Average outstanding loans receivable increased $82,413,000 (5.2%) to $1,674,270,000 in 2022 from $1,591,857,000 in 2021, despite a reduction in average PPP loans of $83,038,000. Average total loans outstanding, excluding PPP loans, increased $165,451,000 (11.0%).
The average yield on loans in the third quarter 2022 was 4.91%, up from 4.76% in the third quarter 2021. Excluding PPP loans, the average yield on loans was 4.90% in the third quarter 2022, up from 4.60% excluding PPP loans in the third quarter 2021. The increase in loan yields reflects the impact of higher interest rates on loans originated in 2022 and higher yields on floating-rate loans. Floating-rate loans totaled approximately 18% of gross loans receivable at September 30, 2022.
Interest income from available-for-sale debt securities increased $939,000 in 2022 from 2021. The average balance of available-for-sale debt securities (at amortized cost) increased to $564,920,000 in 2022 from $391,148,000 in 2021. The increase in available-for-sale debt securities reflects the investment of funds, primarily in the fourth quarter 2021 and first quarter 2022, that would otherwise have represented excess cash. The average yield on available-for-sale debt securities was 2.17% for 2022, down slightly from 2.18% in 2021.
Income from interest-bearing due from banks totaled $176,000 in 2022, an increase of $70,000 from 2021. The average yield on interest-bearing due from banks was 2.03% in 2022 and 0.22% in 2021. The average balance of interest-bearing due from banks was $34,465,000 in the third quarter 2022, down from $195,359,000 in the third quarter 2021. Within this category, the largest asset balance in 2022 and 2021 has been interest-bearing deposits held with the Federal Reserve.
INTEREST EXPENSE AND INTEREST-BEARING LIABILITIES
For the three-month periods, interest expense increased $1,217,000 to $2,831,000 in 2022 from $1,614,000 in 2021. Interest expense on deposits increased $909,000, as the average rate on interest-bearing deposits increased to 0.54% in 2022 from 0.30% in 2021. The increase in average rate on deposits includes increases of 0.40% on time deposits, 0.34% on money market accounts and 0.22% on interest checking accounts. The Corporation’s deposit rates have increased in response to the impact on market rates of increases in the Fed Funds Target Rate. The Fed Funds Target Rate ranged from 0% to 0.25% throughout 2021, while the Federal Reserve implemented a series of rate increases in March, May, June, July, and September 2022 resulting in a Fed Funds Target Rate ranging from 3% to 3.25% at September 30, 2022.
Average total deposits increased $61,825,000 (3.2%) to $1,998,583,000 in the third quarter 2022 from $1,936,758,000 in the third quarter 2021. Average time deposits decreased $14,351,000 and average money market accounts decreased $7,615,000, while the average total balance of other categories of noninterest-bearing demand and other deposits increased $83,791,000. The increase in average deposits includes the impact of funding received by consumers, businesses and municipal entities from government stimulus programs as well as growth in commercial deposits from new business.
Interest expense on short-term borrowings in the third quarter 2022 was $179,000 as compared to less than $1,000 in 2021. The average balance of short-term borrowings increased to $33,970,000 in 2022 from $2,185,000 in 2021 reflecting an increase in overnight borrowings to provide temporary funding to support loan growth. The average rate on short-term borrowings was 2.09% in 2022.
Interest expense on long-term borrowings (FHLB advances) increased $245,000 to $332,000 in 2022 from $87,000 in 2021. The average balance of long-term borrowings was $51,628,000 in 2022, up from an average balance of $41,083,000 in 2021. Borrowings are classified as long-term within the Tables based on their term at origination or assumption in business combinations. The average rate on long-term borrowings was 2.55% in 2022 compared to 0.84% in 2021.
Interest expense on subordinated debt decreased $117,000 to $229,000 in 2022 from $346,000 in 2021. The average balance of subordinated debt decreased to $24,566,000 in 2022 from $32,978,000 in 2021. The average rate on subordinated debt decreased to 3.70% in 2022 from 4.16% in 2021. In the second quarter 2022, the Corporation redeemed subordinated debt with aggregate par values of $8.5 million and a weighted average interest rate of 6.29%.
More information regarding the terms of borrowed funds is provided in Note 8 to the unaudited consolidated financial statements.