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JOINT FILING AGREEMENT
Pursuant to Rule 13d-1(k)(1) promulgated pursuant to the Securities Exchange Act of 1934, as amended, the undersigned agree that the attached Schedule 13D/A is being filed on behalf of each of the undersigned.
Dated: March 15, 2010
VENROCK ASSOCIATES
By: A General Partner
VENROCK ASSOCIATES III, L.P.
By: Venrock Management III, LLC
Its: General Partner
VENROCK MANAGEMENT III, LLC
VENROCK ENTREPRENEURS FUND III, L.P.
By: VEF Management III, LLC
Its: General Partner
VEF MANAGEMENT III, LLC
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| By: | David L. Stepp | |
| | Name: | David L. Stepp | |
| | Title: | Authorized Signatory | |
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