ARK Investment Management 13F-HR Quarterly holdings report by institutional manager

The Securities and Exchange Commission has not necessarily reviewed the information in this filing and has not determined if it is accurate and complete.
The reader should not assume that the information is accurate and complete.

UNITED STATES SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549
FORM 13F

FORM 13F INFORMATION TABLE

OMB APPROVAL

OMB Number:		3235-0006
Expires:	Oct 31, 2018
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COLUMN 1	COLUMN 2	COLUMN 3	COLUMN 4	COLUMN 5			COLUMN 6	COLUMN 7	COLUMN 8
			VALUE	SHRS OR	SH/	PUT/	INVESTMENT	OTHER	VOTING AUTHORITY
NAME OF ISSUER	TITLE OF CLASS	CUSIP	(x$1000)	PRN AMT	PRN	CALL	DISCRETION	MANAGER	SOLE	SHARED	NONE
Adobe Inc	Common Stock	00724F101	7,548	23,718	SH		SOLE		23,718	0	0
Aerojet Rocketdyne Holdings In	Common Stock	007800105	238	5,693	SH		SOLE		5,693	0	0
AeroVironment Inc	Common Stock	008073108	22,605	370,821	SH		SOLE		300,283	44,930	25,608
Alibaba Group Holding Ltd	ADR	01609W102	20,415	104,970	SH		SOLE		72,578	10,183	22,209
Align Technology Inc	Common Stock	016255101	258	1,484	SH		SOLE		1,484	0	0
Allegheny Technologies Inc	Common Stock	01741R102	18	2,130	SH		SOLE		2,130	0	0
Alphabet Inc	Common Stock	02079K305	1,571	1,352	SH		SOLE		1,352	0	0
Altair Engineering Inc	Common Stock	021369103	965	36,416	SH		SOLE		36,416	0	0
Amazon.com Inc	Common Stock	023135106	27,846	14,282	SH		SOLE		12,944	0	1,338
AMETEK Inc	Common Stock	031100100	294	4,084	SH		SOLE		4,084	0	0
ANSYS Inc	Common Stock	03662Q105	2,962	12,741	SH		SOLE		12,741	0	0
Apple Inc	Common Stock	037833100	22,200	87,302	SH		SOLE		70,959	0	16,343
AquaBounty Technologies Inc	Common Stock	03842K200	747	458,408	SH		SOLE		458,408	0	0
Howmet Aerospace Inc	Common Stock	443201108	127	9,850	SH		SOLE		9,850	0	0
Arcturus Therapeutics Holdings	Common Stock	03969T109	21,261	1,564,457	SH		SOLE		1,564,457	0	0
Autodesk Inc	Common Stock	052769106	68,359	437,918	SH		SOLE		334,505	44,033	59,380
Baidu Inc	ADR	056752108	1,582	15,694	SH		SOLE		15,694	0	0
Renishaw PLC	Common Stock	G75006117	1,358	34,609	SH		SOLE		34,609	0	0
CareDx Inc	Common Stock	14167L103	17,205	788,144	SH		SOLE		788,144	0	0
Carpenter Technology Corp	Common Stock	144285103	16	844	SH		SOLE		844	0	0
Castle Biosciences Inc	Common Stock	14843C105	6,572	220,452	SH		SOLE		220,452	0	0
Cellectis SA	ADR	15117K103	14,242	1,548,046	SH		SOLE		1,363,749	174,111	10,186
Cellular Biomedicine Group Inc	Common Stock	15117P102	6,322	399,632	SH		SOLE		399,632	0	0
Cerus Corp	Common Stock	157085101	65,124	14,005,056	SH		SOLE		11,619,631	311,251	2,074,174
Codexis Inc	Common Stock	192005106	11,040	989,249	SH		SOLE		989,249	0	0
Deere & Co	Common Stock	244199105	2,410	17,447	SH		SOLE		17,447	0	0
DuPont de Nemours Inc	Common Stock	26614N102	22	648	SH		SOLE		648	0	0
Eastman Chemical Co	Common Stock	277432100	24	512	SH		SOLE		512	0	0
Eastman Kodak Co	Common Stock	277461406	20	11,467	SH		SOLE		11,467	0	0
Editas Medicine Inc	Common Stock	28106W103	105,812	5,335,941	SH		SOLE		4,128,742	445,150	762,049
Eventbrite Inc	Common Stock	29975E109	464	63,530	SH		SOLE		3,314	9,767	50,449
ExOne Co/The	Common Stock	302104104	16,650	2,605,683	SH		SOLE		2,224,864	0	380,819
FLIR Systems Inc	Common Stock	302445101	3,826	119,984	SH		SOLE		119,984	0	0
Facebook Inc	Common Stock	30303M102	1,012	6,066	SH		SOLE		6,066	0	0
FARO Technologies Inc	Common Stock	311642102	332	7,462	SH		SOLE		7,462	0	0
Fate Therapeutics Inc	Common Stock	31189P102	12,328	555,082	SH		SOLE		555,082	0	0
General Electric Co	Common Stock	369604103	208	26,189	SH		SOLE		26,189	0	0
Grayscale Bitcoin Trust BTC	Open-End Fund	389637109	20,226	2,840,675	SH		SOLE		1,561,407	511,706	767,562
Guardant Health Inc	Common Stock	40131M109	17,237	247,651	SH		SOLE		247,651	0	0
Guidewire Software Inc	Common Stock	40171V100	975	12,292	SH		SOLE		12,292	0	0
HDFC Bank Ltd	ADR	40415F101	2,031	52,809	SH		SOLE		52,809	0	0
HP Inc	Common Stock	40434L105	1,329	76,567	SH		SOLE		76,567	0	0
Hexcel Corp	Common Stock	428291108	19	512	SH		SOLE		512	0	0
HubSpot Inc	Common Stock	443573100	6,791	50,989	SH		SOLE		50,989	0	0
Illumina Inc	Common Stock	452327109	274,220	1,004,026	SH		SOLE		786,687	85,525	131,814
Incyte Corp	Common Stock	45337C102	14,960	204,284	SH		SOLE		204,284	0	0
Inovio Pharmaceuticals Inc	Common Stock	45773H201	12,186	1,637,926	SH		SOLE		1,637,926	0	0
Intellia Therapeutics Inc	Common Stock	45826J105	125,226	10,239,286	SH		SOLE		8,559,768	740,852	938,666
Interactive Brokers Group Inc	Common Stock	45841N107	1,309	30,321	SH		SOLE		30,321	0	0
Intercontinental Exchange Inc	Common Stock	45866F104	3,168	39,232	SH		SOLE		39,232	0	0
Intuit Inc	Common Stock	461202103	1,961	8,526	SH		SOLE		8,526	0	0
Intuitive Surgical Inc	Common Stock	46120E602	2,796	5,647	SH		SOLE		5,647	0	0
OC Oerlikon Corp AG	Common Stock	H59187106	215	26,791	SH		SOLE		26,791	0	0
Invitae Corp	Common Stock	46185L103	220,015	16,094,750	SH		SOLE		12,826,319	1,049,935	2,218,496
Ionis Pharmaceuticals Inc	Common Stock	462222100	5,659	119,681	SH		SOLE		119,681	0	0
Iovance Biotherapeutics Inc	Common Stock	462260100	26,558	887,179	SH		SOLE		887,179	0	0
Iridium Communications Inc	Common Stock	46269C102	36,575	1,637,951	SH		SOLE		1,268,741	157,278	211,932
JD.com Inc	ADR	47215P106	3,117	76,962	SH		SOLE		76,962	0	0
SGL Carbon SE	Common Stock	D6949M108	21	7,817	SH		SOLE		7,817	0	0
Kaiser Aluminum Corp	Common Stock	483007704	200	2,887	SH		SOLE		2,887	0	0
Komatsu Ltd	ADR	500458401	3,356	204,514	SH		SOLE		204,514	0	0
Kratos Defense & Security Solu	Common Stock	50077B207	3,832	276,862	SH		SOLE		276,862	0	0
Henkel AG & Co KGaA	Preference	D3207M110	31	386	SH		SOLE		386	0	0
Bertrandt AG	Common Stock	D1014N107	175	5,157	SH		SOLE		5,157	0	0
LendingClub Corp	Common Stock	52603A208	43,180	5,500,633	SH		SOLE		4,644,507	127,913	728,213
LendingTree Inc	Common Stock	52603B107	87,236	475,685	SH		SOLE		360,881	38,659	76,145
Dassault Systemes SE	Common Stock	F2457H472	1,220	8,237	SH		SOLE		8,237	0	0
Lincoln Electric Holdings Inc	Common Stock	533900106	217	3,141	SH		SOLE		3,141	0	0
Siemens AG	Common Stock	D69671218	200	2,399	SH		SOLE		2,399	0	0
Materialise NV	ADR	57667T100	62,751	3,379,164	SH		SOLE		3,072,602	73,645	232,917
MercadoLibre Inc	Common Stock	58733R102	23,366	47,825	SH		SOLE		31,403	7,808	8,614
Microsoft Corp	Common Stock	594918104	1,345	8,527	SH		SOLE		8,527	0	0
Bezeq The Israeli Telecommunic	Common Stock	M2012Q100	560	774,147	SH		SOLE		774,147	0	0
Bet Shemesh Engines Holdings 1	Common Stock	M20119109	352	14,025	SH		SOLE		14,025	0	0
Moog Inc	Common Stock	615394202	163	3,224	SH		SOLE		3,224	0	0
Rakuten Inc	Common Stock	J64264104	1,158	152,213	SH		SOLE		152,213	0	0
Danel Adir Yeoshua Ltd	Common Stock	M27013107	438	6,275	SH		SOLE		6,275	0	0
Hilan Ltd	Common Stock	M5299H123	418	13,286	SH		SOLE		13,286	0	0
Nano Dimension Ltd	ADR	63008G203	71	75,731	SH		SOLE		75,731	0	0
NanoString Technologies Inc	Common Stock	63009R109	73,392	3,051,660	SH		SOLE		2,787,727	255,664	8,269
Elbit Systems Ltd	Common Stock	M3760D101	2,369	18,551	SH		SOLE		18,551	0	0
SBI Holdings Inc/Japan	Common Stock	J6991H100	1,893	129,300	SH		SOLE		129,300	0	0
Partner Communications Co Ltd	Common Stock	M78465107	508	136,332	SH		SOLE		136,332	0	0
Netflix Inc	Common Stock	64110L106	13,697	36,477	SH		SOLE		27,492	1,894	7,091
Nice Ltd	ADR	653656108	473	3,295	SH		SOLE		3,295	0	0
FirstRand Ltd	Common Stock	S5202Z131	58	13,580	SH		SOLE		13,580	0	0
NVIDIA Corp	Common Stock	67066G104	41,757	158,409	SH		SOLE		121,655	17,034	19,720
NuVasive Inc	Common Stock	670704105	198	3,907	SH		SOLE		3,907	0	0
Matrix IT Ltd	Common Stock	M6859E153	433	26,419	SH		SOLE		26,419	0	0
Organovo Holdings Inc	Common Stock	68620A104	9,394	22,912,766	SH		SOLE		22,842,197	70,569	0
Toray Industries Inc	Common Stock	J89494116	25	5,808	SH		SOLE		5,808	0	0
PTC Inc	Common Stock	69370C100	1,089	17,786	SH		SOLE		17,786	0	0
Pacific Biosciences of Califor	Common Stock	69404D108	12,831	4,193,059	SH		SOLE		4,193,059	0	0
One Software Technologies Ltd	Common Stock	M9866H102	486	7,376	SH		SOLE		7,376	0	0
PayPal Holdings Inc	Common Stock	70450Y103	10,424	108,882	SH		SOLE		104,332	4,550	0
Personalis Inc	Common Stock	71535D106	8,910	1,104,065	SH		SOLE		1,104,065	0	0
Straumann Holding AG	Common Stock	H8300N119	1,199	1,606	SH		SOLE		1,606	0	0
Pinduoduo Inc	ADR	722304102	7,678	213,112	SH		SOLE		213,112	0	0
Pinterest Inc	Common Stock	72352L106	86,602	5,608,922	SH		SOLE		4,561,180	419,526	628,216
Pluristem Therapeutics Inc	Common Stock	72940R300	2,423	660,150	SH		SOLE		660,150	0	0
PolyOne Corp	Common Stock	73179P106	22	1,171	SH		SOLE		1,171	0	0
Proto Labs Inc	Common Stock	743713109	122,012	1,602,676	SH		SOLE		1,229,553	150,407	222,716
Wirecard AG	Common Stock	D22359133	1,302	11,536	SH		SOLE		11,536	0	0
Rakuten Inc	ADR	75102W108	102	13,623	SH		SOLE		13,623	0	0
Redhill Biopharma Ltd	ADR	757468103	436	96,151	SH		SOLE		96,151	0	0
Regeneron Pharmaceuticals Inc	Common Stock	75886F107	43	89	SH		SOLE		89	0	0
Roku Inc	Common Stock	77543R102	147,566	1,686,850	SH		SOLE		1,326,491	147,920	212,439
Rubius Therapeutics Inc	Common Stock	78116T103	1,416	318,121	SH		SOLE		28,130	270,697	19,294
salesforce.com Inc	Common Stock	79466L302	19,896	138,189	SH		SOLE		50,749	39,789	47,651
Charles Schwab Corp/The	Common Stock	808513105	1,204	35,805	SH		SOLE		35,805	0	0
Seres Therapeutics Inc	Common Stock	81750R102	30,871	8,647,311	SH		SOLE		7,152,461	649,867	844,983
Shopify Inc	Common Stock	82509L107	168	402	SH		SOLE		402	0	0
Snap Inc	Common Stock	83304A106	12,667	1,065,328	SH		SOLE		1,065,328	0	0
SolarEdge Technologies Inc	Common Stock	83417M104	465	5,683	SH		SOLE		5,683	0	0
Splunk Inc	Common Stock	848637104	95,287	754,867	SH		SOLE		588,884	67,951	98,032
Square Inc	Common Stock	852234103	250,317	4,778,869	SH		SOLE		3,703,086	405,410	670,373
Stryker Corp	Common Stock	863667101	236	1,415	SH		SOLE		1,415	0	0
Syros Pharmaceuticals Inc	Common Stock	87184Q107	34,983	5,899,398	SH		SOLE		5,143,384	92,721	663,293
Taiwan Semiconductor Manufactu	ADR	874039100	3,404	71,236	SH		SOLE		71,236	0	0
Teladoc Health Inc	Common Stock	87918A105	19,701	127,094	SH		SOLE		73,708	26,795	26,591
Tencent Holdings Ltd	ADR	88032Q109	22,811	464,671	SH		SOLE		423,600	38,335	2,736
Teradyne Inc	Common Stock	880770102	39,112	722,022	SH		SOLE		571,292	78,618	72,112
Tesla Inc	Common Stock	88160R101	312,644	596,649	SH		SOLE		535,039	58,845	2,765
Teva Pharmaceutical Industries	ADR	881624209	504	56,123	SH		SOLE		56,123	0	0
Trade Desk Inc/The	Common Stock	88339J105	5,778	29,939	SH		SOLE		29,939	0	0
3D Systems Corp	Common Stock	88554D205	1,123	145,604	SH		SOLE		145,604	0	0
3M Co	Common Stock	88579Y101	32	232	SH		SOLE		232	0	0
TransUnion	Common Stock	89400J107	1,067	16,123	SH		SOLE		16,123	0	0
Trimble Inc	Common Stock	896239100	5,916	185,873	SH		SOLE		185,873	0	0
Twilio Inc	Common Stock	90138F102	13,632	152,325	SH		SOLE		152,325	0	0
Twist Bioscience Corp	Common Stock	90184D100	19,696	644,093	SH		SOLE		644,093	0	0
Twitter Inc	Common Stock	90184L102	34,284	1,395,937	SH		SOLE		820,007	216,879	359,051
2U Inc	Common Stock	90214J101	143,422	6,758,820	SH		SOLE		5,325,329	531,142	902,349
Ultra Clean Holdings Inc	Common Stock	90385V107	170	12,354	SH		SOLE		12,354	0	0
United Parcel Service Inc	Common Stock	911312106	242	2,586	SH		SOLE		2,586	0	0
Veracyte Inc	Common Stock	92337F107	79,393	3,265,859	SH		SOLE		3,190,818	75,041	0
Verisk Analytics Inc	Common Stock	92345Y106	1,176	8,436	SH		SOLE		8,436	0	0
Workday Inc	Common Stock	98138H101	7,729	59,357	SH		SOLE		59,357	0	0
Xilinx Inc	Common Stock	983919101	120,968	1,552,066	SH		SOLE		1,234,268	124,117	193,681
Xerox Holdings Corp	Common Stock	98421M106	22	1,162	SH		SOLE		1,162	0	0
Zillow Group Inc	Common Stock	98954M101	113,760	3,348,848	SH		SOLE		2,493,679	325,643	529,526
Koninklijke DSM NV	Common Stock	N5017D122	37	329	SH		SOLE		329	0	0
Arkema SA	Common Stock	F0392W125	28	408	SH		SOLE		408	0	0
MGI Digital Graphic Technology	Common Stock	F4090S113	1,089	29,116	SH		SOLE		29,116	0	0
Sandvik AB	Common Stock	W74857165	31	2,212	SH		SOLE		2,212	0	0
Hexagon AB	Common Stock	W40063104	312	7,347	SH		SOLE		7,347	0	0
Cellcom Israel Ltd	Common Stock	M2196U109	535	182,654	SH		SOLE		182,654	0	0
Evonik Industries AG	Common Stock	D2R90Y117	30	1,430	SH		SOLE		1,430	0	0
Fattal Holdings 1998 Ltd	Common Stock	M4148G105	126	3,668	SH		SOLE		3,668	0	0
Meituan Dianping	Common Stock	G59669104	2,945	243,727	SH		SOLE		243,727	0	0
SLM Solutions Group AG	Common Stock	D6T690109	837	108,313	SH		SOLE		108,313	0	0
ZhongAn Online P&C Insurance C	Common Stock	Y989DF109	2,101	636,100	SH		SOLE		636,100	0	0
Adyen NV	Common Stock	N3501V104	2,432	2,877	SH		SOLE		2,877	0	0
Atlassian Corp PLC	Common Stock	G06242104	6,028	43,915	SH		SOLE		43,915	0	0
Medtronic PLC	Common Stock	G5960L103	229	2,544	SH		SOLE		2,544	0	0
Aptiv PLC	Common Stock	G6095L109	2,715	55,136	SH		SOLE		53,235	1,901	0
Sapiens International Corp NV	Common Stock	G7T16G103	431	22,637	SH		SOLE		22,637	0	0
CRISPR Therapeutics AG	Common Stock	H17182108	181,754	4,285,632	SH		SOLE		3,342,383	337,686	605,563
Spotify Technology SA	Common Stock	L8681T102	6,764	55,695	SH		SOLE		37,241	17,130	1,324
Allot Ltd	Common Stock	M0854Q105	572	60,495	SH		SOLE		60,495	0	0
AudioCodes Ltd	Common Stock	M15342104	494	20,677	SH		SOLE		20,677	0	0
Camtek Ltd/Israel	Common Stock	M20791105	387	46,013	SH		SOLE		46,013	0	0
Ceragon Networks Ltd	Common Stock	M22013102	327	259,697	SH		SOLE		259,697	0	0
Check Point Software Technolog	Common Stock	M22465104	507	5,044	SH		SOLE		5,044	0	0
Compugen Ltd	Common Stock	M25722105	74,969	10,326,360	SH		SOLE		9,953,375	212,075	160,910
CyberArk Software Ltd	Common Stock	M2682V108	354	4,133	SH		SOLE		4,133	0	0
Evogene Ltd	Common Stock	M4119S104	302	256,129	SH		SOLE		256,129	0	0
Galmed Pharmaceuticals Ltd	Common Stock	M47238106	305	88,243	SH		SOLE		88,243	0	0
Fiverr International Ltd	Common Stock	M4R82T106	552	21,942	SH		SOLE		21,942	0	0
Mellanox Technologies Ltd	Common Stock	M51363113	585	4,826	SH		SOLE		4,826	0	0
Ituran Location and Control Lt	Common Stock	M6158M104	320	22,516	SH		SOLE		22,516	0	0
Nova Measuring Instruments Ltd	Common Stock	M7516K103	475	14,555	SH		SOLE		14,555	0	0
Radware Ltd	Common Stock	M81873107	473	22,426	SH		SOLE		22,426	0	0
Stratasys Ltd	Common Stock	M85548101	156,232	9,795,137	SH		SOLE		7,517,884	733,741	1,543,512
Taro Pharmaceutical Industries	Common Stock	M8737E108	415	6,779	SH		SOLE		6,779	0	0
Tower Semiconductor Ltd	Common Stock	M87915274	362	22,757	SH		SOLE		22,757	0	0
UroGen Pharma Ltd	Common Stock	M96088105	300	16,821	SH		SOLE		16,821	0	0
Wix.com Ltd	Common Stock	M98068105	408	4,047	SH		SOLE		4,047	0	0
NXP Semiconductors NV	Common Stock	N6596X109	3,281	39,559	SH		SOLE		39,559	0	0
Bayer AG	Common Stock	D0712D163	22	390	SH		SOLE		0	390	0
FANUC Corp	Common Stock	J13440102	30	220	SH		SOLE		0	220	0
Nintendo Co Ltd	Common Stock	J51699106	66	171	SH		SOLE		0	171	0
SoftBank Group Corp	Common Stock	J75963108	39	1,110	SH		SOLE		0	1,110	0
DocuSign Inc	Common Stock	256163106	1,246	13,489	SH		SOLE		13,489	0	0
Zscaler Inc	Common Stock	98980G102	6,639	109,084	SH		SOLE		109,084	0	0
Wirecard AG	ADR	97654L108	202	3,594	SH		SOLE		3,594	0	0
GMO internet Inc	Common Stock	J1822R104	45	2,700	SH		SOLE		2,700	0	0
Discovery Ltd	Common Stock	S2192Y109	841	191,917	SH		SOLE		191,917	0	0
Inmode Ltd	Common Stock	M5425M103	280	13,013	SH		SOLE		13,013	0	0
Perion Network Ltd	Common Stock	M78673114	323	66,012	SH		SOLE		66,012	0	0
Silvergate Capital Corp	Common Stock	82837P408	470	49,251	SH		SOLE		49,251	0	0
Alteryx Inc	Common Stock	02156B103	5,866	61,638	SH		SOLE		61,638	0	0
Meituan Dianping	ADR	58533E103	401	16,871	SH		SOLE		16,871	0	0
Z Holdings Corp	Common Stock	J9894K105	2,868	888,120	SH		SOLE		888,120	0	0
Sea Ltd	ADR	81141R100	9,895	223,322	SH		SOLE		223,322	0	0
Z Holdings Corp	ADR	98877X101	419	66,488	SH		SOLE		66,488	0	0
Pure Storage Inc	Common Stock	74624M102	53,112	4,318,047	SH		SOLE		3,615,952	344,340	357,755
Synopsys Inc	Common Stock	871607107	7,445	57,808	SH		SOLE		57,808	0	0
HUYA Inc	ADR	44852D108	9,384	553,613	SH		SOLE		553,613	0	0
RADA Electronic Industries Ltd	Common Stock	M81863124	306	87,438	SH		SOLE		87,438	0	0
Sol-Gel Technologies Ltd	Common Stock	M8694L103	290	41,098	SH		SOLE		41,098	0	0
PagerDuty Inc	Common Stock	69553P100	45,058	2,607,525	SH		SOLE		2,203,031	89,553	314,941
Bellicum Pharmaceuticals Inc	Common Stock	079481404	0	49	SH		SOLE		49	0	0
Precigen Inc	Common Stock	74017N105	1	281	SH		SOLE		281	0	0
Caterpillar Inc	Common Stock	149123101	3,084	26,577	SH		SOLE		26,577	0	0
Rockwell Automation Inc	Common Stock	773903109	1,471	9,747	SH		SOLE		9,747	0	0
Menlo Therapeutics Inc	Common Stock	586858102	197	73,329	SH		SOLE		73,329	0	0