2.0 million, or 5.3%, due to an increase in the market values for the interest rate swaps and hedges as well as amounts prepaid for state shares tax.
Total deposits increased by $18.2 million, or 1.6%, in the first three months of 2024. This demonstrates customer confidence and the strength and loyalty of our core deposit base. As of March 31, 2024, the 25 largest depositors represented 22.9% of total deposits, which is an increase from December 31, 2023, when it was 22.4%. As of March 31, 2024 and December 31, 2023, the estimated amount of uninsured deposits was $395.9 million and $384.5 million, respectively. The estimate of uninsured deposits was done at the single account level and does not take into account total customer balances in the Bank. It should be noted that approximately 50% of these uninsured deposits relate to public funds from municipalities, government entities, and school districts which by law are required to be collateralized by investment securities or FHLB letters of credit to protect these depositor funds. Total borrowings have decreased by $23.5 million, or 20.3%, since year-end 2023. This change was driven by a decrease in short-term borrowings which was partially offset by an increase in FHLB term advances. Specifically, short-term borrowings decreased by $29.5 million, or 72.0%. Given the high cost of overnight borrowed funds, management has been effectively controlling the usage of this funding source. In addition, the inversion in the yield curve has caused FHLB term advances to have rates that are lower than the cost of overnight borrowed funds. Therefore, management increased usage of FHLB term advances in the first quarter of 2024 leading to an increase in this line item of $4.9 million, or 10.9%.
The Company’s total shareholders’ equity increased by $1.7 million, or 1.6%, during the first three months of 2024. The increase in capital is the result of the Company’s earnings performance during the first quarter of 2024 more than offsetting our common stock dividend payments to shareholders. In addition, the improved market value adjustment on the interest rate hedges had a positive impact on accumulated other comprehensive loss which more than offset the reduced market value of the available for sale investment securities portfolio and the pension adjustment.
The Company continues to be considered well capitalized for regulatory purposes with a total capital ratio of 13.10%, and a common equity tier 1 capital ratio of 9.53% at March 31, 2024. See the discussion of the Basel III capital requirements under the Capital Resources section below. As of March 31, 2024, the Company’s book value per common share was $6.06 and its tangible book value per common share was $5.26(1). When compared to December 31, 2023, book value per common share and tangible book value per common share improved by $0.10 per common share. The improvement in the Company’s book value and tangible book value per share in the first three months of 2024 compared to last year-end reflects the improvement in earnings performance as well as a positive market value adjustment for the interest rate hedges which more than offset a decrease in the fair value of the Company’s available for sale investment securities and a negative pension adjustment. The tangible common equity to tangible assets ratio was 6.58%(1) at March 31, 2024 and increased by 14-basis points when compared to December 31, 2023.
(1) Non-GAAP financial information, see “Reconciliation of Non-GAAP Financial Measures” later in this MD&A.