Verona Pharma (VRNA) SC 13G/A Beneficial ownership report (amended)

Exhibit 99.1

Joint Filing Agreement

The undersigned persons, on February 12, 2019, agree and consent to the joint filing on their behalf of this Schedule 13G in connection with their beneficial ownership of the Common Stock of Verona Pharma plc on December 31, 2018.

Tekla Capital Management LLC
By	/s/ Laura Woodward
Laura Woodward
Daniel R. Omstead
By	/s/ Daniel R. Omstead
Daniel R. Omstead